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Steppe
Terms modified by Steppe Selected AbstractsMid-Holocene and glacial-maximum vegetation geography of the northern continents and AfricaJOURNAL OF BIOGEOGRAPHY, Issue 3 2000I. Colin Prentice Abstract BIOME 6000 is an international project to map vegetation globally at mid-Holocene (6000 14C yr bp) and last glacial maximum (LGM, 18,000 14C yr bp), with a view to evaluating coupled climate-biosphere model results. Primary palaeoecological data are assigned to biomes using an explicit algorithm based on plant functional types. This paper introduces the second Special Feature on BIOME 6000. Site-based global biome maps are shown with data from North America, Eurasia (except South and Southeast Asia) and Africa at both time periods. A map based on surface samples shows the method's skill in reconstructing present-day biomes. Cold and dry conditions at LGM favoured extensive tundra and steppe. These biomes intergraded in northern Eurasia. Northern hemisphere forest biomes were displaced southward. Boreal evergreen forests (taiga) and temperate deciduous forests were fragmented, while European and East Asian steppes were greatly extended. Tropical moist forests (i.e. tropical rain forest and tropical seasonal forest) in Africa were reduced. In south-western North America, desert and steppe were replaced by open conifer woodland, opposite to the general arid trend but consistent with modelled southward displacement of the jet stream. The Arctic forest limit was shifted slighly north at 6000 14C yr bp in some sectors, but not in all. Northern temperate forest zones were generally shifted greater distances north. Warmer winters as well as summers in several regions are required to explain these shifts. Temperate deciduous forests in Europe were greatly extended, into the Mediterranean region as well as to the north. Steppe encroached on forest biomes in interior North America, but not in central Asia. Enhanced monsoons extended forest biomes in China inland and Sahelian vegetation into the Sahara while the African tropical rain forest was also reduced, consistent with a modelled northward shift of the ITCZ and a more seasonal climate in the equatorial zone. Palaeobiome maps show the outcome of separate, independent migrations of plant taxa in response to climate change. The average composition of biomes at LGM was often markedly different from today. Refugia for the temperate deciduous and tropical rain forest biomes may have existed offshore at LGM, but their characteristic taxa also persisted as components of other biomes. Examples include temperate deciduous trees that survived in cool mixed forest in eastern Europe, and tropical evergreen trees that survived in tropical seasonal forest in Africa. The sequence of biome shifts during a glacial-interglacial cycle may help account for some disjunct distributions of plant taxa. For example, the now-arid Saharan mountains may have linked Mediterranean and African tropical montane floras during enhanced monsoon regimes. Major changes in physical land-surface conditions, shown by the palaeobiome data, have implications for the global climate. The data can be used directly to evaluate the output of coupled atmosphere-biosphere models. The data could also be objectively generalized to yield realistic gridded land-surface maps, for use in sensitivity experiments with atmospheric models. Recent analyses of vegetation-climate feedbacks have focused on the hypothesized positive feedback effects of climate-induced vegetation changes in the Sahara/Sahel region and the Arctic during the mid-Holocene. However, a far wider spectrum of interactions potentially exists and could be investigated, using these data, both for 6000 14C yr bp and for the LGM. [source] Range expansion of the globally Vulnerable Karamoja apalis Apalis karamojae in the Serengeti ecosystemAFRICAN JOURNAL OF ECOLOGY, Issue 3 2010Philip Shaw Abstract The underlying causes of change in geographic range size are less well understood in African birds than in north temperate species. Here, we examine factors associated with range expansion in the Karamoja apalis (Apalis karamojae), a globally Vulnerable warbler confined to north-east Uganda, north-central Tanzania and southern Kenya. In Tanzania, it was originally known only from the Wembere Steppe, but since 1993 (and possibly as early as 1983) has extended its range into the Serengeti ecosystem, c. 140 km to the north, reaching southern Kenya by 2004. Changes in the warbler's range within the Serengeti have broadly reflected a cyclical change in the density of its main habitat, Acacia drepanolobium woodland, which was low in the 1970s, high during the 1980s and 1990s, and declined in the early 2000s. Karamoja apalis records in the Serengeti showed a 5 year time lag behind A. drepanolobium density, which was in turn negatively correlated with the area of grassland burnt 10 years earlier. Previous studies in the Serengeti have also linked Acacia regeneration to changes in grazing pressure, as increasing wildebeest (Connochaetes taurinus) numbers have reduced the volume of combustible material present, and hence the frequency of damaging ,hot burns'. We conclude that this globally threatened warbler appears to have benefited from changes in ungulate populations in the Serengeti, which have influenced burning intensity and hence tree regeneration. The warbler's range now appears to be declining, however, following a recent reduction in the density and annual survival of A. drepanolobium in the northern Serengeti. Résumé Les causes sous-jacentes du changement de la taille d'une distribution géographique sont moins bien connues pour les oiseaux africains que pour les espèces du nord tempéré. Nous examinons ici des facteurs liés à l'expansion de l'aire de répartition de l'apalis de Karamoja Apalis karamojae, un sylviidé classé comme Vulnérable au niveau mondial, confiné au NE de l'Ouganda, au centre-nord de la Tanzanie et au sud du Kenya. En Tanzanie, on ne le connaissait à l'origine que dans la steppe de Wembere mais depuis 1993, et peut-être même dès 1983, il a étendu son aire de répartition dans l'écosystème du Serengeti, environ 140 km plus au nord, et atteint le sud du Kenya en 2004. Les changements de l'aire de répartition de ce sylviidé dans le Serengeti reflètent largement un changement cyclique de la densité de son habitat principal, la forêt d'Acacia drepanolobium, qui était faible dans les années 1970, forte pendant les années 1980 et 1990, et qui a diminué au début des années 2000. Les rapports sur l'apalis de Karamoja au Serengeti montrent un retard de cinq ans par rapport à l'évolution de la densité d'A. drepanolobium, elle-même étant négativement liée à la zone de prairie brûlée 10 ans plus tôt. Des études antérieures faites au Serengeti lient aussi la régénération des acacias aux changements de la pression du pâturage, étant donné que le nombre croissant de gnous, Connochaetes taurinus, a réduit le volume des matières combustibles restantes et donc la fréquence des feux trop chauds qui causent beaucoup de dégâts. Nous concluons que cet oiseau menacé au niveau mondial semble avoir bénéficié des changements des populations d'ongulés au Serengeti, qui ont influencé l'intensité des feux et donc la régénération des arbres. L'aire de répartition de ce sylviidé semble pourtant en train de se réduire suite à une récente réduction de la densité et de la survie annuelle d'A. drepanolobium dans le nord du Serengeti. [source] Holocene vegetation and climate changes in Hoton-Nur basin, northwest MongoliaBOREAS, Issue 2 2000PAVEL TARASOV Radiocarbon-dated pollen and diatom records from Hoton-Nur lake (48°40'N, 88°18'E), northwest Mongolia are used to reconstruct Holocene vegetation and climate changes. Steppe, which covered the area some time before 9000 BP (all ages are given in 14C BP), was replaced by boreal conifer forest-steppe by 9000,8500 BP. At the same time, planktonic diatoms increased in abundance from 5 to 45%. After 4000 BP there was a sharp reduction in forest and re-establishment of steppe. Changes in the pollen composition were accompanied by