Synonym

Distribution by Scientific Domains
Distribution within Life Sciences

Kinds of Synonym

  • junior synonym
  • subjective synonym


  • Selected Abstracts


    RESURRECTION OF MESOPLODON TRAVERSII (GRAY, 1874), SENIOR SYNONYM OF M. BAHAMONDI REYES, VAN WAEREBEEK, CÁRDENAS AND YÁÑEZ, 1995 (CETACEA: ZIPHIIDAE)

    MARINE MAMMAL SCIENCE, Issue 3 2002
    Anton L. van Helden
    Abstract Mesoplodon traversii (Gray, 1874) is shown to be a senior synonym of the recently described beaked whale Mesoplodon hahamondi Reyes et al., 1995 on the basis of a phylogenetic analysis of mitochondrial DNA control region sequences. The mandible and teeth of M. traversii, first reported in 1873 by Hector as Dolichodon layardii. are redescribed. The species can be distinguished by features of the calvaria; including the large jugal, broad rostrum, and small distance between premaxillary foramina. The male teeth, which are large and spade-shaped with a strong terminal denticle, are also diagnostic. M. traversii is known only from Pitt Island and White Island, New Zealand and Robinson Crusoe Island, Juan Fernandez Archipelago, Chile. [source]


    A new look at Kedarnatha P.K.Mukh.

    FEDDES REPERTORIUM, Issue 3-4 2004
    & Constance (Umbelliferae)
    A field investigation of Kedarnatha sanctuarii P.K.Mukh. & Constance in its locus classicus and subsequent carpological analysis show the identity of the species and Trachydium garhwalicum H.Wolff, described from the same region of Tehri Garhwal (Uttaranchal, India). As this species differs in some essential characters from Trachydium, Trachyspermum, Chamaesium, Chamaesciadium, Schulzia, the genus Kedarnatha must be retained, and IndoschulziaPimenov & Kljuykov is to be considered as its latter synonym. A new nomenclatural combination, K. garhwalica (H.Wolff) Pimenov & Kljuykov comb. nov. have been proposed to name the type species. Cortia oreomyrrhiformisFarille & Malla and Trachydium hamelianumFarille & Malla are transferred to Kedarnatha [K. oreomyrrhiformis (Farille & Malla) Pimenov & Kljuykov comb. nov. and K. hameliana (Farille & Malla) Pimenov & Kljuykov comb. nov.]. Two new additional species, K. meifoliaPimenov & Kljuykov sp. nova from India and K. vaginataPimenov & Kljuykov sp. nova from Burma, are described. As a result, the genus contains five species, distributed from Himachal Pradesh (India) to Burma. (© 2004 WILEY-VCH Verlag GmbH & Co. KGaA, Weinheim) Eine Neufassung von KedarnathaP.K.Mukh. & Constance (Umbelliferae) Eine Feldstudie an Kedarnatha sanctuarii P.K.Mukh. & Constance am locus classicus und die anschließende karpologische Analyse erbrachten die Identität dieser Art mit TrachydiumgarhwalicumH.Wolff, das ebenfalls aus der Region Tehri Garhwal (Uttaranchal, Indien) beschrieben wurde. Da diese Art sich in einigen wesentlichen Merkmalen von Trachydium, Trachyspermum, Chamaesium, Chamaesciadium und Schulzia unterscheidet, muss die Gattung Kedarantha beibehalten werden. Die Gattung IndoschulziaPimenov & Kljuykov ist somit ein späteres Synonym. Die neue nomenklatorische Kombination, K. garhwalica(H.Wolff) Pimenov & Kljuykov comb. nov. wird als Typusart vorgeschlagen. Cortia oreomyrrhiformisFarille & Malla und Trachydium hamelianumFarille & Malla werden in die Gattung Kedarnatha überführt [K. oreomyrrhiformis (Farille & Malla) Pimenov & Kljuykov comb. nov. und K. hameliana(Farille & Malla) Pimenov & Kljuykov comb. nov.]. Zwei weitere Arten werden neu beschrieben: K. meifoliaPimenov & Kljuykov sp. nova aus Indien und K. vaginataPimenov & Kljuykov sp. nova aus Burma. Damit umfasst die Gattung fünf Arten, die von Himachal Pradesh (Indien) bis Burma verbreitet sind. [source]


    Placebo psychotherapy: Synonym or oxymoron?

    JOURNAL OF CLINICAL PSYCHOLOGY, Issue 7 2005
    Irving Kirsch
    Contrary to some recent claims, the placebo effect is real and in some cases very substantial. Placebo effects can be produced or enhanced by classical conditioning, but consistent with virtually all contemporary conditioning theories, these effects are generally mediated by expectancy. Expectancy can also produce placebo effects that are inconsistent with conditioning history. Although expectancy also plays an important role in psychotherapy outcome, the logic of placebo-controlled trials does not map well onto psychotherapy research. The idea of evaluating the efficacy of psychotherapy by controlling for nonspecific or placebo factors is based on a flawed analogy and should be abandoned. © 2005 Wiley Periodicals, Inc. J Clin Psychol 61: 791,803, 2005. [source]


    Fagus sylvatica forest vegetation in Greece: Syntaxonomy and gradient analysis

    JOURNAL OF VEGETATION SCIENCE, Issue 1 2001
    Erwin Bergmeier
    Abstract. This is the first comprehensive study on Fagus sylvatica s.l. forest communities in Greece. It is based on the multivariate analysis of more than 1100 relevés throughout the distribution area of Fagus in Greece, 60 % of which have been sampled within the last 15 years. The data were compiled from all relevant literature and from unpublished field studies. Prior to data analysis, taxonomic and floristic inaccuracies in older literature have been corrected, and some taxa aggregated. Status and distribution of Fagus in Greece, as well as historical and silvicultural aspects, are outlined. Classification resulted in 16 vegetation types defined by species composition, of which 10 units are ranked as association (of which two were described as new) or community, with six subcommunities included. Synonyms from the literature are given to all types. The floristic structure of the data set is displayed by a synoptic table and by DCA ordination. Community differentiation follows a complex edaphic-climatic-phytogeographic pattern. Apart from Fagus sylvatica ssp. orientalis forests of warm and dry habitats and low to medium altitudes which are related to deciduous Quercus forests, four Fagion community groups are distinguished which represent different habitat types: (1) cool, humid, mesotrophic; (2) cool, acidic; (3) high altitude, calcareous; (4) medium altitude, moderately warm and dry. The communities may be assigned to three regional suballiances, viz. (a) Doronico orientalis-Fagenion at medium altitudes of North-Central and East-Central Greece; (b) Doronico columnae-Fagenion in the far north and at high altitudes southward to East-Central Greece; (c) Geranio striati-Fagenion in the Northwest. Various types of thermophilous beech forest are assigned to (a) and (c), acidic forest to (b), and mesic Fagus forest to (b) and (c). [source]


