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Rocky Mountains (rocky + mountain)
Kinds of Rocky Mountains Selected AbstractsDEVONIAN CARBONATES OF THE NIGEL PEAK AREA, ROCKY MOUNTAINS, CANADA: A FOSSIL PETROLEUM SYSTEM?JOURNAL OF PETROLEUM GEOLOGY, Issue 3 2008J. Köster In this study we report on Devonian (Frasnian , Famennian) limestones and dolostones exposed near Nigel Peak in the Main Ranges of the Canadian Rocky Mountains. These carbonates are a proximal facies of the Southesk-Cairn Carbonate Complex. The investigated strata are stratigraphically equivalent to the oil- and gas bearing Nisku Formation in the subsurface of the Western Canada Sedimentary Basin, about 300 km to the east. The rocks were investigated by polarisation and cathodoluminescence microscopy, total organic carbon analysis, Rock-Eval pyrolysis, solid bitumen reflectance measurements, gas chromatography and fluid inclusion analysis. Thin section analyses showed that silt-grade quartz and saddle dolomite increase upward from the base of the stratigraphic section, and that porosities are generally low. This is due to reduction of pore space due to early cementation and extensive dolomitization. Cathodoluminescence identified up to four generations of calcite cements. TOC values ranged from 0.2 to 2.4 %. Rock-Eval pyrolysis of carbonate samples resulted in measurable S1 peaks but not S2 peaks, indicating that there was no residual petroleum generation potential. Organic petrographic analyses identified dispersed kerogen and migrabitumen, and calculated vitrinite reflectance values were around 4 % on average which implies peak temperatures of 234,262 °C (due to deep burial) or 309,352 °C (due to short term hydrothermal heating). Fluid inclusion data indicates at least one pulse of hot fluids with elevated homogenization temperatures of > 300 °C, and this may explain the high thermal maturity of the studied rocks. [source] Growth of Creative Occupations in U.S. Metropolitan Areas: A Shift-Share AnalysisGROWTH AND CHANGE, Issue 3 2006TODD M. GABE ABSTRACT This article uses a shift-share model to investigate the growth of creative occupations in U.S. metropolitan areas during the 1990s. Empirical findings indicate that the performance of the creative economy varied widely across the U.S., and that the highest competitive growth rates of the creative workforce occurred in the Rocky Mountain, Southeast, and Southwest regions. Further analysis focused on whether high competitive workforce growth between 1990 and 2000 translated into high competitive employment growth from 1999 to 2003. The results show that many of the areas with the highest competitive growth rates of creative economy employment from 1999 to 2003 were some of the weakest regions in terms of creative workforce growth during the 1990s. This raises questions about the extent to which jobs follow people in the creative economy, and suggests that an initial strong presence in the creative economy is not a prerequisite for future growth. [source] Microsatellite analysis of North American wapiti (Cervus elaphus) populationsMOLECULAR ECOLOGY, Issue 10 2000Renee O. Polziehn Abstract Eleven populations of wapiti (Cervus elaphus) were analysed for genetic diversity using 12 microsatellite loci. Samples were taken from Vancouver Island, British Columbia; Burwash and French River herds in Ontario; Ya Ha Tinda Ranch, Alberta; and Banff, Elk Island, Jasper, Kootenay, Riding Mountain, Yellowstone and Yoho National Parks. Overall, wapiti populations have on average three to four alleles per locus and an average expected heterozygosity that ranged from 25.75 to 52.85%. The greatest genetic distances were observed between the Vancouver population and all other populations. Using the assignment test, Roosevelt wapiti (C. e. roosevelti Merriam 1897) assigned only to the Vancouver Island population. The distance and assignment values suggest a divergence of the Roosevelt wapiti from other populations and support the subspecific status for the Vancouver Island population. No evidence was found for the existence of unique Eastern wapiti (C. e. canadensis Erxleben 1777) in the Burwash or French River herds in Ontario. The overlapping distribution of genotypes from indigenous populations from Riding Mountain, Elk Island and Yellowstone National Parks suggests that wapiti were once a continuous population before settlers decimated their numbers. The lack of differentiation between these populations raises questions about the status of Manitoban (C. e.manitobensis Millais 1915) and Rocky Mountain (C. e.nelsoni Bailey 1935) subspecies. [source] Reach-scale channel geometry of a mountain riverEARTH SURFACE PROCESSES AND LANDFORMS, Issue 8 2004Ellen Wohl Abstract Mountain rivers can be subject to strong constraints imposed by changes in gradient and grain size supplied by processes such as glaciation and rockfall. Nonetheless, adjustments in the channel geometry and hydraulics of mountain rivers at the reach scale can produce discernible patterns analogous to those in fully alluvial rivers. Mountain rivers can differ in that imposed reach-scale gradient is an especially important control on reach-scale channel characteristics, as indicated by examination of North St Vrain Creek in Colorado. North St Vrain Creek drains 250 km2 of the Rocky Mountains. We used 25 study reaches within the basin to examine controls on reach-scale channel geometry. Variables measured included channel geometry, large woody debris, grain size, and mean velocity. Drainage area at the study reaches