Home About us Contact
|
Home About us Contact | ||
|
|
||
Systematics
Kinds of Systematics Selected AbstractsRAPHIDOPHYCEAE [CHADEFAUD EX SILVA] SYSTEMATICS AND RAPID IDENTIFICATION: SEQUENCE ANALYSES AND REAL-TIME PCR ASSAYS,JOURNAL OF PHYCOLOGY, Issue 6 2006Holly A. Bowers Species within the class Raphidophyceae were associated with fish kill events in Japanese, European, Canadian, and U.S. coastal waters. Fish mortality was attributable to gill damage with exposure to reactive oxygen species (peroxide, superoxide, and hydroxide radicals), neurotoxins, physical clogging, and hemolytic substances. Morphological identification of these organisms in environmental water samples is difficult, particularly when fixatives are used. Because of this difficulty and the continued global emergence of these species in coastal estuarine waters, we initiated the development and validation of a suite of real-time polymerase chain reaction (PCR) assays. Sequencing was used to generate complete data sets for nuclear encoded small-subunit ribosomal RNA (SSU rRNA; 18S); internal transcribed spacers 1 and 2, 5.8S; and plastid encoded SSU rRNA (16S) for confirmed raphidophyte cultures from various geographic locations. Sequences for several Chattonella species (C. antiqua, C. marina, C. ovata, C. subsalsa, and C. verruculosa), Heterosigma akashiwo, and Fibrocapsa japonica were generated and used to design rapid and specific PCR assays for several species including C. verruculosa Hara et Chihara, C. subsalsa Biecheler, the complex comprised of C. marina Hara et Chihara, C. antiqua Ono and C. ovata, H. akashiwo Ono, and F. japonica Toriumi et Takano using appropriate loci. With this comprehensive data set, we were also able to perform phylogenetic analyses to determine the relationship between these species. [source] SYSTEMATICS OF THE HILDENBRANDIALES (RHODOPHYTA): GENE SEQUENCE AND MORPHOMETRIC ANALYSES OF GLOBAL COLLECTIONS,JOURNAL OF PHYCOLOGY, Issue 2 2003Alison R. Sherwood Fifty-seven collections of marine and freshwater Hildenbrandia from North America, South America, Europe, and Africa were compared with 21 type and historically important specimens using multivariate morphometrics. Additionally, phylogenetic analyses of 48 specimens of Hildenbrandia and two specimens of Apophlaea were carried out based on sequences of the rbcL chloroplast gene and the nuclear 18S rRNA gene. Morphometric analyses based on vegetative cell and filament dimensions distinguished two groups of freshwater Hildenbrandia specimens, the first corresponding to those collections from North America and the Philippines and the second to those from Europe and the Canary Islands. The first group had smaller mean cell and filament dimensions (cells 4.0 × 4.4 ,m, filaments 46.5 ,m) and corresponded to H. angolensis, whereas the second group had larger mean dimensions (cells 5.8 × 6.6 ,m, filaments 55.3 ,m) and represented H. rivularis. Marine specimens were morphometrically distinguishable into two groups based on tetrasporangial division pattern as well as other thallus characters. However, measurements and character determinations of some type specimens differed greatly from the original descriptions, and thus further work to determine the stability of these characters is required. Phylogenetic reconstruction based on the 18S rRNA gene and rbcL gene sequence data generally demonstrated separation of the marine and freshwater forms of Hildenbrandia, with some marine taxa forming monophyletic groups (e.g. H. lecannellieri and H. occidentalis) and others forming paraphyletic groups (e.g. H. rubra). The two specimens of Apophlaea formed a monophyletic group within the paraphyletic genus Hildenbrandia. [source] PHYLOGENETIC SYSTEMATICS OF THE ULVACEAE (ULVALES, ULVOPHYCEAE) USING CHLOROPLAST AND NUCLEAR DNA SEQUENCES,JOURNAL OF PHYCOLOGY, Issue 6 2002Hillary S. Hayden Systematic hypotheses for the Ulvaceae were tested using phylogenetic analysis of sequences for the gene encoding the large subunit of RUBISCO, small subunit rDNA and a combined data matrix. Representatives of eight putative ulvaceous genera and twelve additional taxa from the Ulvophyceae and Trebouxiophyceae were included in analyses using maximum parsimony and maximum likelihood criteria. Molecular data supported hypotheses for the Ulvaceae that are based on the early development of vegetative thalli and motile cell ultrastructure. Ulvaceae sensu Floyd and O'Kelly, including Percursaria Bory de Saint-Vincent, Ulvaria Ruprecht and a complex of closely related species of Chloropelta Tanner, Enteromorpha Link and Ulva L. was supported; however, monophyly of Enteromorpha and Ulva was not supported. The Ulvales and Ulotrichales sensu Floyd and O'Kelly were monophyletic. Blidingia Kylin and Kornmannia Bliding were allied with the former and Capsosiphon Gobi with the latter, although relationships among these and other taxa in these orders remain uncertain. The Ulvales are characterized by an isomorphic life history pattern, gametangia and sporangia that are identical in structure and development, motile cells with bilobed terminal caps and proximal sheaths consisting of two equal subunits. Method of motile cell release and the gross morphology of vegetative thalli are not systematically reliable characters. [source] A REEXAMINATION OF THE SYSTEMATICS OF THE ACROCHAETIALES (RHODOPHYTA) USING LARGE-SUBUNIT RDNA SEQUENCE DATAJOURNAL OF PHYCOLOGY, Issue 2000J.T. Harper The taxonomic history of the red algal order Acrochaetiales is chaotic. There is no consensus in the literature as to how many genera should be recognized or in the assignment of the over 400 species to these genera. Morphological and anatomical studies have provided a suite of possible characters to delineate genera within this order, but there have been major discrepancies in the assessment and use of these features. The phylogenetic placement of the Acrochaetiales has also been the focus of debate. Once thought to be the most ancestral florideophyte lineage, recent molecular systematic studies have illustrated that this order is a derived lineage closely related to the Nemaliales and Palmariales. Phylogenies using sequences of the small-subunit (SSU) rDNA have strongly supported two very divergent lineages within a possibly polyphyletic Acrochaetiales. The relationships