    Phenotypic plasticity, polymorphism and phylogeny within placoderms

    ACTA ZOOLOGICA, Issue 2009
    K. Trinajstic
    Abstract Intraspecies variation, polymorphism and asymmetric traits are observed within two families of Arthrodira, the Incisoscutidae and Camuropiscidae, from the Gogo Formation in northern Western Australia. Individual plates of the head and trunk shield show considerable variation between individuals. Plates that show the greatest degree polymorphic traits are the rostral (R), marginal (M), submarginal (SM), preorbital (PrO), anterior dorsolateral, anterior median ventral (AMV) and posterior ventrolateral (PVL) plates. The paths of the sensory line canals are the most variable feature and the dermal plates of the cheek show the greatest asymmetry. It is apparent that if anatomical data in arthrodires are to be interpreted with greater precision, detailed knowledge of intraspecies variation, polymorphic and asymmetric traits is essential. How these variables are treated in cladistic analysis is also critical. Here multistate characters were coded differently in five discrete analyses, each analysis yielding a different number of trees and relationships. It was concluded that including and coding for multistate characters gave the most robust tree. In addition, further morphological characters from a new specimen of Gogosteus sarahae Long (1994) indicates many of the characters used to separate this genus from Incisoscutum are inconsistent and so it is here considered that the genus Gogosteus is a junior synonym of Incisoscutum. [source]


    A redescription of the Early Carboniferous acanthodian Acanthodes lopatini Rohon, 1889

    ACTA ZOOLOGICA, Issue 2009
    Pavel Beznosov
    Abstract Investigation of numerous well-preserved specimens of Early Carboniferous acanthodians collected over recent decades from southern central Siberia allowed their redescription as Acanthodes lopatini Rohon, 1889. The morphological characteristics supporting this classification, some peculiarities of ontogeny and a new reconstruction of this species are given. The genus Carycinacanthus Miles 1966 is regarded as a junior synonym of Acanthodes. Homalacanthus bergi (Obruchev 1962) is regarded as a junior synonym of A. lopatini. [source]


    Taxonomic Study of Korean Oedipodinae (Orthoptera: Caelifera: Acrididae)

    ENTOMOLOGICAL RESEARCH, Issue 2 2005
    Tae-Woo KIM
    ABSTRACT As a result of Korean Oedipodinae is taxonomically revised, eleven species are recognized and Microgastrimargus taeguensis Lee and Park 1992 is a new synonym of Oedaleus infernalis Saussure 1884. A key to Korean species of Oedipodinae, and description of a little known species Epacromius japonicus (Shiraki 1910) are provided. [source]


    Section Neurocentrae Bunge from genus Cousinia Cass. (Asteraceae) in Iran

    FEDDES REPERTORIUM, Issue 1-2 2009
    S. B. Djavadi
    This paper is based on experience from field studies, herbarium materials and images of type specimens belonging to sect. Neurocentrae Bunge from genus Cousinia Cass. Section Leucocaulon Tscherneva is considered as synonym of sect. Neurocentrae Bunge, according to the morphological characters of C. turcomanica C.Winkl. (the type specimen of sect. Leucocaulon) and close affinities that shows with the species of sect. Neurocentrae. Section Ramosissimae is described as a section new to science. Differences between sect. Neurocentrae and sect. Ramosissimae are discussed and a key to the species of the two sections are provided separately. Besides, C. deserti var. longispinosa is described as a new member for the sect. Neurocentrae Bunge. (© 2009 WILEY-VCH Verlag GmbH & Co. KGaA, Weinheim) Die Sektion Neurocentrae Bunge der Gattung Cousinia Cass. (Asteraceae) in Iran Diese Arbeit basiert auf Erfahrungen aus Geländearbeit, Studien an Herbar-Material und Untersuchungen an Typus-Material der Sektion Neurocentrae Bunge der Gattung Cousinia Cass. Die Sektion Leucocaulon Tscherneva wird als Synonym zur Sekt. Neurocentrae betrachtet und zwar auf Grund der morphologischen Merkmale von C. turcomania C.Winkl. (Typus-Art der Sekt. Leucocaulon) und der engen Verwandtschaft dieser Art zur Sektion Neurocentrae. Die Sekt. Ramosissimae wird hier als neue Sektion beschrieben. Die Unterschiede zwischen der Sekt. Neurocentrae und der Sekt. Ramosissimae werden betrachtet und ein Schlüssel für beide Sektionen beigefügt. Außerdem wird Cousinia deserti var. longispinosa als neue Art in der Sektion Neurocentrae beschrieben. [source]


    Rediscovery of Achillea boissieriHausskn. ex Boiss. later 140 years

    FEDDES REPERTORIUM, Issue 7-8 2006
    T. Arabac
    This study includes taxonomical improvements of Achillea boissieriHausskn. ex Boiss. and distributions in Turkey which was known only from the type specimen and regarded as a synonym in relevant studies. After collected second times from the field, A. boissieri decided as a separate species. Furthermore, description, localities, distribution map and key of allied species are given. (© 2006 WILEY-VCH Verlag GmbH & Co. KGaA, Weinheim) Die Wiederentdeckung von Achillea boissieriHausskn. ex Boiss. nach 140 Jahren Diese Arbeit beschreibt die taxonomische Entwicklung und die Verbreitung von Achillea boissieriHausskn. ex Boiss. in der Türkei, das nur durch den Typus-Beleg bekannt ist und in den diesbezüglichen Untersuchungen synonym geführt wird. Nachdem es während Felduntersuchungen ein zweites Mal gefunden wurde, war festzustellen, dass A. boissieri als getrennte Art bestehen bleiben muss. Ferner werden Beschreibung, Fundorte, Verbreitungskarte und ein Unterscheidungs-Schlüssel zu verwandten Arten vorgelegt. [source]


    A new look at Kedarnatha P.K.Mukh.