a decrease in the abundance of planktonic diatoms from 35,50% (between 9000 and 5500 BP) to less than 10% soon after 4000 BP. These and other published data from Mongolia suggest wetter conditions during the early and middle Holocene than those of today. This pattern probably reflects an expansion of the Pacific monsoon recorded in geological data from China and simulated in climate-model experiments, and suggests that the summer monsoon influenced a larger area than might appear if the Chinese records only are taken into consideration. [source] Performance of Greater Sage-Grouse Models for Conservation Assessment in the Interior Columbia Basin, U.S.A.CONSERVATION BIOLOGY, Issue 5 2002Michael J. Wisdom Consequently, we evaluated the performance of two models designed to assess landscape conditions for Greater Sage-Grouse across 13.6 million ha of sagebrush steppe in the interior Columbia Basin and adjacent portions of the Great Basin of the western United States (referred to as the basin). The first model, the environmental index model, predicted conditions at the scale of the subwatershed (mean size of approximately 7800 ha) based on inputs of habitat density, habitat quality, and effects of human disturbance. Predictions ranged on a continuous scale from 0 for lowest environmental index to 2 for optimal environmental index. The second model, the population outcome model, predicted the composite, range-wide conditions for sage grouse based on the contribution of environmental index values from all subwatersheds and measures of range extent and connectivity. Population outcomes were expressed as five classes (A through E) that represented a gradient from continuous, well-distributed populations (outcome A) to sparse, highly isolated populations with a high likelihood of extirpation (outcome E). To evaluate performance, we predicted environmental index values and population outcome classes in areas currently occupied by sage grouse versus areas where extirpation has occurred. Our a priori expectations were that models should predict substantially worse environmental conditions ( lower environmental index) and a substantially higher probability of extirpation ( lower population outcome class) in extirpated areas. Results for both models met these expectations. For example, a population outcome of class E was predicted for extirpated areas, as opposed to class C for occupied areas. These results suggest that our models provided reliable landscape predictions for the conditions tested. This finding is important for conservation planning in the basin, where the models were used to evaluate management of federal lands for sage grouse. Resumen: Modelos válidos de hábitat y de poblaciones del urogallo (Centrocercus urophasianus) son una necesidad crítica para su manejo debido a la creciente preocupación por la viabilidad de la población. Por lo tanto, evaluamos el funcionamiento de dos modelos diseñados para evaluar las condiciones del paisaje para el urogallo en 13.6 millones de ha de la estepa de artemisa en la Cuenca Columbia Interior y las porciones adyacentes de la Gran Cuenca de los Estados Unidos occidentales (referidos como cuenca). El primer modelo (modelo de índice ambiental) predijo condiciones a escala de la subcuenca (tamaño promedio , 7800 ha) basado en información de la densidad del hábitat, calidad del hábitat y efectos de la perturbación humana. Las predicciones variaron en una escala continua a partir de 0 (el índice ambiental más bajo) a 2 (índice ambiental óptimo). El segundo modelo (modelo del resultado de la población) predijo las condiciones compuestas, de amplio rango, para el urogallo con base en la contribución de los valores de índice ambiental de todas las subcuencas y las medidas de extensión y de conectividad de la pradera. Los resultados de la población fueron expresados en cinco clases (A - E) que representan un gradiente de poblaciones continuas, bien-distribuidas (resultado A) a poblaciones escasas, altamente aisladas con una alta probabilidad de extirpación (resultado E). Para evaluar el funcionamiento, predijimos valores de índice ambiental y resultados de la población en las áreas actualmente ocupadas por urogallos versus áreas donde ha ocurrido la extirpación. Nuestras expectativas a priori eran que los modelos deben predecir condiciones ambientales substancialmente peores (índice ambiental más bajo) y una probabilidad de extirpación sustancialmente mayor (menor resultado de la población) en áreas extirpadas. Los resultados para ambos modelos cumplieron estas expectativas. Por ejemplo, se predijo un resultado de la población de la clase E para áreas extirpadas, en comparación con la clase C para áreas ocupadas. Estos resultados sugieren que nuestros modelos proporcionaron predicciones de paisaje confiables para las condiciones probadas. Este hallazgo es importante para planeación de la conservación de la cuenca, donde los modelos fueron utilizados para evaluar el manejo de terrenos federales para urogallos. [source] Local and regional-scale responses of ant diversity to a semiarid biome transitionECOGRAPHY, Issue 4 2001Brandon T. Bestelmeyer The locations of biome transitions and ecotones are frequently defined by the rapid shift from one form of dominant vegetation to another. The composition of animal taxa is predicted to shift in parallel with that of dominant plants and species diversity is predicted to he greater in transitional zones than in adjacent areas. We asked whether ant species diversity and composition supported these predictions across a biome transition between shortgrass steppe and Chihuahuan desert vegetation. Neither species richness nor diversity was highest at the biome transition region as a whole, or within habitats in the biome transition. The biome transition region was not intermediate in ant species composition or in the representation of different faunal complexes. The community similarity between matched habitats shared between the biome transition zone and adjacent regions was less than that between distinct habitats occurring within regions. A zoogeographic transition for ants may occur to the north of the phytogeographic transition and may be coincident with the northern limits of monsoonal precipitation patterns. In contrast, the phytogeographic transition may be related to less extreme climatic variation within the monsoonal region occurring further south. [source] Paleoindian environmental change and landscape response in Barger Gulch, Middle Park, ColoradoGEOARCHAEOLOGY: AN INTERNATIONAL JOURNAL, Issue 6 2005James H. Mayer Middle Park, a high-altitude basin in the Southern Rocky Mountains of north-central Colorado, contains at least 59 known Paleoindian localities. At Barger Gulch Locality B, an extensive Folsom assemblage (,10,500 14C yr B.P.) occurs within a buried soil. Radiocarbon ages of charcoal and soil organic matter, as well as stratigraphic positions of artifacts, indicate the soil is a composite of a truncated, latest-Pleistocene soil and a younger mollic epipedon formed between ,6000 and 5200 14C yr B.P. and partially welded onto the older soil following erosion and truncation. Radiocarbon ages from an alluvial terrace adjacent to the excavation area indicate that erosion followed by aggradation occurred between ,10,200 and 9700 14C yr B.P., and that the erosion is likely related to truncation of the latest-Pleistocene soil. Erosion along the main axis of Barger Gulch occurring between ,10,000 and 9700 14C yr B.P. was followed by rapid aggradation between ,9700 and 9550 14C yr B.P., which, along with the erosion at Locality B, coincides with the abrupt onset of monsoonal precipitation following cooling in the region ,11,000,10,000 14C yr B.P. during the Younger Dryas oscillation. Buried soils dated between ,9500 and 8000 14C yr B.P. indicate relative landscape stability and soil formation throughout Middle Park. Morphological characteristics displayed by early Holocene soils suggest pedogenesis under parkland vegetation in areas currently characterized by sagebrush steppe. The expansion of forest cover into lower elevations during the early Holocene may have resulted in lower