between these two groups and among other closely related rhodophyte orders were not resolved. We have generated large-subunit (LSU) rDNA sequence data for representatives of the Acrochaetiales and related taxa. Distance and parsimony phylogenies based on LSU and combined SSU and LSU data will be presented. The increased phylogenetic signal afforded by this approach will shed light on previous conundrums in the systematics of this group. [source] A CANDID ASSESSMENT OF SYSTEMATICS IN THE LAMINARIALES: MUDDLED MORPHOLOGIES AND SKETCHY SEQUENCESJOURNAL OF PHYCOLOGY, Issue 2000G.W. Saunders Algae of the Alariaceae, Laminariaceae and Lessoniaceae are the largest, most complex, and, arguably, most fascinating of the seaweeds. It is, therefore, a profound paradox that these species remain in taxonomic chaos despite the contemporary emphases on systematics. Setchell and Gardner established the classification system in 1925, recognizing these families on the basis of clear morphological features. At that time, however, they acknowledged that some species had features consistent with placement in two families, or that obscured logical placement in any of the families. Ironically, the problems noted by Setchell and Gardner have been ignored and the system has become entrenched in kelp literature. Initial molecular studies highlighted the shortcomings noted by Setchell and Gardner, and further indicated that little of the morphology-based system was natural. It was obvious that the diagnostic morphological features, presence or absence of sporophylls and ontogenetic splitting, were ,noisy' being gained and lost independently many times in kelp evolution. Despite the insights of the initial molecular studies, they had limitations and key relationships remained unresolved. The investigations used the Internal Transcribed Spacers (ITS) of the ribosomal cistron that have phylogenetic limitations owing to their short length, which is exacerbated by the many variable regions that can't be aligned confidently and must be removed prior to phylogenetic analyses. Many molecular publications have appeared subsequent to the first ITS results, and their contribution towards elucidating kelp phylogeny will be assessed. Current investigations using the Large Subunit ribosomal DNA (LSU) for kelp systematics will also be discussed. [source] MOLECULAR SYSTEMATICS OF RIVER DOLPHINS INFERRED FROM COMPLETE MITOCHONDRIAL CYTOCHROME- B GENE SEQUENCESMARINE MAMMAL SCIENCE, Issue 1 2002Guang Yang Abstract 1,140 bp of the complete mitochondrial cytochrome- b gene sequences of baiji (Lipotes vexillifer), franciscana (Pontoporia blainvillei), and Ganges river dolphin (Platanista gangetica gangetica) were determined to address the systematic position and phylogeny of extant river dolphins with combination of homologous sequences of other cetaceans. The neighbor-joining (NJ), maximum parsimony (MP), and maximum likelihood (ML) phylogenetic analyses all identified the river dolphins into three lineages, i. e., Platanista, Lipotes, and Inia+Pontoporia. The Lipotes did not have sister relationship with either Platanista or Inia+Pontoporia, which strongly supported the referral of Lipotes to a separate family, i. e., Lipotidae. There were very high sequence divergences between all river dolphin genera, suggesting a relatively longer period of separation time than those among other odontocete families. [source] HIGHER SYSTEMATICS OF SCORPIONS FROM THE CRATO FORMATION, LOWER CRETACEOUS OF BRAZILPALAEONTOLOGY, Issue 1 2007FEDERICA MENON Abstract:, Several new specimens of Protoischnurus axelrodorum Carvalho and Lourenço and Araripescorpius ligabuei Campos from the Crato Formation, Brazil, are described. The preservation and recognition of new morphological features allows a re-diagnosis of both species and a modification of their familial placement. Protoischnurus axelrodorum is the oldest species belonging to the scorpionoid family Hemiscorpiidae Pocock (= Ischnuridae Simon; =,Liochelidae Fet and Bechly) and the first Cretaceous record. It was originally placed in the extinct family Protoischnuridae Carvalho and Lourenço, which is here synonymized with Hemiscorpiidae. Araripescorpius ligabuei, now assigned to Chactidae Pocock, is the first chactoid recorded for the Cretaceous of Brazil. These findings confirm that the lineages of two modern families date back at least to the Early Cretaceous and, considering their current distribution, were probably present before the break-up of the supercontinent Gondwana. Palaeoecological inferences indicate the presence of tropical habitats in the vicinity of the Crato lake/lagoon. [source] LATE ORDOVICIAN BRACHIOPODS FROM THE DULANKARA FORMATION OF THE CHU-ILI RANGE, KAZAKHSTAN: THEIR SYSTEMATICS, PALAEOECOLOGY AND PALAEOBIOGEOGRAPHYPALAEONTOLOGY, Issue 2 2006LEONID E. POPOV Abstract:, Brachiopods from the late Ordovician (late Caradoc) Dulankara Formation of the Chu-Ili Range, Kazakhstan, are reviewed. Those from the upper two members of the formation, the Degeres and Akkol members, are systematically described. New genera from the Dulankara Formation are Nikitinamena (Plectambonitoidea: Leptellinidae), with type species Nikitinamena bicostata sp. nov., and Weberorthis (Orthoidea: Plectorthidae), with type species Mimella brevis Rukavishnikova. Another new genus is Glyptomenoides (Strophomenoidea: Glyptomenidae), with type species Rafinesquina girvanensis Salmon from the Caradoc of Girvan, Scotland. Other new species from the Dulankara Formation are Holtedahlina orientalis, Platymena tersa, Christiania proclivis, Leangella (Leangella) paletsae, Metambonites subcarinatus, Ogmoplecia nesca and Plectorthis licta. The ecology and assemblages of all three members of the Dulankara Formation are identified or reviewed, and their palaeogeographical significance assessed: the Chu-Ili Terrane (on which the Dulankara Formation was situated during the Ordovician) formed part of the relatively low-latitude peri-Gondwanan complex of terranes, and was probably not far from North and South China. The faunal links suggested between the Dulankara brachiopods and contemporary faunas from Australia are now perceived to be weaker than previously thought. [source] SYSTEMATICS OF THE CAMBRIAN TRILOBITE FAMILY NEPEIDAE, WITH REVISION OF AUSTRALIAN SPECIESPALAEONTOLOGY, Issue 3 