    FEDDES REPERTORIUM, Issue 3-4 2004
    & Constance (Umbelliferae)
    A field investigation of Kedarnatha sanctuarii P.K.Mukh. & Constance in its locus classicus and subsequent carpological analysis show the identity of the species and Trachydium garhwalicum H.Wolff, described from the same region of Tehri Garhwal (Uttaranchal, India). As this species differs in some essential characters from Trachydium, Trachyspermum, Chamaesium, Chamaesciadium, Schulzia, the genus Kedarnatha must be retained, and IndoschulziaPimenov & Kljuykov is to be considered as its latter synonym. A new nomenclatural combination, K. garhwalica (H.Wolff) Pimenov & Kljuykov comb. nov. have been proposed to name the type species. Cortia oreomyrrhiformisFarille & Malla and Trachydium hamelianumFarille & Malla are transferred to Kedarnatha [K. oreomyrrhiformis (Farille & Malla) Pimenov & Kljuykov comb. nov. and K. hameliana (Farille & Malla) Pimenov & Kljuykov comb. nov.]. Two new additional species, K. meifoliaPimenov & Kljuykov sp. nova from India and K. vaginataPimenov & Kljuykov sp. nova from Burma, are described. As a result, the genus contains five species, distributed from Himachal Pradesh (India) to Burma. (© 2004 WILEY-VCH Verlag GmbH & Co. KGaA, Weinheim) Eine Neufassung von KedarnathaP.K.Mukh. & Constance (Umbelliferae) Eine Feldstudie an Kedarnatha sanctuarii P.K.Mukh. & Constance am locus classicus und die anschließende karpologische Analyse erbrachten die Identität dieser Art mit TrachydiumgarhwalicumH.Wolff, das ebenfalls aus der Region Tehri Garhwal (Uttaranchal, Indien) beschrieben wurde. Da diese Art sich in einigen wesentlichen Merkmalen von Trachydium, Trachyspermum, Chamaesium, Chamaesciadium und Schulzia unterscheidet, muss die Gattung Kedarantha beibehalten werden. Die Gattung IndoschulziaPimenov & Kljuykov ist somit ein späteres Synonym. Die neue nomenklatorische Kombination, K. garhwalica(H.Wolff) Pimenov & Kljuykov comb. nov. wird als Typusart vorgeschlagen. Cortia oreomyrrhiformisFarille & Malla und Trachydium hamelianumFarille & Malla werden in die Gattung Kedarnatha überführt [K. oreomyrrhiformis (Farille & Malla) Pimenov & Kljuykov comb. nov. und K. hameliana(Farille & Malla) Pimenov & Kljuykov comb. nov.]. Zwei weitere Arten werden neu beschrieben: K. meifoliaPimenov & Kljuykov sp. nova aus Indien und K. vaginataPimenov & Kljuykov sp. nova aus Burma. Damit umfasst die Gattung fünf Arten, die von Himachal Pradesh (Indien) bis Burma verbreitet sind. [source]


    Alpinites and other Posttornoceratidae (Goniatitida, Famennian)

    FOSSIL RECORD-MITTEILUNGEN AUS DEM MUSEUM FUER NATURKUNDE, Issue 1 2002
    R. Thomas Becker
    Abstract The rediscovery of the supposedly lost type allows a revision of Alpinites Bogoslovskiy, 1971, the most advanced genus of the Posttornoceratidae. The type-species, Alp. kayseri Schindewolf, 1923, is so far only known from the Carnic Alps. Alp. schultzei n. sp. from the eastern Anti-Atlas of Morocco is closely related to Alp. kajraktensis n. sp. (= Alp. kayseri in Bogoslovskiy 1971) from Kazakhstan. A second new and more common species of southern Morocco, Alp. zigzag n. sp., is also known from the Holy Cross Mountains (Poland). The taxonomy and phylogeny of other Posttornoceratidae are discussed. The holotype of Exotornoceras nehdense (Lange, 1929) was recovered and is re-illustrated; it is conspecific with Exot. superstes (Wedekind, 1908). The genus and species is also here first recorded from Morocco. Post. weyeri Korn, 1999 is a subjective synonym of Post. posthumum (Wedekind, 1918) in which strongly biconvex growth lines, as typical for the family, are observed for the first time. Goniatites lenticularis Richter, 1848 is a nomen dubium within Discoclymenia, Clymenia polytrichus in Richter (1948) is a Falcitornoceras. It seems possible to distinguish an extreme thin and trochoid Disco. haueri (Münster, 1840) from the tegoid Disco. cucullata (v. Buch, 1839). Various taxa are excluded from the Posttornoceratidae. Posttornoceras sapiens Korn, 1999 forms the type-species of Maiderocera n. gen., Discoclymenia n. sp. of Müller (1956) is assigned to Maid. muelleri n. sp. Disco. cornwallensis is the type-species of Selwoodites n.gen (Sporadoceratidae) which also occurs in the Rhenish Massive. Durch Auffinden des verschollen geglaubten Typus-Exemplares wird eine Revision der Gattung Alpinites Bogoslovskiy, 1971, der höchst entwickelten Gattung der Posttornoceratidae, möglich. Die Typus-Art, Alp. kayseri Schindewolf, 1923, kommt gesichert bisher nur in den Karnischen Alpen vor. Alp. schultzei n. sp. aus dem östlichen Anti-Atlas (Marokko) ist nah mit Alp. kajraktensis n. sp. aus Kazakhstan (= Alp. kayseri in Bogoslovskiy 1971) verwandt. Eine zweite, aber häufigere neue Art Süd-Marokkos, Alp. zigzag n. sp., wird auch im Heiligkreuzgebirge (Polen) nachgewiesen. Die Taxonomie und Phylogenie anderer Vertreter der Posttornoceratidae werden diskutiert. Der verschollen geglaubte Holotypus von Exotornoceras nehdense (Lange, 1929) wird neu abgebildet und ist conspezifisch mit Exot. superstes (Wedekind, 1908). Die Gattung und Art wird erstmals auch in Marokko nachgewiesen. Post. weyeri Korn, 1999 ist ein subjektives Synonym von Post. posthumum (Wedekind, 1918) bei dem erstmals die Familien-typischen, stark biconvexen Anwachsstreifen beobachtet wurden. Goniatites lenticularis Richter, 1848 ist ein nomen dubium innerhalb von Discoclymenia, Clymenia polytrichus in Richter (1848) ein Falcitornoceras. Es scheint möglich, eine extrem dünne, trochoide Disco. haueri (Münster, 1840) von der tegoiden Disco. cucullata (v. Buch, 1839) zu trennen. Verschiedene Taxa werden aus den Posttornoceratidae ausgeschlossen. Post. sapiens Korn, 1999 wird als Typus-Art zu Maideroceras n. gen. gestellt. Discoclymenia n. sp. in Müller (1956) wird als Maid. muelleri n. sp. neu beschrieben. Disco. cornwallensis Selwood, 1960 bildet die Typusart von Selwoodites n. gen. (Sporadoceratidae), welches auch im Rheinischen Schiefergebirge vorkommt. [source]