productivity in regards to mammalian fauna, and may partly explain the abundance of early Paleoindian sites (,11,000,10,000 14C yr B.P., 76%) relative to late Paleoindian sites (,10,000,8000 14C yr B.P., 24%) documented in Middle Park. © 2005 Wiley Periodicals, Inc. [source] Increased temperature and precipitation interact to affect root production, mortality, and turnover in a temperate steppe: implications for ecosystem C cyclingGLOBAL CHANGE BIOLOGY, Issue 4 2010WENMING BAI Abstract Fine root production and turnover play important roles in regulating carbon (C) cycling in terrestrial ecosystems. In order to examine effects of climate change on root production and turnover, a field experiment with increased temperature and precipitation had been conducted in a semiarid temperate steppe in northern China since April 2005. Experimental warming decreased annual root production, mortality, and mean standing crop by 10.3%, 12.1%, 7.0%, respectively, while root turnover was not affected in 2006 and 2007 by the warming. Annual root production and turnover was 5.9% and 10.3% greater in the elevated than ambient precipitation plots. Changes in root production and mortality in response to increased temperature and precipitation could be largely attributed to the changes in gross ecosystem productivity (GEP) and belowground/aboveground C allocation. There were significant interactive effects of warming and increased precipitation on root productivity, mortality, and standing crop. Experimental warming had positive and negative effects on the three root variables (root production, mortality, standing crop) under ambient and increased precipitation, respectively. Increased precipitation stimulated and suppressed the three root variables in the unwarmed and warmed subplots, respectively. The positive dependence of soil respiration and ecosystem respiration upon root productivity and mortality highlights the important role of root dynamics in ecosystem C cycling. The nonadditive effects of increased temperature and precipitation on root productivity, mortality, and standing crop observed in this study are critical for model projections of climate,ecosystem feedbacks. These findings indicate that carbon allocation is a focal point for future research and that results from single factor experiments should be treated with caution because of factor interactions. [source] Effects of ultraviolet radiation on litter decomposition depend on precipitation and litter chemistry in a shortgrass steppe ecosystemGLOBAL CHANGE BIOLOGY, Issue 10 2007LESLIE A. BRANDT Abstract We examined the effect of altered levels of ultraviolet (UV) radiation (280,400 nm) and different amounts of precipitation on the decomposition rates of litter of contrasting carbon to nitrogen ratio (C : N) in a 3-year field experiment in a shortgrass steppe (SGS) ecosystem. UV radiation was either blocked or passed under clear plastic tents where precipitation was applied to simulate a very dry or very wet year. These treatments minimized or maximized the abiotic component (UV) or the biotic component (biological activity of decomposer organisms) of decomposition to assess potential interactions between the two. Initial litter chemistry varied in response to having been grown under ambient or elevated atmospheric CO2 concentrations. While precipitation and litter chemistry were the most important drivers in decomposition in this system, UV radiation increased decomposition rates under dry conditions in litter with higher C : N ratios. Exposure to UV radiation slightly increased the amount of holocellulose that was lost from the litter. UV exposure did not affect the decomposition of the lignin fraction. Increased decomposition with UV radiation was accompanied by a decrease in N immobilization over the summer months. These results suggest that the effects of UV radiation on decomposition rates may be primarily abiotic, caused by direct photochemical degradation of the litter. Our results demonstrate that the role of UV radiation in litter decomposition in semiarid systems depends on the aridity of the system and the chemistry of the litter. [source] Potential for detecting changes in soil organic carbon concentrations resulting from climate changeGLOBAL CHANGE BIOLOGY, Issue 11 2003Franz Conen Abstract The interaction between soil organic carbon pools and climate change is an important determinant of future atmospheric CO2 concentrations. Much effort has so far been allocated to manipulative process studies and predictive modelling exercises. Here, we examine the potential for directly detecting predicted changes through repeated soil sampling. Two contrasting benchmark plots were selected in the steppe at the Russian,Mongolian border, where soil organic carbon losses are predicted to be around 10% over the first 50 years of climate change. In both plots, 50 samples were taken to 20 and 30 cm depths. The estimated time intervals before re-sampling by the same method that were likely to prove significant soil organic carbon losses (,=0.05; statistical power=0.90) were 43 and 26 years. [source] Organic carbon and carbon isotopes in modern and 100-year-old-soil archives of the Russian steppeGLOBAL CHANGE BIOLOGY, Issue 10 2002Margaret S. Torn Abstract Archived soils can provide valuable information about changes in the carbon and carbon isotope content of soils during the past century. We characterized soil carbon dynamics in a Russian steppe preserve using a 100-year-old-soil archive and modern samples collected from the same site. The site has been protected since 1885 to the present, during which time the region has experienced widespread conversion to cultivation, a decrease in fire frequency, and a trend of increasing precipitation. In the preserve, the amount of organic carbon did not change appreciably between the 1900 and 1997 sampling dates, with 32 kg C/m2 in the top meter and a third of that in the top 20 cm. Carbon and nitrogen stocks varied by less than 6% between two replicate modern soil pits or between the modern sites and the archive. Radiocarbon content decreased with depth in all sites and the modern SOM had positive , values near the surface due to nuclear weapons testing in the early 1960s. In the upper 10 cm, most of the SOM had a turnover time of 6,10 years, according to a model fit to the radiocarbon content. Below about 10 cm, the organic matter was almost all passive material with long (millennial) turnover times. Soil respiration ,14CO2 on a summer day was 106,109,, an isotopic disequilibrium of about 9, relative to atmospheric 14CO2. In both the modern and archive soil, the relative abundance of 13C in organic matter increased with depth by 2, in the upper meter from ,13C = --26, at 5 cm to --24, below a meter. In addition, the slope of ,13C vs. depth below 5 cm was the same for both soils. Given the age of the soil archive, these results give clear evidence that the depth gradients are not due to depletion of atmospheric 13CO2 by fossil fuel emissions but must instead be caused by isotopic fractionation between plant litter inputs and preservation of SOM. Overall, the data show that these soils have a large reservoir of recalcitrant C and stocks had not changed between sampling dates 100 years apart. [source] Anthropogenic and climatic impacts on surface pollen assemblages along a precipitation gradient in north-eastern ChinaGLOBAL ECOLOGY, Issue 5 2010Yun Zhang ABSTRACT Aim, To understand the scenarios of ,anthropogenic biomes' that integrate human and ecological systems, we need to explore the impacts of climate and human disturbance on vegetation in the past and present. Interactions among surface pollen, modern vegetation and human activities along climate and land-use gradients are tested to evaluate the natural and anthropogenic forces shaping the modern vegetation, and hence to aid the reconstruction of vegetation and climate in the past. This in turn will help with future predictions. Location, The North-east China Transect (NECT) in north-eastern China. Methods, We analysed 33 surface pollen samples and 213 quadrats across four vegetation zones along the moisture/land-use