2005JOHN R. PATERSON Abstract:, Taxonomy of the Cambrian trilobite family Nepeidae is revised. Morphometric analyses of the genera Nepea and Penarosa demonstrate that use of proportions of linear dimensions of the cranidium to differentiate species is invalid, and that infrageneric variation is continuous. The taxonomy and biostratigraphy of all Australian species of Nepeidae is revised. Species considered valid herein include: Nepea narinosa (type species), N. tonsillata, N. nans, Penarosa retifera (type species), P. elaticeps, P. rhinodelphis, P. netenta, Loxonepea loxophrys (type species) and Ferenepea hispida (type species). Folliceps is considered to be a junior subjective synonym of Nepea, Trinepea is regarded as a junior subjective synonym of Penarosa, and Ascionepea is considered to be a junior subjective synonym of Ferenepea. [source] Bodensystematik und Bodenklassifikation Teil I: GrundbegriffeJOURNAL OF PLANT NUTRITION AND SOIL SCIENCE, Issue 1 2005Christoph Albrecht Abstract Bodenordnungssysteme lassen sich meist nach zwei Prinzipien entwickeln: Entweder werden nur rein bodenkundliche Informationen als kategorisierendes Merkmal verwendet (pedogenetische Faktoren/Prozesse), oder die Kategorienbildung erfolgt problemorientiert anhand ausgewählter Parameter. Die meisten der weltweit verwendeten Bodenordnungssysteme lassen sich nach ihrer Grundausrichtung einem der beiden Typen zuordnen. Diese Betrachtungsweise ist nicht neu und wird in der Literatur mit unterschiedlichen Begriffen und Begriffsinhalten dargestellt. In der vorliegenden Arbeit werden die verschiedenen Definitionen von Systematik, Klassifikation, Taxonomie und Identifizierung zusammengefasst und geordnet. Dabei fällt auf, dass Begriffe mit sehr unterschiedlichen Inhalten oft synonym verwendet werden. Grundgedanke unserer Überlegungen ist die Trennung von Systematik, Klassifikation und Identifizierung. Systematik ist die grundsätzliche wissenschaftlich-deduktive Gliederung von Objekten in systematische Einheiten. Dabei soll das gesamte Wissen eines Fachgebietes in eine überschaubare Form gebracht werden, im Mittelpunkt stehen sowohl die umfassende Beschreibung einzelner Objekte als auch die Beziehungen zwischen den Objekten. Im Gegensatz dazu ist eine Klassifikation die zielorientiert-induktive Gliederung von Objekten. Die entstehenden Klassen werden nur anhand ausgewählter Parameter abgegrenzt, womit ein schneller Überblick bei speziellen Fragestellungen ermöglicht wird. Die Identifizierung ist die Einordnung von neuen Objekten in eine bestehende Systematik oder Klassifikation. Eine zweifelsfreie Identifizierung erfordert die Messbarkeit der kategorisierenden Merkmale. Bei einer genetisch angelegten Bodensystematik sind die Merkmale die Boden bildenden Prozesse und Faktoren. Da sie beim gegenwärtigen Kenntnisstand oft nicht messbar sind, bleiben Versuche, einen Boden in eine Systematik einzuordnen, häufig hypothetisch und dadurch subjektiv. Die Ergebnisse einer Bodensystematisierung sind daher oft anfechtbar, weil sie nicht durch Messwerte verifiziert werden können. Im Gegensatz dazu erlauben Bodenklassifikationen objektive Profilansprachen. Da jedoch die Festlegung der Grenzwerte eher pragmatisch nach Zweckmäßigkeit geschieht und nicht wissenschaftlich anhand von Prozessintensitäten, ist die Verwendung als grundlegendes Ordnungssystem eines Wissenschaftsgebietes nicht möglich. Die Bodenkunde benötigt beide Arten von Ordnungssystemen, um wissenschaftliche und praktische Ansprüche gleichermaßen erfüllen zu können, jedoch erfordern die Vollendung und Verifizierung der Systematik umfangreiche Forschungsarbeiten. Kurzfristig ist dieses Problem nur durch die Entwicklung einer kennwertbasierten Klassifikation lösbar, mit der die Kategorien der bestehenden Systematik so gut wie möglich nachgebildet werden. Langfristig ist die exakte Erforschung und Modellierung der Boden bildenden Prozesse aber unumgänglich. Soil systematics and classification systems Part I: Fundamentals Soil-ordering systems are primarily based and developed on one of two underlying principles: They are either categorized according to soil-forming processes, or the formation of categories develops by chosen parameters. This perspective has already been established in the literature, though it is often confusing as many terms are defined and applied differently. In this contribution, the various definitions of systematics, classification, taxonomy, and identification will be clearly differentiated and summarized. The core of our work is to clearly define and contrast three terms: systematics, classification, and identification. Systematics is the fundamental scientific and deductive ordering of objects into systematic units. The purpose of this approach is to organize the entire spectrum of knowledge within a discipline into a transparent and manageable form. Classification, in direct contrast to systematics, is goal-oriented and an inductive ordering of objects. Thus, the ordering scheme consists of classes which are clearly parameterized. Identification is the ordering of new objects into an already existing systematics or classification system. Close attention is paid to both the differences and the similarities between a systematics and a classification system, especially pertaining to their practical applications. The identification requires that the category-forming characteristics can be measured (e.g., for soil systematics, these are the soil-forming processes and factors). Currently, it is unfortunately not feasible to objectively quantify most soil-forming processes. Thus, most attempts at categorizing soils by systematics are hypothetical and highly subjective in nature. The resulting identification derived from the soil systematics approach is open to questions and contestable, since a graded measuring system does not yet exist to verify these determinations. In contrast, a soil-classification system does allow an objective soil-profile identification, although such systems are conceived pragmatically and designed for a practical purpose (e.g., not scientifically based on process intensities). Unfortunately, such a classification system cannot be applied as a universal scientific categorization system due to this method of conception. Both categorization approaches are required in soil science in order to