    Taxonomy, evolutionary History and Distribution of the middle to late Famennian Wocklumeriina (Ammonoidea, Clymeniida)

    FOSSIL RECORD-MITTEILUNGEN AUS DEM MUSEUM FUER NATURKUNDE, Issue 1 2000
    R. Thomas Becker
    Abstract Old collections, new records, and data from global literature are used for taxonomic revisions and for a new reconstruction of the evolutionary history of the triangularly coiled clymenids, the Wocklumeriaceae, and their ancestors. Epiwocklumeria applanata is first reported from the topmost Wocklum Limestone at Hasselbachtal and Drewer, and this supports the distinction of an applanata Subzone just prior to the global Hangenberg Event which wiped out the whole group. The Wocklumeriaceae and Glatziellidae record of the famous Oberrödinghausen Railway Cut and of other Rhenish sections is revised. The Maïder of Southern Morocco has yielded first Parawocklumeria patens, paprothae, Wocklumeria sphaeroides plana, Kielcensia ingeniens n. sp., and Synwocklumeria mapesi n. sp. Parawocklumeria distributa Czarnocki forms the type-species of Tardewocklumeria n. gen. Lecto- and neotypes for several taxa are designated. The variability and paedomorphic patterns of Wo. sphaeroides are discussed. The Wocklumeriaceae represent the terminal Famennian (Upper Devonian VI-C/D) radiation of a long-ranging lineage which evolved in parallel to other clymenid groups. They are characterized by sutural features and by longidomic and very slowly expanding whorls. The polyphyletic traditional Gonioclymeniina are divided into the suborder Wocklumeriina (with Wocklumeriaceae, Glatziellaceae n.superfam. and Biloclymeniaceae) and into the Gonioclymeniaceae of the Clymeniina. Gyroclymenia Czarnocki is regarded as a junior synonym of Pleuroclymenia Schindewolf which, however, does not include the ,Pleuro.' americana and eurylobica groups. Pleuroclymenia represents the ancestral form of the Wocklumeriina and also the phylogenetical link with Platyclymenia (Varioclymenia) of the Clymeniaceae. The Gonioclymeniaceae had their roots in advanced Platyclymeniidae. Alte Aufsammlungen, Neunachweise und globale Literaturdaten werden für taxonomische Revisionen und für eine neue Rekonstruktion der Evolution der Dreiecksclymenien (Wocklumeriaceae) und ihrer Vorfahren benutzt. Epiwocklumeria applanata wird zum ersten Mal im Hasselbachtal und bei Drewer nachgewiesen, und diese Funde bestätigen die Abtrennung einer applanata -Subzone im unmittelbar Liegenden des Hangenberg-Event, welcher zum Aussterben der gesamten Gruppe führte. Die Verbreitung von Wocklumeriaceae und Glatziellidae im klassischen Profil des Bahneinschnittes bei Oberrödinghausen und an anderen Fundorten im Rheinischen Schiefergebirge wird revidiert. Der Maïder in Süd-Marokko lieferte erstmalig Parawocklumeria patens, paprothae, Wocklumeria sphaeroides plana, Kielcensia ingeniens n. sp. und Synwocklumeria mapesi n. sp. Parawocklumeria distributa Czarnocki bildet die Typus-Art von Tardewocklumeria n.gen. Weiterhin werden die Variabilität und paedomorphe Erscheinungen bei Wo. sphaeroides diskutiert. Die Wocklumeriaceae repräsentieren im höchsten Famennium (UD VI-C/D) die Radiationsphase einer langlebigen phylogenetischen Linie, die sich parallel zu anderen Clymenien entwickelte. Sie ist durch Suturmerkmale und lange Wohnkammern bei niedrigmündigen Gehäusen charakterisiert. Die als polyphyletisch erkannten Gonioclymeniina werden in die Wocklumeriina (mit Wocklumeriaceae, Glatziellaceae n.superfam. und Biloclymeniaceae) und in die Gonioclymeniaceae der Clymeniina geteilt. Gyroclymenia Czarnocki ist ein jüngeres Synonym von Pleuroclymenia Schindewolf, umfasst jedoch nicht die ,Pleuro.' americana -und ,Pleuro.' eurylobica -Artgruppen. Die Gattung ist die ursprünglichste Form der Wocklumeriina und stellt gleichzeitig das Bindeglied zu Platyclymenia (Varioclymenia) der Clymeniaceae dar. Die Gonioclymeniaceae wurzeln in fortgeschrittenen Platyclymeniidae. [source]


    Eastern Beringian biogeography: historical and spatial genetic structure of singing voles in Alaska

    JOURNAL OF BIOGEOGRAPHY, Issue 8 2010
    Marcelo Weksler
    Abstract Aim Pleistocene climatic cycles have left marked signatures in the spatial and historical genetic structure of high-latitude organisms. We examine the mitochondrial (cytochrome b) genetic structure of the singing vole, Microtus miurus (Rodentia: Cricetidae: Arvicolinae), a member of the Pleistocene Beringian fauna, and of the insular vole, Microtus abbreviatus, its putative sister species found only on the St Matthew Archipelago. We reconstruct the phylogenetic and phylogeographical structure of these taxa, characterize their geographical partitioning and date coalescent and cladogenetic events in these species. Finally, we compare the recovered results with the phylogenetic, coalescent and spatial genetic patterns of other eastern Beringian mammals and high-latitude arvicoline rodents. Location Continental Alaska (alpine and arctic tundra) and the St Matthew Archipelago (Bering Sea). Methods We generated and analysed cytochrome b sequences of 97 singing and insular voles (M. miurus and M. abbreviatus) from Alaska. Deep evolutionary structure was inferred by phylogenetic analysis using parsimony, maximum likelihood and Bayesian approaches; the geographical structure of genetic diversity was assessed using analysis of molecular variance and network analysis; ages of cladogenetic and coalescent events were estimated using a relaxed molecular clock model with Bayesian approximation. Results Regional nucleotide diversity in singing voles is higher than in other high-latitude arvicoline species, but intra-population diversity is within the observed range of values for arvicolines. Microtus abbreviatus specimens are phylogenetically nested within M. miurus. Molecular divergence date estimates indicate that current genetic diversity was formed in the last glacial (Wisconsinan) and previous interglacial (Sangamonian) periods, with the exception of a Middle Pleistocene split found between samples collected in the Wrangell Mountains region and all other singing vole samples. Main conclusions High levels of phylogenetic and spatial structure are observed among analysed populations. This pattern is consistent with that expected for a taxon with a long history in Beringia. The spatial genetic structure of continental singing voles differs in its northern and southern ranges, possibly reflecting differences in habitat distribution between arctic and alpine tundra. Our phylogenetic results support the taxonomic inclusion of M. miurus in its senior synonym, M. abbreviatus. [source]