gradients of the NECT. Detrended correspondence analysis (DCA) and redundancy analysis (RDA) of 52 pollen taxa and three environmental variables were used to distinguish anthropogenic and climatic factors that affect surface pollen assemblages along the NECT. Results, The 33 surface samples are divided into four pollen zones (forest, meadow steppe, typical steppe and desert steppe) corresponding to major vegetation types in the NECT. Variations in pollen ratios of fern/herb (F/H), Artemisia/Chenopodiaceae (A/C) and arboreal pollen/non-arboreal pollen (AP/NAP) represent the vegetation and precipitation gradient along the NECT. DCA and RDA analyses suggest that surface pollen assemblages are significantly influenced by the precipitation gradient. Changes in the abundance of Chenopodiaceae pollen are related to both human activities and precipitation. Main conclusions, Surface pollen assemblages, fossil pollen records, archaeological evidence and historical documents in northern China show that a large increase of Chenopodiaceae pollen indicates human-caused vegetation degradation in sandy habitats. The A/C ratio is a good indicator of climatic aridity, but should be used in conjunction with multiple proxies of human activities and climate change in the pollen-based reconstruction of anthropogenic biomes. [source] Large-scale pattern of biomass partitioning across China's grasslandsGLOBAL ECOLOGY, Issue 2 2010Yuanhe Yang ABSTRACT Aim, To investigate large-scale patterns of above-ground and below-ground biomass partitioning in grassland ecosystems and to test the isometric theory at the community level. Location, Northern China, in diverse grassland types spanning temperate grasslands in arid and semi-arid regions to alpine grasslands on the Tibetan Plateau. Methods, We investigated above-ground and below-ground biomass in China's grasslands by conducting five consecutive sampling campaigns across the northern part of the country during 2001,05. We then documented the root : shoot ratio (R/S) and its relationship with climatic factors for China's grasslands. We further explored relationships between above-ground and below-ground biomass across different grassland types. Results, Our results indicated that the overall R/S of China's grasslands was larger than the global average (6.3 vs. 3.7). The R/S for China's grasslands did not show any significant trend with either mean annual temperature or mean annual precipitation. Above-ground biomass was nearly proportional to below-ground biomass with a scaling exponent (the slope of log,log linear relationship between above-ground and below-ground biomass) of 1.02 across various grassland types. The slope did not differ significantly between temperate and alpine grasslands or between steppe and meadow. Main conclusions, Our findings support the isometric theory of above-ground and below-ground biomass partitioning, and suggest that above-ground biomass scales isometrically with below-ground biomass at the community level. [source] What caused the mid-Holocene forest decline on the eastern Tibet-Qinghai Plateau?GLOBAL ECOLOGY, Issue 2 2010Ulrike Herzschuh ABSTRACT Aim, Atmospheric CO2 concentrations depend, in part, on the amount of biomass locked up in terrestrial vegetation. Information on the causes of a broad-scale vegetation transition and associated loss of biomass is thus of critical interest for understanding global palaeoclimatic changes. Pollen records from the north-eastern Tibet-Qinghai Plateau reveal a dramatic and extensive forest decline beginning c. 6000 cal. yr bp. The aim of this study is to elucidate the causes of this regional-scale change from high-biomass forest to low-biomass steppe on the Tibet-Qinghai Plateau during the second half of the Holocene. Location, Our study focuses on the north-eastern Tibet-Qinghai Plateau. Stratigraphical data used are from Qinghai Lake (3200 m a.s.l., 36°32,,37°15, N, 99°36,,100°47, E). Methods, We apply a modern pollen-precipitation transfer function from the eastern and north-eastern Tibet-Qinghai Plateau to fossil pollen spectra from Qinghai Lake to reconstruct annual precipitation changes during the Holocene. The reconstructions are compared to a stable oxygen-isotope record from the same sediment core and to results from two transient climate model simulations. Results, The pollen-based precipitation reconstruction covering the Holocene parallels moisture changes inferred from the stable oxygen-isotope record. Furthermore, these results are in close agreement with simulated model-based past annual precipitation changes. Main conclusions, In the light of these data and the model results, we conclude that it is not necessary to attribute the broad-scale forest decline to human activity. Climate change as a result of changes in the intensity of the East Asian Summer Monsoon in the mid-Holocene is the most parsimonious explanation for the widespread forest decline on the Tibet-Qinghai Plateau. Moreover, climate feedback from a reduced forest cover accentuates increasingly drier conditions in the area, indicating complex vegetation,climate interactions during this major ecological change. [source] Plant species richness in continental southern Siberia: effects of pH and climate in the context of the species pool hypothesisGLOBAL ECOLOGY, Issue 5 2007Milan Chytrý ABSTRACT Aim, Many high-latitude floras contain more calcicole than calcifuge vascular plant species. The species pool hypothesis explains this pattern through an historical abundance of high-pH soils in the Pleistocene and an associated opportunity for the evolutionary accumulation of calcicoles. To obtain insights into the history of calcicole/calcifuge patterns, we studied species richness,pH,climate relationships across a climatic gradient, which included cool and dry landscapes resembling the Pleistocene environments of northern Eurasia. Location, Western Sayan Mountains, southern Siberia. Methods, Vegetation and environmental variables were sampled at steppe, forest and tundra sites varying in climate and soil pH, which ranged from 3.7 to 8.6. Species richness was related to pH and other variables using linear models and regression trees. Results, Species richness is higher in areas with warmer winters and at medium altitudes that are warmer than the mountains and wetter than the lowlands. In treeless vegetation, the species richness,pH relationship is unimodal. In tundra vegetation, which occurs on low-pH soils, richness increases with pH, but it decreases in steppes, which have high-pH soils. In forests, where soils are more acidic than in the open landscape, the species richness,pH relationship is monotonic positive. Most species occur on soils with a pH of 6,7. Main conclusions, Soil pH in continental southern Siberia is strongly negatively correlated with precipitation, and species richness is determined by the opposite effects of these two variables. Species richness increases with pH until the soil is very dry. In dry soils, pH is high but species richness decreases due to drought stress. Thus, the species richness,pH relationship is unimodal in treeless vegetation. Trees do not grow on the driest soils, which results in a positive species richness,pH relationship in forests. If modern species richness resulted mainly from the species pool effects, it would suggest that historically common habitats had moderate precipitation and slightly acidic to neutral soils. [source] Climate change and grasslands through the ages: an overviewGRASS & FORAGE SCIENCE, Issue 2 2007L. 