satisfy both the practical and the scientific aspects of the field. However, substantial research must be done to complete and verify systematics. The only viable short-term solution is through the development of a graded classification system where the categories of the system are directly derived from the current systematics approach. In the long run both the exact investigation and the detailed modeling of the soil-forming processes are inevitable. [source] Systematics and phylogeny of the Asclerosibutia -group of genera (Coleoptera, Oedemeridae), with description of a new genus and seven new species from tropical AfricaMITTEILUNGEN AUS DEM MUSEUM FUER NATURKUNDE IN BERLIN-DEUTSCHE ENTOMOLOGISCHE ZEITSCHRIFT, Issue 1 2004Xavier A. Vázquez Abstract The phylogenetic relationships of the Asclerosibutia-group are studied, based on the examination of the type species of all genera previously described. The phylogenetic hypothesis supports that the Asclerosibutia-group is a monophyletic assemblage. On the base of the cladistic analysis, three genera can be recognized, i.e., Idgiomimula Blair, 1926, gen. rediv., Monosigynes gen. n. and Asclerosibutia Pic, 1914, with the following relationships: Idgiomimula (Monosigynes (Asclerosibutia)). Seven new species are described: Idgiomimula uhligi sp. n., Monosigynes difformis sp. n., Monosigynes marci sp. n., Monosigynes zarazagai sp. n., Monosigynes sphyrninus sp. n., Asclerosibutia girardi sp. n. and Asclerosibutia beardae sp. n. Danerces semipicea Karsch. 1881. Microsessinia notatipennis Pic, 1922, Microsessinia rufithorax Pic, 1958, are transferred to the genus Monosigynes. Microsessinia cyanescens Pic, 1922 is transferred to Asclerosibutia and Asclerosibutia terminalis Blair, 1926 is transferred to Idgiomimula. The following new synonyms are established: Microsessinia Pic, 1922 (= Asclerosibutia Pic, 1914), Microsessinia notatipennis var. atrimembris Pic, 1935 (= Monosigynes notatipennis (Pic, 1922)), Asclerosibutia gabonica Pic, 1914. Nacerdes latenigra Pic, 1922, Asclerosibutia lineaticollis var. rufipennis Blair, 1926, Asclerosibutia reducta Pic, 1930, Asclerosibutia divisa Pic. 1952, Asclerosibutia lineaticollis var. ealensis Pic, 1952 (all synonyms of Asclerosibutia lineaticollis Pic, 1914). and Asclerosibutia abdominalis Blair, 1926 (= Asclerosibutia marshalli Blair, 1926). Keys to the genera and species are given. All descriptions are accompanied by male and female genital illustrations. Lectotypes for the following species are designated: Monosigynes rufithorax Pic, 1958, Asclerosibutia violacea Blair, 1926, and Asclerosibutia lineaticollis var. rufipennis Blair. 1926. [source] Act and Event in Rahner and von Balthasar: A Case Study in Catholic Systematics,NEW BLACKFRIARS, Issue 1019 2008Cyrus P. Olsen Abstract In Benedict XVI's God is Love§1 humanity's relationship with God is described as an encounter with an ,event', the Christ-event. I argue that this shift in Catholic theology towards language of act and event signalled a relationship to existentialism, which, taken broadly, entailed an emphasis upon subjectivity and freedom uncharacteristic of the focus upon objectivity common to neo-Scholastic thought. As we shall see with Karl Rahner and Hans Urs von Balthasar, however, this newfound emphasis on subjectivity and freedom did not necessitate an abandonment of all the elements of the more ,objective' perspective. Rather, the task during the ascendancy of existentialist thought became the integration of human subjectivity with the objective and independent reality of the world and God. I suggest that Rahner and von Balthasar use notions such as act and event as a way of being mindful of the role of the subject's creativity and freedom in its encounter with the world, God, and other persons, without thereby undermining the freedom and creativity of that which is other than the subject. [source] Systematics of Chaetocerotaceae (Bacillariophyceae).PHYCOLOGICAL RESEARCH, Issue 2 2003SUMMARY In order to construct a model of evolutionary relationships within the diatom family Chaetocerotaceae, 37 species of Chaetoceros Ehrenberg, representing all subgenera and 21 of 22 subgeneric sections of the genus, plus three Bacteriastrum Shadbolt species, representing both of its subgeneric sections, were subjected to cladistic analysis. One species each of Eucampia Ehrenberg, Cerataulina Peragallo, Hemiau-lus Ehrenberg, Attheya West and Gonioceros H. & M. Peragallo were used as outgroups. A matrix of 65 binary and multistate morphological characters was constructed, with data being gathered from original observation of material in the light and electron microscopes, and from the published literature. The analysis yielded 36 most-parsimonious cladograms of 316 steps; incongruence between trees is largely restricted to some taxa representing undersampled sections of Chaetoceros subg. Hyalochaete. The robustness of this hypothesis was examined in several ways. To assess the effect of character weighting, the bootstrap was used to randomly weight characters. The parsimony criterion was relaxed via a decay index, and finally, the tree length was compared to that of trees randomly generated from the data matrix. The majority of investigated species of Chaetoceros subg. Phaeoceros, Chaetoceros subg. Hyalochaete and Bacteriastrum appear to belong to a continuous grade, rather than comprising individual clades. Chaetoceros is paraphyletic. Thus, the traditional classification does not accurately reflect the hypothesized phylogenetic relationships of this family. [source] The Triune God: Systematics , By Bernard LonerganRELIGIOUS STUDIES REVIEW, Issue 1 2008Don Schweitzer No abstract is available for this article. [source] Mineral Paragenesis, Fluid Inclusions and Sulfur Isotope Systematics of the Lepanto Far Southeast Porphyry Cu-Au Deposit, Mankayan, PhilippinesRESOURCE GEOLOGY, Issue 3 2000Akira IMAI Abstract: The Lepanto Far Southeast porphyry Cu-Au deposit is located beneath and to the southeast of the Lepanto enargite-luzonite Cu,Au deposit in Mankayan, Benguet Province, Philippines. The principal orebody consists of potassic alteration subjected to partial retrograde chlorite alteration that rims stock-work of quartz-anhydrite veinlets. Fluid inclusions found in stockwork quartz and anhydrite in the biotitized