    The effect of environmental diversification on species diversification in New Caledonian caddisflies (Insecta: Trichoptera: Hydropsychidae)

    JOURNAL OF BIOGEOGRAPHY, Issue 5 2010
    Marianne Espeland
    Abstract Aim, To test whether environmental diversification played a role in the diversification of the New Caledonian Hydropsychinae caddisflies. Location, New Caledonia, south-west Pacific. Methods, The phylogeny of the New Caledonian Hydropsychinae caddisflies was hypothesized using parsimony and Bayesian methods on molecular characters. The Bayesian analysis was the basis for a comparative analysis of the correlation between phylogeny and three environmental factors: geological substrate (ultrabasic, non-ultrabasic), elevation and precipitation. Phylogenetic divergence times were estimated using a relaxed clock method, and environmental factors were mapped onto a lineage-through-time plot to investigate the timing of environmental diversification in relation to species radiation. The correlation between rainfall and elevation was tested using independent contrasts, and the gamma statistic was calculated to infer the diversification pattern of the group. Results, The diversification of extant Orthopsyche,Caledopsyche species began in the Middle,Late Oligocene, when much of the island of New Caledonia was covered by ultrabasic substrate and mountain forming was prevalent. Most lineages originated in the Middle,Late Miocene, a period associated with long-term climate oscillation. Optimization of environmental factors on the phylogeny demonstrated that the New Caledonian Hydropsychinae group adapted to ultrabasic substrate early in its evolutionary history. The clade living mostly on ultrabasic substrate was far more species-rich than the clade living mostly on non-ultrabasic substrate. Elevation and rainfall were significantly correlated with each other. The lineage-through-time plot revealed that the main environmental diversification preceded species diversification. A constant speciation through time was rejected, and the negative gamma indicates that most of the diversification occurred early in the history of the clade. According to the inferred phylogeny, the genus Orthopsyche McFarlane is a synonym under Caledopsyche Kimmins, and Abacaria caledona Oláh & Barnard should also be included in Caledopsyche. Main conclusions, The age of the radiation does not support a vicariance origin of New Caledonian Hydropsychinae caddisflies. Environmental diversification pre-dates lineage diversification, and thus environmental heterogeneity potentially played a role in the diversification of the group, by providing a variety of fragmented habitats to disperse into, promoting speciation. The negative gamma indicates that the speciation rate slowed as niches started to fill. [source]


    Phylogenetics of European cyprinids: insights from allozymes

    JOURNAL OF FISH BIOLOGY, Issue 2 2000
    B. Häunfling
    Allozyme data from 23 central European cyprinid taxa suggest that the two sub-families Alburninae and Leuciscinae are paraphyletic. However together, both families seem to form a monophyletic group. By using a molecular clock, the radiation of this group is estimated to be as old as 20 million years. This is consistent with the fossil record. The genus Leuciscus is not monophyletic. The commonly used genus Blicca is a junior synonym to Abramis and both genera should become merged. The genus Tinca is rather distinct from the Alburninae Leuciscinae lineage and clearly does not belong to this group. These conclusions are consistent with published phylogenetic analyses of sequence data. These analyses underline that despite the developments in sequencing allozymes are a cheap and valuable tool in phylogenetics. Combining the estimate of the phylogenetic tree and biogeographic data, there was a decrease of distributional overlap when moving from the tips of the tree to the root. This pattern is consistent with sympatric speciation. [source]


    Marine macroalgae analyzed by mass spectrometry are rich sources of polyunsaturated fatty acids,

    JOURNAL OF MASS SPECTROMETRY (INCORP BIOLOGICAL MASS SPECTROMETRY), Issue 12 2005
    L. De Angelis
    Abstract Algae from cold water (Canada) and warm water (China) were analyzed for their total lipid content, and for their fatty acid (FA) composition and content. The major findings are that FA from Canadian algae are generally richer in polyunsaturated FA (PUFA), with a higher n-3/n-6 FA ratio, and a higher degree of total unsaturation. The 18 C, 4 double bonds FA (18 : 4 stearidonic acid, morotic acid as synonym) was detected in greater amounts in cold water samples. The high levels of total PUFA, and especially of n-3 FA in Canadian algae, suggests their possible utilizations for nutritional purposes. Copyright © 2005 John Wiley & Sons, Ltd. [source]


    Integrin ,3,1 interacts with I1PP2A/lanp and phosphatase PP1

    JOURNAL OF NEUROSCIENCE RESEARCH, Issue 8 2006
    Diana Mutz
    Abstract Integrin ,3,1 is a receptor for the extracellular matrix component laminin 5. To elucidate possible signaling pathways induced by integrin ,3,1, we looked for proteins that interact with the cytoplasmic part of the ,3A integrin subunit. We identified several multifunctional proteins by affinity chromatography and subsequent MALDI-TOF-MS and focused on the inhibitor 1 of serine/threonine phosphatase PP2A (I1PP2A, synonym: lanp) which also plays a role during the development of the mouse cerebellum. I1PP2A/lanp colocalizes with the ,3A integrin subunit in differentiated PC12 cells in the cell body and in neurites as well as in Purkinje cells of mouse cerebellum. Overexpression of GFP-I1PP2A/lanp in PC12 cells leads to markedly reduced neurite length on laminin 5 after induction with nerve growth factor. By affinity chromatography the protein phosphatase PP1 can also be identified as a ,3A/cyto-binding protein. PP1 and integrin ,3,1 can be pulled down by GST-I1PP2A/lanp from cell lysates of differentiated and undifferentiated PC12 cells. The phosphatase binds to the cytoplasmic membrane-proximal conserved GFFKR motif of the , integrin subunit, whereas I1PP2A/lanp requires a longer sequence for binding. PP1 but not PP2A is able to dephosphorylate precipitated integrin ,3,1 in vitro. Furthermore, PP1 releases phosphate from T1046 of phosphopeptides that mimic the phosphorylation consensus sequence in the cytoplasmic part of the ,3A integrin subunit. These data suggest that I1PP2A/lanp forms a complex with PP1 and the ,3A integrin subunit and might possibly regulate the phosphorylation status of integrin ,3,1 and/or integrin downstream targets. © 2006 Wiley-Liss, Inc. [source]