't Mannetje Summary Change from cool to warm temperatures and vice versa have occurred throughout geological time. During the Jurassic and Cretaceous periods (206,65 million years ago, Ma) the climate was more uniformly warm and moist than at present and tropical rainforests were widespread. Grasses evolved during the Jurassic period and they expanded greatly as the climate differentiated with reduced rainfall and temperatures. C4 -grasses probably arose during the Oligocene period (24,35 Ma). During the Miocene period (23·8,5·3 Ma) grasslands expanded into huge areas (e.g. prairies in the USA, steppe in Eurasia, and pampas and llanos in South America). During the Quaternary period (1·8 Ma till now) some twenty-two different ice ages with periodicities of about 100 000 years occurred. Eighteen-thousand years ago, north-western Europe had a polar climate with tundra vegetation and the Mediterranean region was covered by steppe. During that time Amazonia was so dry that it was covered in extensive areas of savanna and the Sahara expanded rapidly. Only in the last 10 000 years has a closed rainforest covered the Amazonian region again. However, 9000 years ago a brief period of global warming caused excessive rains, which caused the sea and river levels to rise in north-western Europe with tremendous loss of life. The present period of extreme dryness in the Sahara only started some 5000 years ago and then the desert expanded rapidly into the Sahel. Before that the Sahara was covered by steppe. Global warming took place between about ad 900 and about ad 1200 or 1300 just before the Little Ice Age (1550,1700 ad). The article concludes with a description of temperature and vegetation changes that are occurring in Europe at present. It is predicted that C4 -grasses, which are already present in southern Europe, will further expand but that, in the short term, land abandonment will have much more deleterious effects than temperature change due to increased wild fires, loss of biodiversity and desertification. [source] The genetic diversity of perennial Leymus chinensis originating from ChinaGRASS & FORAGE SCIENCE, Issue 1 2007Z. P. Liu Summary Leymus chinensis is an economically and ecologically important grass that is widely distributed across eastern areas of the Eurasian steppe. A better knowledge of genetic diversity of L. chinensis could be valuable in the efficient utilization, conservation and management of germplasm collections. Genetic diversity in thirty-seven morphological characters of 293 accessions was assessed in three successive years. Based on these qualitative and quantitative characters, the genetic diversity indices (Shannon indices) of traits and geographical populations were estimated, and a principal coordinates analysis and a path analysis were undertaken. Compared with the yellow-green type of L. chinensis, the grey-green type had significantly (P < 0·05) more genetic diversity. In addition, the path analysis suggested that the combined effects of genetic diversity and vegetative traits could explain 0·206 of the total variance in plant reproductive traits. The highest Shannon genetic diversity index of accessions (H = 2·252) was observed in accessions from the region of longitude of 124,128°E, suggesting the most abundant germplasm of L. chinensis in this region. [source] Demographic variation and biomass allocation of Agropyron cristatum grown on steppe and dune sites in the Hunshandake Desert, North ChinaGRASS & FORAGE SCIENCE, Issue 1 2005R. Z. Wang Abstract Demographic and biomass allocation patterns of Agropyron cristatum were measured on steppe and dune sites in the Hunshandake desert of North China in 2001 and 2002. Total plant population, reproductive shoot densities and its differentiation rates were significantly higher in the steppe sites in both years. Plant heights for both vegetative and reproductive shoots were greater in the year with the higher rainfall. The dune sites had a higher biomass allocation to vegetative shoots and roots, while the steppe sites had a higher biomass allocation to reproductive shoots and seed production. It is suggested that the population demography and biomass allocation of the species responded to the differences in the soil variables in the steppe and dune sites. [source] Vegetation of the Stipa loess steppe in Ningxia (northern China) in relation to grazing intensityGRASSLAND SCIENCE, Issue 3 2007Yingzhong Xie Abstract Large areas of northern and western China are covered with steppe vegetation which is grazed with different intensities. Areas which are not or only slightly grazed are rare and mostly exist in nature reserves. We investigated floristic composition and species diversity of the steppe vegetation of the autonomous region Ningxia Hui with emphasis on the Yunwushan dry steppe nature reserve in relation to grazing intensity; all 77 plots were analyzed using ordination techniques, and the relationship between the variation in species composition and environmental conditions was analyzed. In Ningxia, two types of loess grass steppes prevail: in areas with lower altitudes from approximately 1650,1950 m a.s.l., the Stipa bungeana steppe occurs whereas the Stipa grandis steppe is typical for higher altitudes between 1900 and 2100 m a.s.l. For both vegetation types, three subtypes can be distinguished according to different levels of grazing intensity. With increasing grazing intensity, the number of plant species per plot strongly declines and the typical steppe vegetation types are dominated by Artemisia species (A. frigida in the Stipa grandis community and A. sacrorum in the Stipa bungeana community). In contrast to studies carried out in neighboring areas, in our study area the Artemisia steppes do not represent the final state of degradation. Instead, Convolvulus ammanii dominates in heavily grazed areas of both Stipa steppe types. Detrended correspondence analysis confirms the classification and shows that the heavily grazed subtypes of the Stipa grandis and the Stipa bungeana steppe with Convolvulus ammanii are more similar to each other than the other vegetation types. The first axis of the detrended correspondence analysis is strongly related to the grazing intensity (r = 0.93). Our results show that grazing intensity is an important factor for the differentiation of the steppe grasslands and a severe threat to species diversity. [source] The influence of climatic change and human activity on erosion processes in sub-arid watersheds in southern East SiberiaHYDROLOGICAL PROCESSES, Issue 16 2003Leonid M. Korytny Abstract A LUCIFS model variant is presented that represents the influence of climate and land use change on fluvial systems. The study considers trends of climatic characteristics (air temperature, annual precipitation totals, rainfall erosion index, aridity and continentality coefficients) for the steppe and partially wooded steppe watersheds of the south of East Siberia (the Yenisey River macro-watershed). It also describes the influence of these characteristics on erosion processes, one indicator of which is the suspended sediment yield. Changes in the river network structure (the order of rivers, lengths, etc.) as a result of agricultural activity during the 20th century are investigated by means of analysis of maps of different dates for one of the watersheds, that of the Selenga River, the biggest tributary of Lake Baikal. The study reveals an increase of erosion process intensity in the first two-thirds of the century in the Selenga River watershed and a reduction of this intensity in the last third of the century, both in the Selenga River watershed and in most of the other watersheds of the study area. Copyright © 2003 John Wiley & Sons, Ltd. [source] Hydrological regime analysis of the Selenge River basin, MongoliaHYDROLOGICAL PROCESSES, Issue 14 2003X. Ma Abstract Arid and semi-arid regions are very vulnerable to environmental changes. Climate change studies indicate that the environment in such areas will steadily deteriorate with global warming; inland lakes will shrink and desert areas will expand. Mongolia is a landlocked country in north-central Asia that contains a unique ecological system consisting of taiga, steppe, and desert from north to south. The Selenge River basin (280 000 km2) in northern Mongolia is a semi-arid region underlain by permafrost, between latitudes 46 and 52°N, and longitudes 96 and 109°E. The issue of sustainable development of the basin is very important owing to its limited natural resources, including fresh water, forest, and rangeland. To examine the water cycle processes in the basin, a hydrological analysis was carried out using a simple scheme for the interaction between the land surface and atmosphere (big-leaf model) coupled to a hydrological