orebody center are dominated by polyphase inclusions that homogenize at temperatures of >500C. Sulfur isotopic thermometry applied to the sulfides-anhydrite pairs suggests around 500C. The principal ore minerals associated with quartz-anhydrite stockworks are chalcopyrite and pyrite with minor bornite and Bi,Te,bearing tennantite, with trace of native gold. Rounded pyrite grains appear fractured and corroded and are interpreted as remnants of primary intermediate solid solution + pyrite assemblage. A breccia pipe truncates the deposit. Mineralization in the breccia pipe is brought by quartz-anhydrite veinlets and infilling in the interstices between clasts. Chalcopyrite-Au mineralization associated with molybdenite is recognized in the deeper zone in the breccia pipe. Fluid inclusion microthermometry on polyphase inclusions in veinlet quartz as well as sulfur isotope thermometry applied for the pair of anhydrite and sulfides suggests >450C. Fluid inclusions in veinlet quartz and anhydrite in the fringe advanced argillic alteration are chiefly composed of coexisting liquid-rich inclusions and gas-rich inclusions, in addition to coexisting polyphase inclusions and gas-rich inclusions. These inclusions exhibit a wide range of homogenization temperatures, suggesting heterogeneous entrapping in the two-fluid unmixing region. Sulfur isotopes of aqueous sulfide and sulfate exhibit a general trend from the smallest fractionation pairs (about 11%) in the biotitized orebody center to the largest fractionation (about 25%) pairs in the fringe advanced argillic alteration, suggesting a simple evolution of hydrothermal system. The slopes of arbitrary regression lines in ,34S versus 34S[SO4 = ,H2S] diagram suggest that the abundance ratio of aqueous sulfate to sulfide in the hydrothermal fluid has been broadly constant at about 1:3 through temperature decrease. The intersection of these two regression lines at the ,34S axis indicates that the bulk ,34S is about +6%. Thus, the Lepanto FSE deposit is a further example which confirms enrichment in 34S in the hydrous intermediate to silicic magmas and associated magmatic hydrothermal deposits in the western Luzon arc. [source] Systematics and phylogeography of a threatened tortoise, the speckled padloperANIMAL CONSERVATION, Issue 3 2010S. R. Daniels Abstract This study investigated the systematics and phylogeography of a threatened tortoise of South Africa, the speckled padloper Homopus signatus. Sixty three specimens were collected from 17 localities that covered the distributional range of the two subspecies in western South Africa and a north-eastern population that was recently discovered near Pofadder. The Pofadder sample could not be assigned to either subspecies based on morphology. The samples were sequenced for two partial mtDNA fragments, nicotinamide adenine dinucleotide dehydrogenase component four and cytochrome b, which yielded ,1.1 kb, while a subset of the samples were sequenced for a 390 bp nuclear DNA (nDNA) fragment of prolactin. Phylogenetic analyses of mtDNA using minimum evolution, maximum parsimony and Bayesian inferences supported the monophyly of H. signatus and revealed that the Pofadder specimen was basal in the topology and sister to the remainder. The phylogenetic analyses did not support the recognition of two subspecies; there was statistical support for a Homopus signatus signatus clade but Homopus signatus cafer was not monophyletic. The nDNA analysis showed no difference between the subspecies and placed the Pofadder sample distant but not distinct from H. s. signatus. The mtDNA and the nDNA data suggest that the subspecies are invalid taxonomic units. The structure of the mtDNA network corresponded to the geographical distribution of populations. The north-western populations formed one haplocluster, corresponding to H. s. signatus, whereas the south-western populations formed three haploclusters, corresponding to H. s. cafer. The Pofadder sample was unconnected to the network. The morphology of the northern and southern morphotypes probably reflects selection for crypsis on the different substrate types of the regions, granites and sedimentary rocks, respectively. These results highlight that subspecies designations should be authenticated by molecular techniques because morphological plasticity can obfuscate phylogenetic relationships. We consider the western H. signatus populations as one taxonomic unit and recommend wider sampling of the Pofadder locality to clarify the taxonomic status of this lineage. [source] Systematics of the soft X-ray characteristics of various stellar types in RASS dataASTRONOMISCHE NACHRICHTEN, Issue 1-2 2003E.J.A. Meurs No abstract is available for this article. [source] Systematics and biology of the Australian species of Balaustium von Heyden (Acari: Erythraeidae)AUSTRALIAN JOURNAL OF ENTOMOLOGY, Issue 4 2001RB Halliday Abstract The presence of Balaustium medicagoense Meyer and Ryke (Erythraeidae) in Australia is confirmed. Australian specimens previously identified as Balaustium murorum (Hermann) are shown to belong to B. medicagoense Balaustium medicagoense has been regarded as a beneficial predator in southern Australia, but it can also be a significant pest of cereal crops. It appears to have been introduced into Australia from South Africa. Some species previously described as Balaustium are transferred to Abrolophus, as Abrolophus newmani (Womersley) comb. n., Abrolophus glauerti (Womersley) comb. n., Abrolophus littorale (Womersley) comb. n., Abrolophus brevum (Womersley) comb. n., and Abrolophus ripicola (Womersley) comb. n. [source] Systematics and evolutionary relationships of the mountain lizard Liolaemus monticola (Liolaemini): how morphological and molecular evidence contributes to reveal hidden species diversityBIOLOGICAL JOURNAL OF THE LINNEAN SOCIETY, Issue 3 2009FERNANDO TORRES-PÉREZ The delimitation of species is a major issue in systematic biology and has been a re-emerging discipline in the last decade. A number of studies have shown that the use of multiple data sets is critical for the identification of cryptic species, particularly in groups with complex evolutionary histories. Liolaemus monticola is a montane lizard species distributed in central Chile (32°,42°S), with four described subspecies in a latitudinal gradient from