ranged from 2·2 to 245 km2, and gradient from 0·013 to 0·147 m m,1. We examined correlations among (1) potential reach-scale response variables describing channel bankfull dimension and shape, hydraulics, bedform wavelength and amplitude, grain size, ,ow resistance, standard deviation of hydraulic radius, and volume of large woody debris, and (2) potential control variables that change progressively downstream (drainage area, discharge) or that are likely to re,ect a reach-speci,c control (bed gradient). We tested the hypothesis that response variables correlate most strongly with local bed gradient because of the segmented nature of mountain channels. Results from simple linear regression analyses indicate that most response variables correlate best with gradient, although channel width and width/depth ratio correlate best with discharge. Multiple regression analyses using Mallow's Cp selection criterion and log-transformation of all variables produced similar results in that most response variables correlate strongly with gradient. These results suggest that the hypothesis is partially supported: channel bed gradient is likely to be a good predictor for many reach-scale response variables along mountain rivers, but discharge is also an important predictor for some response variables. Copyright © 2004 John Wiley & Sons, Ltd. [source] Effects of Acer platanoides invasion on understory plant communities and tree regeneration in the northern Rocky MountainsECOGRAPHY, Issue 5 2005Kurt O. Reinhart Quantitative studies are necessary to determine whether invasive plant species displace natives and reduce local biodiversity, or if they increase local biodiversity. Here we describe the effects of invasion by Norway maple Acer platanoides on riparian plant communities and tree regeneration at two different scales (individual tree vs stand scales) in western Montana, USA, using both descriptive and experimental approaches. The three stands differed in community composition with the stand most dominated by A. platanoides invasion being more compositionally homogenous, and less species rich (,67%), species even (,40%), and diverse (,75%) than the two other stands. This sharp decrease in community richness and diversity of the highly invaded stand, relative to the other stands, corresponded with a 28-fold increase in A. platanoides seedlings and saplings. The dramatic difference between stand 1 vs 2 and 3 suggests that A. platanoides invasion is associated with a dramatic change in community composition and local loss of species diversity; however, other unaccounted for differences between stands may be the cause. These whole-stand correlations were corroborated by community patterns under individual A. platanoides trees in a stand with intermediate levels of patchy invasion. At the scale of individual A. platanoides canopies within a matrix of native trees, diversity and richness of species beneath solitary A. platanoides trees declined as the size of the trees increased. These decreases in native community properties corresponded with an increase in the density of A. platanoides seedlings. The effect of A. platanoides at the stand scale was more dramatic than at the individual canopy scale; however, at this smaller scale we only collected data from the stand with intermediate levels of invasion and not from the stand with high levels of invasion. Transplant experiments with tree seedlings demonstrated that A. platanoides seedlings performed better when grown beneath conspecific canopies than under natives, but Populus and Pinus seedlings performed better when grown beneath Populus canopies, the dominant native. Our results indicate that A. platanoides trees suppress most native species, including the regeneration of the natural canopy dominants, but facilitate conspecifics in their understories. [source] Transpiration and stomatal conductance across a steep climate gradient in the southern Rocky MountainsECOHYDROLOGY, Issue 3 2008Nate G. McDowell Abstract Transpiration (E) is regulated over short time periods by stomatal conductance (Gs) and over multi-year periods by tree- and stand-structural factors such as leaf area, height and density, with upper limits ultimately set by climate. We tested the hypothesis that tree structure, stand structure and Gs together regulate E per ground area (Eg) within climatic limits using three sites located across a steep climatic gradient: a low-elevation Juniperus woodland, a mid-elevation Pinus forest and a high-elevation Picea forest. We measured leaf area : sapwood area ratio (Al : As), height and ecosystem sapwood area : ground area ratio (As : Ag) to assess long-term structural adjustments, tree-ring carbon isotope ratios (,13C) to assess seasonal gas exchange, and whole-tree E and Gs to assess short-term regulation. We used a hydraulic model based on Darcy's law to interpret the interactive regulation of Gs and Eg. Common allometric dependencies were found only in the relationship of sapwood area to diameter for pine and spruce; there were strong site differences for allometric relationships of sapwood area to basal area, Al : As and As : Ag. On a sapwood area basis, E decreased with increasing elevation, but this pattern was reversed when E was scaled to the crown using Al : As. Eg was controlled largely by As : Ag, and both Eg and Gs declined from high- to low-elevation sites. Observation-model comparisons of Eg, Gs and ,13C were strongest using the hydraulic model parameterized with precipitation, vapour pressure deficit, Al : As, height, and