model for the period 1988,92 to estimate the hydrological regime of the basin. Annual precipitation in this period averaged 298 mm, ranging from 212 to 352 mm at a 1 ° × 1 ° resolution based on data from 10 gauges, and the estimated annual evapotranspiration averaged 241 mm, ranging between 153 and 300 mm. This indicates that evapotranspiration accounts for the overwhelming majority of the annual precipitation, averaging 81% and ranging between 64 and 96%. The annual potential evapotranspiration in the basin averaged 2009 mm; the ratio of evapotranspiration (actual to potential evapotranspiration) was 0·12 and the wetness index (annual precipitation to potential evapotranspiration) was 0·15. Copyright © 2003 John Wiley & Sons, Ltd. [source] Effects of temperature on the post-diapause embryonic development and the hatching time in three grasshopper species (Orth., Acrididae)JOURNAL OF APPLIED ENTOMOLOGY, Issue 2 2004S. G. Hao Abstract: A study was conducted to determine the effects of six constant temperatures (15, 20, 25, 30, 35 and 40°C) on the post-diapause embryonic development and the hatching time in three grasshopper species ,Omocestus haemorrhoidalis (Charp.), Calliptamus abbreviatus Ikonn. and Chorthippus fallax (Zub.) , from the Inner Mongolian steppe. The results indicate that the species differ in the developmental rates, survival curves and cumulative hatching probabilities. The eggs of O. haemorrhoidalis had the fastest developmental rate with a low developmental threshold temperature of 9.9°C and the sum of effective temperature (SET) 211.2 degree-days (DD). The corresponding values were 10.9°C and 210.6 DD for C. abbreviatus, 10.5°C and 240.2 DD for Ch. fallax respectively. The SET at which 50% of post-diapause eggs hatched were 252.0 DD for O. haemorrhoidalis, 262.8 DD for C. abbreviatus, and 273.3 DD for Ch. fallax. The predicted maximal hatch ability of O. haemorrhoidalis (91.17%), C. abbreviatus (75.67%) and Ch. fallax (94.07%) occurred at 23.7, 29.0 and 31.3°C, respectively. The thermal death points of each species were reached at 43.3, 45.0 and 48.6°C. The optimal temperature ranges were 12.2,35.2°C for O. haemorrhoidalis, 21.7,36.3°C for C. abbreviatus and 20.9,41.7°C for Ch. fallax respectively. These results suggest that O. haemorrhoidalis adapt to hatch at a lower temperature range, C. abbreviatus adapt to mid-temperature range, while Ch. fallax adapt to hatch at a higher temperature range. Although the SET of Ch. fallax is more than that of the other two species, it is not sufficient to explain the hatching sequence of the species in springtime. The results also indicate that Ch. fallax and O. haemorrhoidalis have wider adaptive temperature range than C. abbreviatus. [source] Infectivity of soilborne Frankia and mycorrhizae in Discaria trinervis along a vegetation gradient in Patagonian soilJOURNAL OF BASIC MICROBIOLOGY, Issue 4 2006Eugenia Esther Chaia The infective capacities of the nitrogen fixing Actinomycete Frankia and arbuscular mycorrhizal fungi from soils near watercourses, along a vegetation gradient, were studied using plant bioassays. Frankia and arbuscular mycorrhizas capable of infecting Discaria trinervis were found at seventeen sites sampled. More specific enumeration of the infective capacities of both microorganisms in relation to environmental factors was performed in seven representative soils of the analysed vegetation zones (rainforest, xeric forest and steppe) using the most probable number method. The highest nodulation capacities ranged from 340 infective units g,1 soil, in a steppe marsh devoid of actinorhizas, to 61 in a coastal actinorhizal scrub (in xeric forest). The highest number of infective mycorrhizal units , also found in marsh , was 145. In general, rainforest soils had the lowest values for both microorganisms. Infective units of Frankia and arbuscular mycorrhizal fungi in soil were positively correlated (r = 0.89, P < 0.05). Both soilborne symbionts showed the highest infective capacity in semi-arid conditions nearby watercourse and at the valley bottom location. Tripartite symbiosis was effective in plants inoculated with steppe and xeric forest soils and plants inoculated with Frankia BCU110501 and Glomus mosseae. Interaction between both symbionts and influence of environmental conditions, in general, would contribute to define comparable trends of their infective capacities. (© 2006 WILEY-VCH Verlag GmbH & Co. KGaA, Weinheim) [source] The ,oriental' component of the Balkan flora: evidence of presence on the Thracian Plain during the Weichselian late-glacialJOURNAL OF BIOGEOGRAPHY, Issue 5 2008E. K. Magyari Abstract Aim, To obtain palaeobotanical evidence enabling evaluation of the viability of the hypothesis that the ,oriental' element of the Balkan flora reached south-east Europe from Turkey prior to the Holocene, probably via the Thracian Plain during a late Quaternary glacial stage but no later than the late Weichselian. Location, Ezero wetland, northern Thracian Plain, Bulgaria. Methods, We undertook analyses of pollen and microspores, plant macrofossils, wood fragments and molluscs recovered from sediments deposited in the Ezero wetland during the late Weichselian and Weichselian late-glacial. Sediment chronology was determined using radiocarbon age estimates. Results, Six metres of sediments were recovered from the basin, of which the lower 3 m, extending from c. 15,450 cal yr bp to the early Allerød, was analysed. A major hiatus occurred after c. 13,900 cal yr bp, the overlying sediments being of late Holocene age. Palaeobotanical evidence indicates predominantly open vegetation during the Weichselian late-glacial, although macrofossil remains of woody taxa demonstrate the local presence of patches of wooded steppe and gallery forest. Changes in the composition of the steppe vegetation, and in the nature of the sediments deposited in the basin, indicate changes in climatic conditions, especially in the hydrological regime and in the moisture available to vegetation. After an initially relatively moister phase, the final centuries of the late Weichselian were drier, as was a short interval that may correlate with the Older Dryas. Moister conditions characterize intervals corresponding to the Bølling and Allerød sub-units of the Weichselian late-glacial interstadial. Although the pollen evidence is thus consistent with that from previous studies of this period in south-east Europe and south-west Asia, indicating predominantly open steppe vegetation, the macrofossil evidence indicates the persistent local presence of woody taxa. The woody taxa recorded include Celtis tournefortii -type and Juniperus cf. J. excelsa, two taxa today characteristic of the wooded steppes of Anatolia and members of the ,oriental' element of the southern Balkan flora, as well as Rosaceae Subfams. Maloideae and Prunoideae, Alnus and Fraxinus. Main conclusions, The late Weichselian vegetation of the northern Thracian Plain included patches of wooded steppe that supported members of the ,oriental' element of the modern Balkan flora. The presence of such taxa renders viable the hypothesis that they could have reached south-east Europe from Turkey via the Thracian Plain during glacial times. Such hypotheses in historical biogeography can be evaluated critically using the evidence obtained from plant macrofossil analyses in combination with that from pollen analysis. [source] The Branchiopoda (Crustacea: Anomopoda, Ctenopoda and Cyclestherida) of the rain forests of Cameroon, West Africa: low abundances, few endemics and a boreal,tropical disjunctionJOURNAL OF BIOGEOGRAPHY, Issue 9 2005George Y. Chiambeng Abstract Aim, We provide the first in-depth study of the Branchiopoda of the rain forests of Cameroon and also of the African continent. Location, Surface water environments, Cameroon. Methods, Qualitative plankton samples were collected in all types of surface water environments present, ranging from big lakes to water collected in rock crevices or fallen fruit cavities. A tow or hand-held plankton net of mesh size 100 ,m was used, and water volumes filtered were at least several m3 in large water bodies, or half