north to south: L. m. monticola, L. m. chillanensis, L. monticola ssp. and L. m. villaricensis. In order to test the systematic status and phylogenetic relationships of the taxa included in the L. monticola group, we analysed morphological (morphometric and meristic) and molecular (allozyme and mitochondrial DNA) data sets. The results of the morphological analyses showed that meristic variables correctly assigned individuals with higher accuracy than did morphometric characters. The results of the analyses of allozyme data revealed eight diagnostic loci that are evidence for significant differences among the four L. monticola subspecies. Phylogenetic analyses with mitochondrial DNA data, including additional species, showed that the L. monticola group is polyphyletic. We postulate that the four current subspecies represent independent evolutionary lineages and must be raised to the specific level as L. monticola, L. chillanensis and L. villaricensis. The taxonomic status of the unnamed L. monticola ssp. remains unresolved, although we provide a preliminary proposal. © 2009 The Linnean Society of London, Biological Journal of the Linnean Society, 2009, 96, 635,650. [source] Molecular systematics of cowries (Gastropoda: Cypraeidae) and diversification patterns in the tropicsBIOLOGICAL JOURNAL OF THE LINNEAN SOCIETY, Issue 3 2003CHRISTOPHER P. MEYER This study produces a nearly comprehensive phylogeny for the marine gastropod group Cypraeidae (cowries) and uses this topology to examine diversification patterns in the tropics. The dataset is based on molecular sequence data from two mitochondrial genes and includes 210 evolutionary significant units (ESUs) from 170 recognized species (>80%). Systematics for the group is revised based on well-supported clades, and tree topology is generally consistent with previously proposed classification schemes. Three new genera are introduced (Cryptocypraea gen. nov, Palmulacypraea gen. nov, and Contradusta gen. nov) and two previous genera are resurrected (Perisserosa and Eclogavena). One new tribe is proposed (Bistolidini). Topologies produced by a range of transition:transversion (Ti:Tv) weighting schemes in parsimony are pooled and evaluated using maximum likelihood criteria. Extensive geographical coverage shows persistent, large-scale geographical structure in sister-groups. Genetic divergence between subspecies is often equivalent or even greater than that between recognized species. Using ESUs as a metric, diversity throughout the Indo-West Pacific (IWP) increases by 38%. Intra- and inter-regional diversification patterns show that the IWP is the centre for speciation in cowries. The other major tropical regions of the world are inhabited by a predominantly relictual fauna; from a cowrie's eye-view. Good dispersal ability begets larger ranges, increased extinction resistance and morphological stasis; whereas shorter larval duration results in smaller ranges, higher speciation rates, but also higher turnover. Larval duration and dispersal ability appear correlated with ocean productivity as taxa with longer-lived larvae are associated with oligotrophic conditions; whereas taxa with shorter larval durations are associated with eutrophic, continental conditions. This tendency is carried to the extreme in temperate or upwelling regions where a planktonic phase is completely lost and crawl-away larvae evolve multiple times. A strong phylogenetic trend supports these observations as lineages leading up to and including the derived Indo-West Pacific Erroneinae clade contain taxa predominantly restricted to continental habitats and have undergone the greatest evolutionary radiations in their respective regions. © 2003 The Linnean Society of London, Biological Journal of the Linnean Society, 2003, 79, 401, 459. [source] Systematics of the Nasa poissoniana group (Loasaceae) from Andean South AmericaBOTANICAL JOURNAL OF THE LINNEAN SOCIETY, Issue 3 2009TILO HENNING The monophyletic Nasa poissoniana group (Loasaceae, subfamily Loasoideae) is revised on the basis of extensive field studies, observations in cultivation and the revision of herbarium specimens. A core of taxa has been considered as closely allied on the basis of morphology in the past, but several additional taxa have been recognized as allied to this group on the basis of molecular data. One species, N. raimondii, formerly placed in the N. stuebeliana group, is now transferred to the N. poissoniana complex as a result of the molecular findings. Nasa poissoniana ssp. glandulifera is described as new to science on the basis of morphologically divergent collections of N. poissoniana from two northern departments of Peru. Numerous new localities, often far from the previously known distribution area, are reported for several species. The N. poissoniana group has its centre of diversity in the inner Andean valleys of southern Peru, quite unlike all other groups of Nasa, with centres of diversity in the Amotape,Huancabamba Zone in northern Peru and southern Ecuador. Descriptions, drawings and a distribution map and key to all taxa are provided, and detailed information on habitat and distribution is given for each species. © 2009 The Linnean Society of London, Botanical Journal of the Linnean Society, 2009, 161, 278,301. [source] Systematics and biogeography of Klasea (Asteraceae,Cardueae) and a synopsis of the genusBOTANICAL JOURNAL OF THE LINNEAN SOCIETY, Issue 4 2006LUDWIG MARTINS Klasea, traditionally treated as a section in Serratula, is now widely accepted at the generic level. A classification of the genus is presented here, accommodating the 46 species in ten sections based on nuclear ribosomal DNA external and internal transcribed spacer sequence data and morphology. New combinations for five species and ten subspecies are published, and a new hybrid species is described. The genus ranges from the Iberian Peninsula and north Africa through southern and eastern Europe, west and central Asia to the Himalayas, and the Far East of Russia and China. The ancestral area is in west Asia, most probably eastern Anatolia and northern and western Iran. In this region, representatives of all sections are present. The largest section Klasea diversified most likely in the mountains of central Asia. A key to all Klasea species is provided. © 2006 The Linnean Society