As : Ag, supporting the concept that climate, Gs, tree- and stand-structure interact to regulate Eg. Copyright © 2008 John Wiley & Sons, Ltd. [source] Persistent organic pollutants in air and vegetation from the canadian rocky mountainsENVIRONMENTAL TOXICOLOGY & CHEMISTRY, Issue 3 2004Deborah A. Davidson Abstract The exchange of chlorinated organic pollutants between air and vegetation in cold, mountain environments was investigated through the extraction of coniferous vegetation and high-volume air samples collected from the Canadian Rocky Mountains during the summers of 1999 and 2000. Concentrations of several compounds in vegetation increased as temperatures decreased, whereas atmospheric concentrations were not related to temperature. Daily cycling of these compounds between air and vegetation as a result of diurnal temperature changes was not observed. Compared with concentrations in vegetation from the Canadian Rocky Mountains, plant samples from the western valley in British Columbia (Canada) showed higher pollutant levels. Chemical partitioning between vegetation and air was not correlated with temperature, indicating that air contamination is governed by long-range transport and not by local revolatilization events. Based on these observations, we show that both deposition at higher altitudes and long-range atmospheric transport influence chemical accumulation in vegetation from the Canadian Rocky Mountains. [source] ESTIMATING A GEOGRAPHICALLY EXPLICIT MODEL OF POPULATION DIVERGENCEEVOLUTION, Issue 3 2007L. Lacey Knowles Patterns of genetic variation can provide valuable insights for deciphering the relative roles of different evolutionary processes in species differentiation. However, population-genetic models for studying divergence in geographically structured species are generally lacking. Since these are the biogeographic settings where genetic drift is expected to predominate, not only are population-genetic tests of hypotheses in geographically structured species constrained, but generalizations about the evolutionary processes that promote species divergence may also be potentially biased. Here we estimate a population-divergence model in montane grasshoppers from the sky islands of the Rocky Mountains. Because this region was directly impacted by Pleistocene glaciation, both the displacement into glacial refugia and recolonization of montane habitats may contribute to differentiation. Building on the tradition of using information from the genealogical relationships of alleles to infer the geography of divergence, here the additional consideration of the process of gene-lineage sorting is used to obtain a quantitative estimate of population relationships and historical associations (i.e., a population tree) from the gene trees of five anonymous nuclear loci and one mitochondrial locus in the broadly distributed species Melanoplus oregonensis. Three different approaches are used to estimate a model of population divergence; this comparison allows us to evaluate specific methodological assumptions that influence the estimated history of divergence. A model of population divergence was identified that significantly fits the data better compared to the other approaches, based on per-site likelihood scores of the multiple loci, and that provides clues about how divergence proceeded in M. oregonensis during the dynamic Pleistocene. Unlike the approaches that either considered only the most recent coalescence (i.e., information from a single individual per population) or did not consider the pattern of coalescence in the gene genealogies, the population-divergence model that best fits the data was estimated by considering the pattern of gene lineage coalescence across multiple individuals, as well as loci. These results indicate that sampling of multiple individuals per population is critical to obtaining an accurate estimate of the history of divergence so that the signal of common ancestry can be separated from the confounding influence of gene flow,even though estimates suggest that gene flow is not a predominant factor structuring patterns of genetic variation across these sky island populations. They also suggest that the gene genealogies contain information about population relationships, despite the lack of complete sorting of gene lineages. What emerges from the analyses is a model of population divergence that incorporates both contemporary distributions and historical associations, and shows a latitudinal and regional structuring of populations reminiscent of population displacements into multiple glacial refugia. Because the population-divergence model itself is built upon the specific events shaping the history of M. oregonensis, it provides a framework for estimating additional population-genetic parameters relevant to understanding the processes governing differentiation in geographically structured species and avoids the problems of relying on overly simplified and inaccurate divergence models. The utility of these approaches, as well as the caveats and future improvements, for estimating population relationships and historical associations relevant to genetic analyses of geographically structured species are discussed. [source] HISTORIC CYCLES OF FRAGMENTATION AND EXPANSION IN PARNASSIUS SMINTHEUS (PAPILIONIDAE) INFERRED USING MITOCHONDRIAL DNAEVOLUTION, Issue 1 2004Eric G. DeChaine Abstract Climate oscillations of the Quaternary drove the repeated expansion and contraction of ecosystems. Alpine organisms were probably isolated in sky island