to whole water volume in small water bodies. Results, We recorded 61 species (53 first records for the country), based on 700+ samples collected between September 1998 and March 2002. Anomopoda (92%) was the dominant order, followed by Ctenopoda (6.5%) and Cyclestherida (1.5%). Chydoridae (67%) was the most speciose family followed by Macrothricidae (6.5%) and Daphniidae (5%). Alona (11%) was the dominant genus followed by Chydorus (10%) and Pleuroxus (8%). Several species of Chydorinae, especially of the genus Pleuroxus, are shared with continental Eurasia,North America, but are absent from the Mediterranean and desert,steppe,savanna zones of Africa (boreal,tropical disjunction). Daphnia was absent, as in most tropical lowlands. No single species was really abundant, and a majority were rare to very rare, and of restricted occurrence within the rain forest patches. Comparing Africa, South America and Southeast Asia, we found a current total of 196 species for the combined rain forest areas, out of a world total of 500+ species. Systematic trends in richness at three taxonomic levels were the same for all continents: Anomopoda,Ctenopoda,Cyclestherida at ordinal level, Chydoridae,Daphniidae,Macrothricidae,Sididae at family level and Alona,Chydorus,Macrothrix,Diaphanosoma at genus level. Southeast Asia was richest (111 species, 14 endemics) with South America a close second (110 species, 27 endemics). Africa was the most species-poor (95 species, of which only 5 are endemics). Main conclusions, We hypothesize that the post-Miocene cooling and aridization of the world climate hit the freshwater biota of Africa particularly hard, with more extinction here than elsewhere, and little recolonization. Most extinction occurred in the savanna-desert belt, and eight disjunct boreal species (four Pleuroxus, Picripleuroxus laevis, Kurzia latissima, Alonella exigua, and Monospilus dispar) survive morphologically unchanged since pre-Pleistocene times in the Cameroon rain forest. Slow evolution thus appears typical of these cyclic parthenogenetic branchiopods in which sexual recombination occurs only at intervals. Illustrative of the same slow evolution is the fact that the two endemic cladocerans of Cameroon (Nicsmirnovius camerounensis and Bryospilus africanus) belong to tropicopolitan genera of Gondwanan age. [source] Community organization and species richness of ants (Hymenoptera/Formicidae) in Mongolia along an ecological gradient from steppe to Gobi desertJOURNAL OF BIOGEOGRAPHY, Issue 12 2003Martin Pfeiffer Abstract Aim, Ants (Hymenoptera/Formicidae) have strong influences on ecosystems especially in arid regions. However, little is known about ants of the vast steppe and desert regions of Central Asia. Here we provide the first comprehensive study of ant communities in Mongolia, conducted along a north-to-south gradient in climate. We examined ants' distribution patterns, assessed the impact of climatic parameters on community structure and species diversity and investigated the influence of the corresponding communities of plants. Location, Mongolia (Central Asia). Methods, We observed 31,956 ants at seed baits at 11 study sites along a transect from steppe to Gobi desert for which we attained meteorological data (mean yearly precipitation: 197 to 84 mm). Extra sampling was conducted at sugar and protein baits and by the inspection of different microhabitats. Vegetation patterns of each plot were recorded. Statistical evaluation comprised ordination and correlation. Results, We observed 15 species of ants at seed baits. Three faunal complexes of ants could be distinguished by detrended correspondence analysis (DCA): (1) in steppe baits were dominated by Formica - and Myrmica -species, (2) in semi desert we found mostly species of Tetramorium, Myrmica, Proformica, Plagiolepis, and Leptothorax, and (3) in desert Cataglyphis aenescens and Messor aciculatus dominated, and Lasius was exclusively found there. Another 11 rare ant species were sampled by hand and at sugar baits. Altogether five ant species were new to the Mongolian fauna: Cardiocondyla koshewnikovi, Myrmica koreana, Myrmica pisarskii, Polyergus nigerrimus, and Proformica kaszabi. Assignment of taxa to functional groups showed that in steppe cold climate specialists dominated, in semi desert we found mainly opportunists, and in desert hot climate specialists. Several functional groups know from arid zones in other parts of the world were missing. In desert certain species were highly dominant. First DCA scores of ant- and plant-communities were highly correlated with each other and with climatic parameters. While plant species diversity was positively correlated with increasing northern latitude, ant diversity and ant species richness were not correlated with latitude and responded neither to precipitation, nor to any other climatic parameter. Semi desert was a transition zone between steppe and desert, with high species richness. Ant genus composition of the ecotone overlapped with both other regions. However, beta diversity between pairs of plots within this zone was low, indicating a small-scale mosaic pattern. Main conclusions, The ant communities in the Mongolian steppe and desert zones were strongly influenced by low temperatures and differed in many aspects from the ant fauna in other arid ecosystems, especially in terms of species richness, diversity of feeding guilds, and richness of functional groups. [source] The Middle Asian Element in the Southern Rocky Mountain Flora of the western United States: a critical biogeographical reviewJOURNAL OF BIOGEOGRAPHY, Issue 5 2003William A. Weber Abstract Aim, Presentation of an hypothesis suggesting that the extraordinarily similarity of the Russian Altai and the American Southern Rocky Mountain Flora represents an Oroboreal Flora; that had to have had an essential continuity across the northern part of the world in the Tertiary period, constituting a highland and steppe component of the better-known Arcto-Tertiary Flora of eastern and far-western North America and eastern Asia. Location, North America and Middle (Altai) Asia. Methods, Summarization of the author's field and herbarium studies of whole floras over a period of over 60 years, consisting of successive specializations in vascular plants, lichens, and bryophytes. Main conclusions, (1) The modern alpine and associated marginal steppe and montane floras contain taxa of Tertiary age. (2) The floras of the southern mountains antedate those of the present-day Arctic. (3) The Middle Asiatic and the North American floras once enjoyed a contiguous existence over a broad area involving connections between North America and Asia across the North Pole by way of Greenland. Their present disjunctions are products of extinction and attrition of ranges, not of long-distance migration or dispersal mechanisms. (4) North-eastern North American disjunctions of so-called Cordilleran species (the Nunatak hypothesis) need not require explanations involving long-distance dispersal or migration, but represent relictual populations of the once widely distributed Oroboreal flora. [source] Modern pollen precipitation from an elevational transect in central Jordan and its relationship to vegetationJOURNAL OF BIOGEOGRAPHY, Issue 10 2001Caroline P. Davies Aim To explore the relationship between modern pollen precipitation and vegetation patterns in an arid region of the Middle East. Location Data come from the central Jordan Rift from 1700 m elevation in the highlands to 300 m below sea level in the Dead Sea basin. Methods Modern pollen samples and descriptive vegetation data were collected from twenty-one locations at 100 m elevational intervals from the highest elevations on the eastern side of the rift valley, where woodlands grow, to the lowest elevation desert on earth, characterized by drought and salt tolerant plants. Pollen percentage data from each vegetation zone are compared descriptively and numerically using cluster and Principal Components Analyses (PCA). Results Three major vegetation zones: woodland, shrub steppe, and desert scrub, are identified by their dominant plant species. The widely spaced