of London, Botanical Journal of the Linnean Society, 2006, 152, 435,464. [source] Phylogenetic Species, Nested Hierarchies, and Character FixationCLADISTICS, Issue 4 2000Paul Z. Goldstein Cladistic mechanics and ramifications of various species concepts rooted in phylogenetic theory are explored. Published discussions of the phylogenetic species concept (PSC) have been hampered by persistent misconceptions surrounding its ontology and applicability, and by confusion of various incompatible versions of species concepts claiming to follow from Hennig's (1966), Phylogenetic Systematics, Univ. of Illinois Press, Urbana work. Especially problematic are topology- or tree-based versions of species diagnosis, which render diagnoses dependent on relationships depicted as hierarchically structured regardless of any lack of underlying hierarchy. Because the applicability of concepts such as monophyly, paraphyly, and polyphyly rests ultimately on the underlying hierarchical distribution of characters, representations of tokogenetic or reticulating systems as nested hierarchies are necessarily inaccurate. And since hierarchical representations,even if accurate,of nonrecombining genetic elements need not coincide with the organisms that bear them, tree-based diagnoses are further hampered, except potentially as retrospective tools. The relationship between tree-based species delineations and the criterion of character fixation is explored. Fixation of characters by which one identifies phylogenetic species is further distinguished from the fixation of character state differences, and the implications of that distinction are explored with reference to the interpretation of speciation events. It is demonstrated that character fixation in alternative species need not coincide with the achievement of reciprocal monophyly. While the PSC retains shortcomings, some of the more frequently criticized aspects of the PSC are functions of sampling that are no more problematic than for any basic systematic endeavor. [source] On the Other "Phylogenetic Systematics"CLADISTICS, Issue 3 2000Kevin C. Nixon De Queiroz and Gauthier, in a serial paper, argue that biological taxonomy is in a sad state, because taxonomists harbor "widely held belief" systems that are archaic and insufficient for modern classification, and that the bulk of practicing taxonomists are essentialists. Their paper argues for the scrapping of the current system of nomenclature, but fails to provide specific rules for the new "Phylogenetic Systematics",instead we have been presented with a vague and sketchy manifesto based upon the assertion that "clades are individuals" and therefore must be pointed at with proper names, rather than diagnosed by synapomorphies. They claim greater stability for "node pointing," yet even their own examples show that the opposite is true, and their node pointing system is only more stable in a purely metaphysical sense detached from characters, evidence, usage of names, and composition of groups. We will show that the node pointing system is actually far LESS stable than the existing Linnaean System when stability is measured by the rational method of determining the net change in taxa (species) included in a particular group under different classifications. [source] Systematics and Biogeography of Hard Ticks, a Total Evidence ApproachCLADISTICS, Issue 1 2000J.S.H. Klompen Systematic relationships among the basal Ixodidae are examined using one morphological and three molecular data sets, 18S and 28S nuclear and 16S mitochondrial rDNA. Although different combinations of partitions are incompatible in a partition homogeneity test, combining them produces similar or better support for most major lineages through both additive and complementary effects. The different data sets are not complete for all taxa, but inclusion or exclusion of taxa with missing data for one or more data sets (8 of 29 ingroup taxa) does not influence overall tree topology and only weakly affects support levels. The only notable effect was based on gap treatment in the 28S data set. Gap treatment completely changes the arrangement and support levels for one basal node. The combined analyses show strong support for the Metastriata, a lineage including most endemic Australian Ixodes, and a lineage including the remaining Ixodes, but not for the Prostriata (=Ixodes s.l.). The distribution pattern of endemic Australian taxa (nearly all included in three exclusively Australian basal lineages) suggests that these lineages, and by extension the Ixodidae, originated after the isolation of Australia in the late Cretaceous, much more recently than previously indicated. [source] A new platysomid from the Upper Carboniferous of Kansas (USA) and remarks on the systematics of deep-bodied lower actinopterygiansACTA ZOOLOGICA, Issue 2009Kathryn E. Mickle Abstract Scales and presumptive tooth plates from deep-bodied platysomid actinopterygians have previously been reported from the Lower Permian of Kansas (USA), but until now, an articulated specimen from this state has not been recovered. The first articulated deep-bodied platysomid fish from the Upper Carboniferous of Kansas is described herein. This specimen is compared to other known species from the genus Platysomus, with special attention paid to the North American taxa. The cranial osteology of this new specimen is closest to Platysomus schultzei from the Upper Carboniferous of New Mexico. The newly described platysomid species highlights the need for a revision of the fish included within the genus Platysomus. [source] Searching for new morphological characters in the systematics of scleractinian reef corals: comparison of septal teeth and granules between Atlantic and Pacific MussidaeACTA ZOOLOGICA, Issue 2 2009Ann F. Budd Abstract Recent molecular analyses have challenged the traditional classification of scleractinian corals at all taxonomic levels suggesting that new morphological characters are needed. Here we tackle this problem for the family Mussidae, which is polyphyletic. Most of its members belong to two molecular clades composed of: (1) Atlantic Mussidae and Faviidae (except Montastraea) and (2) Pacific Mussidae (Cynarina, Lobophyllia, Scolymia, Symphyllia) and Pectiniidae. Other Pacific mussids (e.g. Acanthastrea) belong to additional clades. To