refugia during warm interglacials, such as now, and expanded their range by migrating down-slope during glacial periods. We used population genetic and phylogenetic approaches to infer how paleoclimatic events influenced the distribution of genetic variation in the predominantly alpine butterfly Parnassius smintheus. We sequenced a 789 bp region of cytochrome oxidase I for 385 individuals from 20 locations throughout the Rocky Mountains, ranging from southern Colorado to northern Montana. Analyses revealed at lease two centers of diversity in the northern and southern Rocky Mountains and strong population structure. Nested clade analysis suggested that the species experienced repeated cycles of population expansion and fragmentation. The estimated ages of these events, assuming a molecular clock, corresponded with paleoclimatic data on habitat expansion and contraction over the past 400,000 years. We propose that alpine butterflies persisted in an archipelago of isolated sky islands during interglacials and that populations expanded and became more connected during cold glacial periods. An archipelago model implies that the effects of genetic drift and selection varied among populations, depending on their latitude, area, and local environment. Alpine organisms are sensitive indicators of climate change and their history can be used to predict how high-elevation ecosystems might respond to further climate warming. [source] Do male and female black-backed woodpeckers respond differently to gaps in habitat?EVOLUTIONARY APPLICATIONS (ELECTRONIC), Issue 3 2010Jennifer C. Pierson Abstract We used population- and individual-based genetic approaches to assess barriers to movement in black-backed woodpeckers (Picoides arcticus), a fire-specialist that mainly occupies the boreal forest in North America. We tested if male and female woodpeckers exhibited the same movement patterns using both spatially implicit and explicit genetic analyses to define population structure and movement patterns of both sexes among populations. Three genetic groups were identified, a large, genetically continuous population that spans from the Rocky Mountains to Quebec, a small isolated population in South Dakota and a separate population in the western portion of their distribution (Oregon). Patterns of genetic diversity suggest extensive gene flow mediated by both males and females within the continuous boreal forest. However, male-mediated gene flow is the main form of connectivity between the continuously distributed group and the smaller populations of South Dakota and Oregon that are separated by large areas of unforested habitat, which apparently serves as a barrier to movement of female woodpeckers. [source] Interregional comparisons of sediment microbial respiration in streamsFRESHWATER BIOLOGY, Issue 2 2000B. H. Hill Summary 1The rate of microbial respiration on fine-grained stream sediments was measured at 371 first to fourth-order streams in the Central Appalachian region (Maryland, Pennsylvania, Virginia, and West Virginia), Southern Rocky Mountains (Colorado), and California's Central Valley in 1994 and 1995. 2Study streams were randomly selected from the United States Environmental Protection Agency's (USEPA) River Reach File (RF3) using the sample design developed by USEPA's Environmental Monitoring and Assessment Program (EMAP). 3Respiration rate ranged from 0 to 0.621 g O2 g -1 AFDM h -1 in Central Appalachian streams, 0-0.254 g O2 g -1 AFDM h -1 in Rocky Mountain streams, and 0-0.436 g O2 g -1 AFDM h -1 in Central Valley streams. 4Respiration was significantly lower in Southern Rocky Mountain streams and in cold water streams (< 15 °C) of the Central Appalachians. 5Within a defined index period, respiration was not significantly different between years, and was significantly correlated with stream temperature and chemistry (DOC, total N, total P, K, Cl, and alkalinity). 6The uniformity of respiration estimates among the three study regions suggests that sediment microbial respiration may be collected at any number of scales above the site-level for reliable prediction of respiration patterns at larger spatial scales. [source] Temporal coherence of two alpine lake basins of the Colorado Front Range, U.S.A.FRESHWATER BIOLOGY, Issue 3 2000J. I. L. L. S. Baron 1. Knowledge of synchrony in trends is important to determining regional responses of lakes to disturbances such as atmospheric deposition and climate change. We explored the temporal coherence of physical and chemical characteristics of two series of mostly alpine lakes in nearby basins of the Colorado Rocky Mountains. Using year-to-year variation over a 10-year period, we asked whether lakes more similar in exposure to the atmosphere be-haved more similarly than those with greater influence of catchment or in-lake processes. 2. The Green Lakes Valley and Loch Vale Watershed are steeply incised basins with strong altitudinal gradients. There are glaciers at the heads of each catchment. The eight lakes studied are small, shallow and typically ice-covered for more than half the year. Snowmelt is the dominant hydrological event each year, flushing about 70% of the annual discharge from each lake between April and mid-July. The lakes do not thermally stratify during the period of open water. Data from these lakes included surface water temper-ature, sulphate, nitrate, calcium, silica, bicarbonate alkalinity and conductivity. 