    GEITLERINEMA SPECIES (OSCILLATORIALES, CYANOBACTERIA) REVEALED BY CELLULAR MORPHOLOGY, ULTRASTRUCTURE, AND DNA SEQUENCING,

    JOURNAL OF PHYCOLOGY, Issue 3 2009
    Maria Do Carmo Bittencourt-Oliveira
    Geitlerinema amphibium (C. Agardh ex Gomont) Anagn. and G. unigranulatum (Rama N. Singh) Komárek et M. T. P. Azevedo are morphologically close species with characteristics frequently overlapping. Ten strains of Geitlerinema (six of G. amphibium and four of G. unigranulatum) were analyzed by DNA sequencing and transmission electronic and optical microscopy. Among the investigated strains, the two species were not separated with respect to cellular dimensions, and cellular width was the most varying characteristic. The number and localization of granules, as well as other ultrastructural characteristics, did not provide a means to discriminate between the two species. The two species were not separated either by geography or environment. These results were further corroborated by the analysis of the cpcB- cpcA intergenic spacer (PC-IGS) sequences. Given the fact that morphology is very uniform, plus the coexistence of these populations in the same habitat, it would be nearly impossible to distinguish between them in nature. On the other hand, two of the analyzed strains were distinct from all others based on the PC-IGS sequences, in spite of their morphological similarity. PC-IGS sequences indicate that these two strains could be a different species of Geitlerinema. Using morphology, cell ultrastructure, and PC-IGS sequences, it is not possible to distinguish G. amphibium and G. unigranulatum. Therefore, they should be treated as one species, G. unigranulatum as a synonym of G. amphibium. [source]


    Synthesis and degradation of biomedical materials based on linear and star shaped polyglycidols

    JOURNAL OF POLYMER SCIENCE (IN TWO SECTIONS), Issue 13 2009
    Helmut Keul
    Abstract Linear and star shaped polyglycidols (synonym with polyglycerols) are prepared in a controlled ring opening polymerization of protected glycidols. Beside the molar mass and the polydispersity, the architecture of the polyglycidols is controlled by using mono- and multifunctional mono- and polydispers initiators. Copolymers of dissimilarly protected glycidols as well as copolymers with nonfunctional oxiranes were prepared by means of anionic polymerization while copolymers of protected glycidol with tetrahydrofuran were prepared by means of cationic polymerization. Polyethers with functional groups in the side chains (functional polyethers) with special emphasis on polyglycidols (containing hydroxymethyl groups in the side chains) were used to prepare multifunctional polymers and (hetero)grafted polymer brushes via chemical and enzyme catalyzed reaction. The potential of poly(glycidol- graft -,-caprolactone)- co -glycidol) prepared via enzyme catalyzed grafting of polyglycidols using ,-caprolactone as a monomer and Lipase B from Candida antarctica as a catalyst is presented. Finally, comparative degradation studies of densely and loosely grafted polyglycidols are presented and discussed. © 2009 Wiley Periodicals, Inc. J Polym Sci Part A: Polym Chem 47: 3209,3231, 2009 [source]


    Design , an inappropriate concept in evolutionary theory,

    JOURNAL OF ZOOLOGICAL SYSTEMATICS AND EVOLUTIONARY RESEARCH, Issue 1 2009
    W. J. Bock
    Abstract The concept of accident in evolution refers to causes which are stochastic with respect to selective demands arising from the external environment and acting on the organism, while the concept of design refers to causes which meet the requirement of these selective demands. The condition ,with respect to selective demands' is generally forgotten so that evolutionary changes are described as being design modifications. Design is an invalid synonym for adaptation. Further it implies a designer and has been used by some authors since before Darwin to argue that design in organisms demonstrates the existence of a designer and hence a plan. Yet if evolution depends on two simultaneously acting causes, one of which is accidental, then the process of evolution and all attributes of organisms are accidental. The concept of design is inappropriate in biology and should be eliminated from all biological explanations. [source]


    Towards a phylogenetic classification of dendrocoelid freshwater planarians (Platyhelminthes): a morphological and eclectic approach