discover new characters that would better serve as phylogenetic markers, we compare the skeletal morphology of mussid genera in different molecular-based clades. Three sets of characters are considered: (1) macromorphology (budding; colony form; size and shape of corallites; numbers of septal cycles), (2) micromorphology (shapes and distributions of septal teeth and granules), and (3) microstructure (arrangement of calcification centres and thickening deposits within costosepta). Although most traditional macromorphological characters exhibit homoplasy, several new micromorphological characters are effective at distinguishing clades, including the shapes and distribution of septal teeth and granules, the area between teeth, and the development of thickening deposits. Arrangements of calcification centres and fibres differ among clades, but the fine-scale structure of thickening deposits does not. [source] Anatomy and systematics of the minute syrnolopsine gastropods from Lake Tanganyika (Caenogastropoda, Cerithioidea, Paludomidae)ACTA ZOOLOGICA, Issue 4 2008Ellen E. Strong Abstract The minute syrnolopsine gastropods endemic to Lake Tanganyika have been allied to a number of freshwater, marine and terrestrial groups as a consequence of superficial conchological similarity. Although early anatomical studies confirmed the cerithioid organization of this clade, their close relationship to other lake species was not consistently recognized. In several recent cladistic analyses based on molecular data, the higher taxonomic placement and sister group relationships of syrnolopsines have been unstable. The present analysis confirms that syrnolopsines possess a spermatophore-forming organ , a synapomorphy of the Paludomidae , corroborating their placement in this family. Consistent with the molecular data, syrnolopsine monophyly is supported by two characters that occur exclusively in this group (salivary gland ducts that bypass the nerve ring and a linear albumen gland). Several characters in Martelia tanganyicensis, the most diminutive syrnolopsine , are only evident in the smallest lake species thus far investigated (Bridouxia, Stormsia) namely reduction of ctenidial leaflets, sorting area, intestine length and number of statoconia. These features are interpreted as being correlated with reduction in size. Nevertheless, close examination reveals differences in detail that allow more refined hypotheses of homology and are consistent with their independent origin. [source] Fungivore host-use groups from cluster analysis: patterns of utilisation of fungal fruiting bodies by ciid beetlesECOLOGICAL ENTOMOLOGY, Issue 6 2005Glenda M. Orledge Abstract., 1.,Ciid beetles typically live and breed in the fruiting bodies of lignicolous basidiomycete fungi. This study was undertaken to address the lack of an objective examination of patterns of host use by ciids. 2.,Cluster analysis of ciid host-use datasets from Britain, Germany, North America, and Japan, and subsequent cross-dataset comparisons, demonstrated the existence of ciid host-use patterns of wide geographical occurrence. These patterns were formalised as ciid host-use groups. 3.,Six Holarctic ciid host-use groups, and two host-use subgroups, were identified, and are described. Each host-use group comprises an assemblage of fungal genera and the breeding ciids that it supports. Each taxon belongs to only a single host-use group, but may be associated with several members of that group. There is a strong tendency for closely related taxa to belong to the same host-use group. 4.,It is suggested that ciid host-use groups are defined ultimately by host chemistry, with the ciids that belong to a particular group recognising, and responding positively to, emitted volatiles characterising the fungi belonging to that group. 5.,The idea of the host-use group bears comparison with the concepts of niche and guild, but is not equivalent to either. 6.,Ciid host-use groups have a valuable role to play in underpinning future studies of ciid ecology, also the systematics of both ciids and their fungal hosts. [source] IS A NEW AND GENERAL THEORY OF MOLECULAR SYSTEMATICS EMERGING?EVOLUTION, Issue 1 2009Scott V. Edwards The advent and maturation of algorithms for estimating species trees,phylogenetic trees that allow gene tree heterogeneity and whose tips represent lineages, populations and species, as opposed to genes,represent an exciting confluence of phylogenetics, phylogeography, and population genetics, and ushers in a new generation of concepts and challenges for the molecular systematist. In this essay I argue that to better deal with the large multilocus datasets brought on by phylogenomics, and to better align the fields of phylogeography and phylogenetics, we should embrace the primacy of species trees, not only as a new and useful practical tool for systematics, but also as a long-standing conceptual goal of systematics that, largely due to the lack of appropriate computational tools, has been eclipsed in the past few decades. I suggest that phylogenies as gene trees are a "local optimum" for systematics, and review recent advances that will bring us to the broader optimum inherent in species trees. In addition to adopting new methods of phylogenetic analysis (and ideally reserving the term "phylogeny" for species trees rather than gene trees), the new paradigm suggests shifts in a number of practices, such as sampling data to maximize not only the number of accumulated sites but also the number of independently segregating genes; routinely using coalescent or other models in computer simulations to allow gene tree heterogeneity; and understanding better the role of concatenation in influencing topologies and confidence in phylogenies. By building on the foundation laid by concepts of gene trees and coalescent theory, and by taking cues from recent trends in multilocus phylogeography, molecular systematics stands to be enriched. Many of the challenges and lessons learned for estimating gene trees will carry over to the challenge of estimating species trees, although adopting the species tree paradigm will clarify many issues (such as the nature of polytomies and the star tree paradox), raise conceptually new challenges, or provide new answers to old questions. [source] | ||||||||||||||||||||||||||||||||||||||||||||||