3. Coherence was estimated by Pearson's correlation coefficient between lake pairs for each of the different variables. Despite close geographical proximity, there was not a strong direct signal from climatic or atmospheric conditions across all lakes in the study. Individual lake characteristics overwhelmed regional responses. Temporal coherence was higher for lakes within each basin than between basins and was highest for nearest neighbours. 4. Among the Green Lakes, conductivity, alkalinity and temperature were temporally coherent, suggesting that these lakes were sensitive to climate fluctuations. Water tem-perature is indicative of air temperature, and conductivity and alkalinity concentrations are indicative of dilution from the amount of precipitation flushed through by snowmelt. 5. In Loch Vale, calcium, conductivity, nitrate, sulphate and alkalinity were temporally coherent, while silica and temperature were not. This suggests that external influences are attenuated by internal catchment and lake processes in Loch Vale lakes. Calcium and sulphate are primarily weathering products, but sulphate derives both from deposition and from mineral weathering. Different proportions of snowmelt versus groundwater in different years could influence summer lake concentrations. Nitrate is elevated in lake waters from atmospheric deposition, but the internal dynamics of nitrate and silica may be controlled by lake food webs. Temperature is attenuated by inconsistently different climates across altitude and glacial meltwaters. 6. It appears that, while the lakes in the two basins are topographically close, geologically and morphologically similar, and often connected by streams, only some attributes are temporally coherent. Catchment and in-lake processes influenced temporal patterns, especially for temperature, alkalinity and silica. Montane lakes with high altitudinal gradients may be particularly prone to local controls compared to systems where coherence is more obvious. [source] Ecological implications of xylem cavitation for several Pinaceae in the Pacific Northern USAFUNCTIONAL ECOLOGY, Issue 5 2000J. Piñol Abstract 1.,Xylem hydraulic properties and vulnerability to cavitation (determined using the air-injection method) were studied in six Pinaceae of the northern Rocky Mountains: Pinus ponderosa, Pseudotsuga menziesii, Larix occidentalis, Pinus contorta, Pinus albicaulis and Abies lasiocarpa. We tested whether species extending into drier habitats exhibited increased resistance to water stress-induced cavitation, and whether there is a trade-off between xylem transport capacity and resistance to cavitation. 2.,At lower elevations the more drought-tolerant P. ponderosa was much less resistant to cavitation than the codominant P. menziesii. Greater vulnerability to cavitation in P. ponderosa was compensated for, at least in part, by increased stomatal control of water loss (inferred from carbon isotope discrimination) and by increased sapwood to leaf area ratios. Similar differences, but less pronounced, were found in codominant species at higher elevations. 3.,Leaf specific hydraulic conductivity was negatively correlated with mean cavitation pressure. When species were separated into pines and non-pines, sapwood specific conductivity and mean cavitation pressure were also negatively correlated within each of the two groups. 4.,Our results indicate that within the evergreen conifers examined, greater resistance to water stress-induced cavitation is not required for survival in more xeric habitats, and that there is a trade-off between xylem conductance and resistance to cavitation. [source] Pilot study experiments sourcing quartzite, Gunnison Basin, ColoradoGEOARCHAEOLOGY: AN INTERNATIONAL JOURNAL, Issue 6 2008Bonnie L. Pitblado This paper reports the results of pilot-study efforts to develop methods to profile quartzite, a rock type to which geochemical and other sourcing techniques have only rarely been applied. The long-term goal of the research is to fingerprint sources of quartzite in the Gunnison Basin, southwest Colorado, used by Paleoindian people ca. 11,500,8,000 years ago to make stone tools. Success would facilitate reconstruction of Paleoindian mobility in the Southern Rocky Mountains and potentially anywhere prehistoric people used quartzite. The goals of this paper are more modest: to demonstrate that a small-scale exploration of sourcing techniques suggests reason for optimism that quartzites may be amenable to source discrimination. For the same twenty Gunnison Basin quartzite samples, this study evaluated petrography, ultraviolet fluorescence (UVF), wavelength dispersive X-ray fluorescence (WD-XRF), instrumental neutron activation analysis (INAA), and inductively coupled plasma mass spectrometry,both acid-digestion (AD-ICP-MS) and laser ablation (LA-ICP-MS),as means to differentiate among the specimens and the sources they represent. Although more testing is needed to verify and refine our results, the study suggests there is potential for petrography, INAA, and both versions of ICP-MS to discriminate among quartzites from different source localities in the Gunnison Basin. The greatest potential for discriminating among different sources of quartzite in the Gunnison Basin may lie in a methodology combining petrographic analysis and LA-ICPMS. Future testing is required to evaluate this two-fold approach. © 2008 Wiley Periodicals, Inc. [source] Fire-cracked rock features on sandy landforms in the Northern Rocky Mountains: Toward establishing reliable frames of reference for assessing site integrityGEOARCHAEOLOGY: AN INTERNATIONAL JOURNAL, Issue 5 2007Alston V. Thoms In cool