    JOURNAL OF ZOOLOGICAL SYSTEMATICS AND EVOLUTIONARY RESEARCH, Issue 4 2006
    R. Sluys
    Abstract We explore and review the taxonomic distribution of morphological features that may be used as supporting apomorphies for the monophyletic status of various taxa in future, more comprehensive phylogenetic analyses of the dendrocoelid freshwater planarians and their close relatives. Characters examined are: (i) musculature of the pharynx; (ii) anatomy of the eye; (iii) distribution of testis follicles along the longitudinal axis of the body; (iv) epidermis of the body wall; (v) musculature of the ventral body wall; (vi) presence or absence of a penis papilla; (vii) position of the mouth opening; and (viii) anterior adhesive organ. A phylogenetic hypothesis, in the form of a character state tree, is developed. This hypothesis suggests that the Kenkiidae should be resurrected as a separate family, sharing a sistergroup relationship with the Dendrocoelidae. This sistergroup relationship is supported by the shared presence of an apomorphic anterior adhesive organ. Although the shape of the anterior adhesive organ, both within and between kenkiids and dendrocoelids, may differ considerably between species, they are all constructed according to the same principles. The monophyly of the Kenkiidae is supported by the presence of an apomorphic thickening of the marginal zone of the epidermis and presumably also by the presence of prepharyngeal testes. Within the Kenkiidae, the genus Macrocotyla should be considered as a junior synonym of Kenkia, which is characterized by the presence of an apomorphic ,planariid pharynx-type 2'. The monophyly of the Dendrocoelidae is suggested by the presence of a dendrocoelid type of pharynx. It is shown that multicellular eye cups with numerous photoreceptor cells are not restricted to the Dugesiidae and the Terricola but are also characteristic for the Dendrocoelidae; the phylogenetic implications of this character state distribution are discussed. It is pointed out that in planarians there may be a relationship between: (i) body size, the size of an eye cup and the number of its constituent pigment cells; and (ii) the size of an eye cup and the number of photoreceptor cells that it contains. Some tentative phylogenetic groups are suggested on the basis of the following presumed apomorphic characters: presence of transverse fibres in the ventral subepidermal musculature, presence of an extra longitudinal muscle layer in the outer pharynx musculature, a mouth opening that has shifted anteriad. It is suggested that the presumed monophyly of the genus Bdellocephala is supported by the apomorphic absence of a penis papilla, combined with a penial lumen provided with many plicae. Zusammenfassung Wir untersuchen und begutachten die taxonomische Verbreitung morphologischer Merkmale, die man als Apomorphien für die Monophylie verschiedener Taxa in einer zukünftigen umfangreicheren phylogenetischen Analyse der Dendrocoeliden und ihrer nächsten Verwandten heranziehen könnte. Zu diesen untersuchten Merkmalen gehören die Pharynxmuskulatur, die Augen-Anatomie, die Anordnung der Hodenfollikel entlang der Längsachse des Körpers, die Epidermis, die Muskulatur der ventralen Körperwand, das Vorhandensein oder Fehlen einer Penispapille, die Lage der Mundöffnung sowie des vorderen Hartorgans. Eine phylogenetische Hypothese wurde als character state-Stammbaum entwickelt. Hierin wird vorgeschlagen, die Kenkiidae wieder als eigenständige Familie und als Schwestergruppe der Dendrocoelidae zu betrachten. Dieses Schwestergruppen-Verhältnis wird durch den gemeinsamen Besitz eines apomorphen vorderen Haftorgans gestützt. Obwohl sich die Form dieses Organs sowohl innerhalb als auch zwischen Kenkiiden und Dendrocoeliden beträchtlich zwischen den Arten unterscheiden kann, ist es grundsätzlich gleich aufgebaut. Die Monophylie der Kenkiidae wird durch eine als apomorph betrachtete Verdickung des Epidermisrandes und durch das Auftreten praepharyngealer Hoden gestützt. Innerhalb der Kenkiidae sollte die Gattung Macrocotyla als Juniorsynonym von Kenkia angesehen werden, die durch den apomorphen Planarien-Pharynx-Typ 2 charakterisiert ist. Als Beleg für die Monophylie der Dendrocoelidae wird der Dendrocoeliden-Pharynx-Typ vorgeschlagen. Es wird gezeigt, dass vielzellige Augenbecher mit zahlreichen Photorezeptorzellen nicht auf die Dugesiidae und die Terricola beschränkt, sondern auch für die Dendrocoelidae charakteristisch sind; die phylogenetische Bedeutung dieser Merkmalsverteilung wird diskutiert. Es wird hervorgehoben, dass es in den Planarien wahrscheinlich eine Beziehung zwischen (i) Körpergröße, Größe der Augenbecher und Zahl der Photorezeptorzellen ebenso wie (ii) der Größe eines Augenbechers und der Zahl seiner Photorezeptorzellen gibt. Versuchsweise werden einige phylogenetische Gruppierungen auf der Basis der folgenden vermutlich apomorphen Merkmale vorgeschlagen: Vorhandensein transversaler Fasern in der ventralen subepidermalen Muskulatur, Auftreten einer zusätzlichen Längsmuskelschicht in der äußeren Pharynxmuskulatur und eine nach vorn verschobene Mundöffnung. Es wird weiterhin vorgeschlagen, die vermutete Monophylie der Gattung Bdellocephala durch das Fehlen einer Penispapille als Autapomorphie zu stützen, zusammen mit dem Merkmal eines durch viele Falten gekennzeichneten Penislumens. [source]


    Vibrio harveyi: a significant pathogen of marine vertebrates and invertebrates

    LETTERS IN APPLIED MICROBIOLOGY, Issue 2 2006
    B. Austin
    Abstract Vibrio harveyi, which now includes Vibrio carchariae as a junior synonym, is a serious pathogen of marine fish and invertebrates, particularly penaeid shrimp. In fish, the diseases include vasculitis, gastro-enteritis and eye lesions. With shrimp, the pathogen is associated with luminous vibriosis and Bolitas negricans. Yet, the pathogenicity mechanisms are imprecisely understood, with likely mechanisms involving the ability to attach and form biofilms, quorum sensing, various extracellular products including proteases and haemolysins, lipopolysaccharide, and interaction with bacteriophage and bacteriocin-like substances. [source]


    RESURRECTION OF MESOPLODON TRAVERSII (GRAY, 1874), SENIOR SYNONYM OF M. BAHAMONDI REYES, VAN WAEREBEEK, CÁRDENAS AND YÁÑEZ, 1995 (CETACEA: ZIPHIIDAE)

    MARINE MAMMAL SCIENCE, Issue 3 2002
    Anton L. van Helden
    Abstract Mesoplodon traversii (Gray, 1874) is shown to be a senior synonym of the recently described beaked whale Mesoplodon hahamondi Reyes et al., 1995 on the basis of a phylogenetic analysis of mitochondrial DNA control region sequences. The mandible and teeth of M. traversii, first reported in 1873 by Hector as Dolichodon layardii. are redescribed. The species can be distinguished by features of the calvaria; including the large jugal, broad rostrum, and small distance between premaxillary foramina. The male teeth, which are large and spade-shaped with a strong terminal denticle, are also diagnostic. M. traversii is known only from Pitt Island and White Island, New Zealand and Robinson Crusoe Island, Juan Fernandez Archipelago, Chile. [source]


    Specific validity of Meligethes cyrenaicus Rebmann, 1940, and an updated identification key for the M. carinulatus species-group (Coleoptera, Nitidulidae)

    MITTEILUNGEN AUS DEM MUSEUM FUER NATURKUNDE IN BERLIN-DEUTSCHE ENTOMOLOGISCHE ZEITSCHRIFT, Issue 1 2002
    Paolo Audisio
    Abstract The North African species Meligethes cyrenaicus Rebmann, 1940, a member of the M. carinulatus species-group, previously considered a synonym of the widespread European M. carinulatus Förster, 1849, is resurrected at a specific rank. This taxonomic settlement is chiefly based on an analysis of external morphological characters of the type material of M. cyrenaicus (from Libya, Cyrenaica) and of a large series of southern European specimens of M. carinulatus. An updated key for the identification of the eight members of the Meligethes carinulatus species-group is presented. [source]