tree, Quercus calliprinos Webb, defines the Quercus L. woodland that grows above 1500 m elevation. The shrub steppe can be divided into two subzones found between about 1500 and 900 m elevation: Artemisia herba-alba Asso shrub steppe and Artemisia L. shrub steppe with Juniperus phoenica L. Desert scrub dominates the lower elevation landscape with Hammada salicornia (Moq.) Iljin the dominant shrub between 900 and 200 m and Haloxylon persicum Bge. found below 200 m elevation. Pollen spectra reflect the elevational vegetation zones. In particular, Quercus L., Juniperus L. and Tamarix L. pollen are abundant where the trees grow. Highly variable amounts of non-arboreal pollen taxa , primarily Artemisia L. and Chenopodiaceae Vent. , differentiate shrub steppe from desert scrub vegetation. Cluster and PCA of pollen data support the qualitative vegetation zonation. Main conclusions The main vegetation zones along the Jordan Rift from 1700 to ,300 m elevation can be distinguished by their modern pollen precipitation. Open vegetation types, in particular, can be recognized by their pollen spectra. High amounts of Artemisia L. pollen distinguish the moister upper elevations where Artemisia L. steppe grows. In contrast, greater amounts of Chenopodiaceae Vent. pollen characterize the drier, lower elevation deserts. [source] What is the origin of the carabid beetle fauna of dry, anthropogenic habitats in western Europe?JOURNAL OF BIOGEOGRAPHY, Issue 4 2000Johan Andersen Abstract Aim To study the origin of the carabid beetle fauna of dry, open anthropogenic habitats. Location Various parts of Europe. Methods Collecting by hand in habitats influenced by humans (arable land, sand pits, ruderal places, road verges, meadows) and naturally open habitats such as talus, supralittoral sand dunes, alvar as well as littoral habitats in south and central Scandinavia. Studies made in other parts of Europe were included in the discussion. Results and conclusion The material comprises about 37,700 specimens. About 8% of the Scandinavian species occurring in anthropogenic habitats may have a littoral origin, about 10% may originate from forests whereas about 9% are euryoecious. A majority of the species (about 66%) originate from primarily open, dry habitats and many of them constitute a steppe or steppe-like element. They invaded the bare landscape soon after the deglaciation and probably survived in naturally open habitats (supralittoral sand dunes, talus, alvar, old riverbeds) during the postglacial warm period, but some species have a recent origin in Scandinavia. [source] Mid-Holocene and glacial-maximum vegetation geography of the northern continents and AfricaJOURNAL OF BIOGEOGRAPHY, Issue 3 2000I. Colin Prentice Abstract BIOME 6000 is an international project to map vegetation globally at mid-Holocene (6000 14C yr bp) and last glacial maximum (LGM, 18,000 14C yr bp), with a view to evaluating coupled climate-biosphere model results. Primary palaeoecological data are assigned to biomes using an explicit algorithm based on plant functional types. This paper introduces the second Special Feature on BIOME 6000. Site-based global biome maps are shown with data from North America, Eurasia (except South and Southeast Asia) and Africa at both time periods. A map based on surface samples shows the method's skill in reconstructing present-day biomes. Cold and dry conditions at LGM favoured extensive tundra and steppe. These biomes intergraded in northern Eurasia. Northern hemisphere forest biomes were displaced southward. Boreal evergreen forests (taiga) and temperate deciduous forests were fragmented, while European and East Asian steppes were greatly extended. Tropical moist forests (i.e. tropical rain forest and tropical seasonal forest) in Africa were reduced. In south-western North America, desert and steppe were replaced by open conifer woodland, opposite to the general arid trend but consistent with modelled southward displacement of the jet stream. The Arctic forest limit was shifted slighly north at 6000 14C yr bp in some sectors, but not in all. Northern temperate forest zones were generally shifted greater distances north. Warmer winters as well as summers in several regions are required to explain these shifts. Temperate deciduous forests in Europe were greatly extended, into the Mediterranean region as well as to the north. Steppe encroached on forest biomes in interior North America, but not in central Asia. Enhanced monsoons extended forest biomes in China inland and Sahelian vegetation into the Sahara while the African tropical rain forest was also reduced, consistent with a modelled northward shift of the ITCZ and a more seasonal climate in the equatorial zone. Palaeobiome maps show the outcome of separate, independent migrations of plant taxa in response to climate change. The average composition of biomes at LGM was often markedly different from today. Refugia for the temperate deciduous and tropical rain forest biomes may have existed offshore at LGM, but their characteristic taxa also persisted as components of other biomes. Examples include temperate deciduous trees that survived in cool mixed forest in eastern Europe, and tropical evergreen trees that survived in tropical seasonal forest in Africa. The sequence of biome shifts during a glacial-interglacial cycle may help account for some disjunct distributions of plant taxa. For example, the now-arid Saharan mountains may have linked Mediterranean and African tropical montane floras during enhanced monsoon regimes. Major changes in physical land-surface conditions, shown by the palaeobiome data, have implications for the global climate. The data can be used directly to evaluate the output of coupled atmosphere-biosphere models. The data could also be objectively generalized to yield realistic gridded land-surface maps, for use in sensitivity experiments with atmospheric models. Recent analyses of vegetation-climate feedbacks have focused on the hypothesized positive feedback effects of climate-induced vegetation changes in the Sahara/Sahel region and the Arctic during the mid-Holocene. However, a far wider spectrum of interactions potentially exists and could be investigated, using these data, both for 6000 14C yr bp and for the LGM. [source] Extending point pattern analysis for objects of finite size and irregular shapeJOURNAL OF ECOLOGY, Issue 4 2006THORSTEN WIEGAND Summary 1We use a grid- and simulation-based approach to extend point pattern analysis to deal with plants of finite size and irregular shape, and compare the results of our approach with that of the conventional point approximation. The plants are approximated by using an underlying grid and may occupy several adjacent grid cells depending on their size and shape. Null models correspond to that of point pattern analysis but need to be modified to account for the finite size and irregular shape of plants. 2We use a mapped area of a grass-shrub steppe in semi-arid Patagonia, Argentina, to show that the shrub community is essentially randomly structured, but that shrubs facilitate grasses in their immediate neighbourhood. 3The occurrence of this random spatial structure provides important new information on the biology of shrub populations. In general, previous data from semi-arid and arid ecosystems have shown that adult shrubs tend to show over-dispersed patterns, whereas juveniles are clumped. 4We find that the point approximation may produce misleading results (i) if plant size varies greatly, (ii) if the scale of interest is of the same order of magnitude as the size of the plants, and (iii) if the plants of a given pattern are constrained through competition for space by the presence of other plants. The point approximation worked well in all other cases, but usually depicted weaker significant effects than when the size and shape of plants were taken into account. 5Our approach to quantifying small-scale spatial patterns in plant communities has broad applications, including the study of facilitation and competition. Ecologists will be able to use the software available to take advantage of these methods. [source] | ||||||||||||||||||||||||||||||||||