coniferous forest settings of the Northern Rocky Mountains, well-preserved fire-cracked rock (FCR) features within 30 cm of the surface on ostensibly stable, sandy upland landforms date to the last six millennia. Isolated FCR and artifacts sometimes extend a meter or more below surface, which is suggestive of in situ burial. A paucity of intact features in the lower solum, however, is consistent with the downward migration of deeply buried artifacts by biomechanical processes, especially floralturbation. Moreover, an absence of credible sediment source areas usually precludes colluvial deposition, and results of grain-size analysis reported herein are inconsistent with eolian deposition. Site disarticulation rates tend to be faster on landforms in warmer forested regions of south-central North America, given that most intact FCR features there date only to the last two millennia. The very presence of millennia-old FCR features in these diverse settings, however, is a testament to their durability and utility as measures of site integrity. © 2007 Wiley Periodicals, Inc. [source] Paleoindian environmental change and landscape response in Barger Gulch, Middle Park, ColoradoGEOARCHAEOLOGY: AN INTERNATIONAL JOURNAL, Issue 6 2005James H. Mayer Middle Park, a high-altitude basin in the Southern Rocky Mountains of north-central Colorado, contains at least 59 known Paleoindian localities. At Barger Gulch Locality B, an extensive Folsom assemblage (,10,500 14C yr B.P.) occurs within a buried soil. Radiocarbon ages of charcoal and soil organic matter, as well as stratigraphic positions of artifacts, indicate the soil is a composite of a truncated, latest-Pleistocene soil and a younger mollic epipedon formed between ,6000 and 5200 14C yr B.P. and partially welded onto the older soil following erosion and truncation. Radiocarbon ages from an alluvial terrace adjacent to the excavation area indicate that erosion followed by aggradation occurred between ,10,200 and 9700 14C yr B.P., and that the erosion is likely related to truncation of the latest-Pleistocene soil. Erosion along the main axis of Barger Gulch occurring between ,10,000 and 9700 14C yr B.P. was followed by rapid aggradation between ,9700 and 9550 14C yr B.P., which, along with the erosion at Locality B, coincides with the abrupt onset of monsoonal precipitation following cooling in the region ,11,000,10,000 14C yr B.P. during the Younger Dryas oscillation. Buried soils dated between ,9500 and 8000 14C yr B.P. indicate relative landscape stability and soil formation throughout Middle Park. Morphological characteristics displayed by early Holocene soils suggest pedogenesis under parkland vegetation in areas currently characterized by sagebrush steppe. The expansion of forest cover into lower elevations during the early Holocene may have resulted in lower productivity in regards to mammalian fauna, and may partly explain the abundance of early Paleoindian sites (,11,000,10,000 14C yr B.P., 76%) relative to late Paleoindian sites (,10,000,8000 14C yr B.P., 24%) documented in Middle Park. © 2005 Wiley Periodicals, Inc. [source] ALPINE AREAS IN THE COLORADO FRONT RANGE AS MONITORS OF CLIMATE CHANGE AND ECOSYSTEM RESPONSE,GEOGRAPHICAL REVIEW, Issue 2 2002MARK W. WILLIAMS ABSTRACT. The presence of a seasonal snowpack in alpine environments can amplify climate signals. A conceptual model is developed for the response of alpine ecosystems in temperate, midlatitude areas to changes in energy, chemicals, and water, based on a case study from Green Lakes Valley,Niwot Ridge, a headwater catchment in the Colorado Front Range. A linear regression shows the increase in annual precipitation of about 300 millimeters from 1951 to 1996 to be significant. Most of the precipitation increase has occurred since 1967. The annual deposition of inorganic nitrogen in wetfall at the Niwot Ridge National Atmospheric Deposition Program site roughly doubled between 1985,1988 and 1989,1992. Storage and release of strong acid anions, such as those from the seasonal snowpack in an ionic pulse, have resulted in episodic acidification of surface waters. These biochemical changes alter the quantity and quality of organic matter in high-elevation catchments of the Rocky Mountains. Affecting the bottom of the food chain, the increase in nitrogen deposition may be partly responsible for the current decline of bighorn sheep in the Rocky Mountains. [source] Severe Deep Moist Convective Storms: Forecasting and MitigationGEOGRAPHY COMPASS (ELECTRONIC), Issue 1 2008David L. Arnold Small-scale (2,20 km) circulations, termed ,severe deep moist convective storms', account for a disproportionate share of the world's insured weather-related losses. Spatial frequency maximums of severe convective events occur in South Africa, India, Mexico, the Caucasus, and Great Plains/Prairies region of North America, where the maximum tornado frequency occurs east of the Rocky Mountains. Interest in forecasting severe deep moist convective systems, especially those that produce tornadoes, dates to 1884 when tornado alerts were first provided in the central United States. Modern thunderstorm and tornado forecasting relies on technology and theory, but in the post-World War II era interest in forecasting has also been driven by public pressure. The forecasting process begins with a diagnostic analysis, in which the forecaster considers the potential of the atmospheric environment to produce severe convective storms (which requires knowledge of the evolving kinematic and thermodynamic fields, and