    Critical notes on Saussurea pulvinata (Asteraceae) in China

    NORDIC JOURNAL OF BOTANY, Issue 3-4 2008
    Bing-Qiang Xu
    Saussurea ruoqiangensis K. M. Shen is reduced to a synonym of S. pulvinata Maxim., a distribution map and a key to S. pulvinata and related species is provided. [source]


    A key and four new species of Hedysarum (Fabaceae) in Iran

    NORDIC JOURNAL OF BOTANY, Issue 1-2 2008
    M. Ranjbar
    A key, descriptions and new findings are given for some species from of the short-stemmed group Hedysarum sect. Multicaulia, which occur within the area covered by the flora of the north-eastern part of Iran (Khorassan Province). They have short stem, becoming woody at base, standard as long as keel and pods, glabrous or densely covered with grey appressed hairs. In addition, H.longepedunculatum, H. bojnordense, H. paucifoliolatum and H. glabrifoliolatum are described and illustrated as new species. The relationships between the new species and their closest relatives are discussed. Hedysarum fallacinum Rech. f. & Aellen is reduced to a synonym of H. kopetdaghi Boriss. on the basis of their common characters such as arrangement of branches and flowers, shape of leaflet and size of calyx and corolla. [source]


    On Hieracium incurrens (Asteraceae) and similar species in central Sweden

    NORDIC JOURNAL OF BOTANY, Issue 3-4 2007
    Torbjörn Tyler
    Hieracium incurrens Saelan ex Norrl., originally described from Finland, has been reported from Sweden several times and independently by different authors. Still, however, its presence in Sweden has remained dubious and it has become obvious that the different reports are referable to different taxa and the name has thus been, at least partially, misapplied. In the present study, all Swedish material and putative taxa that have been suggested to belong to H. incurrens, as well as some additional closely similar taxa, is critically compared with authentic material of the latter from Finland. It is concluded that H. incurrens do indeed occur in central Sweden, namely in the eastern coastal provinces closest to Finland. However, the relevant material has mostly been referred to H. longilingua Johanss., which is here considered to be a later synonym of H. incurrens. Material previously referred to H. incurrens from other parts of Sweden does, however, represent other taxa. Five accepted species of this affinity are described and discussed in some detail and one species, viz. H. oestmanii T. Tyler is described anew. In addition, the unrelated species H. arnoldii T. Tyler is described anew based on material from the provinces of Hälsingland and Medelpad. [source]


    THE TAXONOMIC AND PHYLOGENETIC POSITION OF THE PLESIOSAUROIDEA FROM THE LOWER JURASSIC POSIDONIA SHALE OF SOUTH-WEST GERMANY

    PALAEONTOLOGY, Issue 3 2007
    FRANZISKA GROßMANNArticle first published online: 17 MAY 200
    Abstract:, The two plesiosauroid species from the Posidonia shale of Holzmaden, ,Plesiosaurus'guilelmiimperatoris and ,Plesiosaurus'brachypterygius, do not belong to Plesiosaurus but form new monotypic genera. The new genus Hydrorion is erected for ,P.'brachypterygius, and the genus Seeleyosaurus is re-established for ,P.'guilelmiimperatoris. The recently described species Plesiopterys wildii is regarded as a junior synonym of S. guilelmiimperatoris. A short phylogenetic analysis shows that S. guilelmiimperatoris and Muraenosaurus are basal elasmosaurs. H. brachypterygius, Occitanosaurus tournemirensis from France and Microcleidus homalospondylus from England form a monophyletic clade, which is the sister taxon to the Cretaceous elasmosaurs. A palaeobiogeographical comparison of plesiosaur localities in the Lower Jurassic shows distinct palaeobiogeographical zones for the Toarcian, with different plesiosaur taxa in England, Germany and France. [source]


    CRANIAL CREST DEVELOPMENT IN THE AZHDARCHOID PTEROSAUR TUPUXUARA, WITH A REVIEW OF THE GENUS AND TAPEJARID MONOPHYLY

    PALAEONTOLOGY, Issue 4 2006
    DAVID M. MARTILL
    Abstract:, A portion of pterosaur skull from the Romualdo Member of the Santana Formation (?Albian,?Turonian, Cretaceous) of north-east Brazil provides new data on the morphology and ontogeny of azhdarchoid pterosaur cranial crests. The specimen consists of parts of the cranial bones posterodorsal to the nasoantorbital fenestra, including partial nasals, lacrimals, frontals and possibly the parietals. A posterodorsally directed premaxillary crest with a concave posterior border is located dorsal to the posterior border of the nasoantorbital fenestra. A well-defined suture indicates overlapping, posterodorsally directed growth of the premaxilla over the skull roof, suggesting that the generation of the premaxillary crest is a late ontogenetic feature and thus probably related to sexual display. The systematics of Tupuxuara and its relationship to other azhdarchoids is reviewed and a cladistic analysis of the group is presented. Tupuxuara is found to be the sister-taxon to Azhdarchidae. Tupuxuara longicristatus Kellner and Campos, 1988 is argued to be the only valid named species in this genus and Thalassodromeus Kellner and Campos, 2002 is considered a junior subjective synonym of this taxon. As originally conceived, Tapejaridae is paraphyletic: a new, more restrictive version of Tapejaridae (including Tapejara and Sinopterus dongi) might exist, but its monophyly is weakly supported. Furthermore, Tapejara was found to be paraphyletic in all trees. [source]


    SYSTEMATICS OF THE CAMBRIAN TRILOBITE FAMILY NEPEIDAE, WITH REVISION OF AUSTRALIAN SPECIES

    PALAEONTOLOGY, Issue 3 2005
    JOHN R. PATERSON
    Abstract:, Taxonomy of the Cambrian trilobite family Nepeidae is revised. Morphometric analyses of the genera Nepea and Penarosa demonstrate that use of proportions of linear dimensions of the cranidium to differentiate species is invalid, and that infrageneric variation is continuous. The taxonomy and biostratigraphy of all Australian species of Nepeidae is revised. Species considered valid herein include: Nepea narinosa (type species), N. tonsillata, N. nans, Penarosa retifera (type species), P. elaticeps, P. rhinodelphis, P. netenta, Loxonepea loxophrys (type species) and Ferenepea hispida (type species). Folliceps is considered to be a junior subjective synonym of Nepea, Trinepea is regarded as a junior subjective synonym of Penarosa, and Ascionepea is considered to be a junior subjective synonym of Ferenepea. [source]