the character of the land surface over which the storms will pass), and the likely character of the storms that may develop. Improvements in forecasting will likely depend on technological advancements, such as the development of phased-array radar systems and finer resolution numerical weather prediction models. Once initiated, the evolution of deep convective storms is monitored by satellite and radar. Mitigation of the hazards posed by severe deep moist convective storms is a three-step process, involving preparedness, response, and recovery. Preparedness implies that risks have been identified and organizations and individuals are familiar with a response plan. Response necessitates that potential events are identified before they occur and the developing threat is communicated to the public. Recovery is a function of the awareness of local, regional, and even national governments to the character and magnitude of potential events in specific locations, and whether or not long-term operational plans are in place at the time of disasters. [source] Non-random patterns in the Yellowstone ecosystem: inferences from mammalian body size, order and biogeographical affinityGLOBAL ECOLOGY, Issue 2 2007Judsen E. Bruzgul ABSTRACT Aim, Our aim was to investigate how the environment, species characteristics and historical factors at the subcontinental scale affect patterns of diversity. We used the assembly of the Yellowstone biota over the past 10,000 years as a natural experiment for investigating the processes that generate a modern non-volant mammal species pool. Location, The data represent species from throughout North America with special attention to the non-volant mammals of Yellowstone National Park, USA. Methods, We used digitized range maps to determine biogeographical affinity for all non-volant mammals in the Rocky Mountains, Deserts and Great Plains biogeographical regions of North America. This biogeographical affinity, along with taxonomic order and body size class, was used to test whether non-random patterns exist in the assemblage of Yellowstone non-volant mammals. These characteristics were also used to investigate the strength of non-random processes, such as habitat or taxon filtering, on particular groups of species or individual species. Results, Our results indicated that the Yellowstone fauna is composed of a non-random subset of mammals from specific body size classes and with particular biogeographical affinities. Analyses by taxonomic order found significantly more Carnivora from the Rocky Mountains region and significantly fewer Rodentia from the Deserts region than expected from random assembly. Analyses using body size classes revealed deviations from expectations, including several significant differences between the frequency distribution of regional body sizes and the distribution of those species found within Yellowstone. Main conclusions, Our novel approach explores processes affecting species pool assembly in the Yellowstone region and elsewhere, and particularly identifies unique properties of species that may contribute to non-random assembly. Focusing on the mechanisms generating diversity, not just current diversity patterns, will assist the design of conservation strategies given future environmental change scenarios. [source] Modified passive capillary samplers for collecting samples of snowmelt infiltration for stable isotope analysis in remote, seasonally inaccessible watersheds 2: field evaluationHYDROLOGICAL PROCESSES, Issue 7 2010Marty D. Frisbee Abstract Twelve modified passive capillary samplers (M-PCAPS) were installed in remote locations within a large, alpine watershed located in the southern Rocky Mountains of Colorado to collect samples of infiltration during the snowmelt and summer rainfall seasons. These samples were collected in order to provide better constraints on the isotopic composition of soil-water endmembers in the watershed. The seasonally integrated stable isotope composition (,18O and ,2H) of soil-meltwater collected with M-PCAPS installed at shallow soil depths < 10 cm was similar to the seasonally integrated isotopic composition of bulk snow taken at the soil surface. However, meltwater which infiltrated to depths > 20 cm evolved along an isotopic enrichment line similar to the trendline described by the evolution of fresh snow to surface runoff from snowmelt in the watershed. Coincident changes in geochemistry were also observed at depth suggesting that the isotopic and geochemical composition of deep infiltration may be very different from that obtained by surface and/or shallow-subsurface measurements. The M-PCAPS design was also used to estimate downward fluxes of meltwater during the snowmelt season. Shallow and deep infiltration averaged 8·4 and 4·7 cm of event water or 54 and 33% of the measured snow water equivalent (SWE), respectively. Finally, dominant shallow-subsurface runoff processes occurring during snowmelt could be identified using geochemical data obtained with the M-PCAPS design. One soil regime was dominated by a combination of slow matrix flow in the shallow soil profile and fast preferential flow at depth through a layer of platy, volcanic rocks. The other soil regime lacked the rock layer and was dominated by slow matrix flow. Based on these results, the M-PCAPS design appears to be a useful, robust methodology to quantify soil-water fluxes during the snowmelt season and to sample the stable isotopic and geochemical composition of soil-meltwater endmembers in remote watersheds. Copyright © 2009 John Wiley & Sons, Ltd. [source] Spatial grouping of United States climate stations using a hybrid clustering approachINTERNATIONAL JOURNAL OF CLIMATOLOGY, Issue 7 2001Arthur T. DeGaetano Abstract The development of a hybrid clustering technique based on the geographic proximity of observing stations and some application-driven measure of statistical similarity (in this case rank correlation) is described. The procedure is then applied to temperature and precipitation data from the United States (US) Historical Climatology Network. The resulting station groups provide some insight into the number of observation stations that are necessary to monitor adequately the climate of the US. Based on temperature data alone, a 287-station subset of the original 1145 sites would be adequate to account for 80% of the spatial variability in seasonal temperature across the US. Geographically the distribution of these stations would be relatively sparse in the centre of the country with higher station density along the East Coast and from the Rocky Mountains to the West Coast. Generally, the temperature clusters match the existing US climate divisions to some extent. To monitor adequately the spatial variability of precipitation, a network of similar size could be used. However, such a network would only account for 65% of the spatial variability in precipitation. In this case, fairly uniform station density is indicated across the country with the highest station density in Florida and the Dakotas. A similar number of stations, but with slightly different geographic groupings would be adequate to monitor precipitation and temperature simultaneously. Copyright © 2001 Royal Meteorological Society [source] Breeding in high-elevation habitat results in shift to slower life-history strategy within a single speciesJOURNAL OF ANIMAL ECOLOGY, Issue 2 2009H. Bears Summary 1Elevational gradients create environmental variation that is hypothesized to promote variation in life-history strategies. We tested whether differences in life-history strategies were associated with elevation in a songbird, the dark-eyed junco (Junco hyemalis; Aves; A.O.U. 1998). 2We monitored birds in four replicated sites per elevation, at 2000 m a.s.l. (high elevation) and 1000 m a.s.l. (low elevation), in the Rocky Mountains of Canada. 3Over 5 years, we measured the following traits and vital rates: egg-laying schedules, morphological indicators of reproductive stage, seasonal reproductive success, indicators of competitive class (age, size, arrival time), and survival rates. 4We found two main patterns: with an increase in breeding elevation, dark-eyed juncos delayed the development of structures necessary for reproduction (e.g. cloacal protuberance in males) and reduced the duration of their reproductive period to less than half of the time used by low-elevation birds; and 5Juncos at high-elevation sites had 55,61% lower annual reproductive success and 15 to 20% higher survival rates. While adult juncos at high elevations produced fewer offspring, those offspring were in better condition. Proportions of age and size classes in high- compared to low-elevation populations were similar, suggesting that a life-history trade-off is present, rather than competition forcing inferior competitors to breed in a peripheral habitat. The apparent trade-off between reproduction and survival corresponded to a shorter period of favourable weather and available food in high- compared to low-elevation habitats. 6Thus, elevation had a strong influence on life-history characteristics of a single species over a short spatial distance, suggesting a shift in life history from a high reproductive strategy at lower elevations to a high survivor strategy at high elevations. 7This is the first paper to show a shift in avian life-history strategies along an elevational gradient (in both genders, of multiple age classes) when region (latitude) and phylogenetic histories are controlled for. [source] Disturbance facilitates rapid range expansion of aspen into higher elevations of the Rocky Mountains under a warming climateJOURNAL OF BIOGEOGRAPHY, Issue 1 2010Simon M. Landhäusser Abstract Aim, Trembling aspen (Populus tremuloides Michx.) is absent in the upper foothills region of west-central Alberta because of the cold conditions and short growing season at this high elevation. However, in recent years it appears that aspen has been establishing from seed in this zone and that it has been doing so mainly as a result of forest harvesting. The objectives of this study were to determine the frequency of and types of microsite required for the successful establishment of aspen seedlings at these higher elevations. Location, Rocky Mountains Upper Foothills Natural Subregion of west-central Alberta, Canada. Methods, The current distribution of mature aspen and the presence and absence of aspen seedlings in harvested areas were determined in an area c. 300 km2 in size, using ground and aerial surveys. In an intensive study, 12 belt transects (180 m long and 5 m wide) were established in areas disturbed by forest harvesting at high elevations where no aspen was present prior to harvesting. Transects were surveyed seven growing seasons after disturbance and the microsites occupied by aspen seedlings were characterized according to their substrate and microtopography. Similarly, the availability of different substrates and microtopographic positions were assessed by systematic point sampling on these sites. Results, On level surfaces, aspen seedling regeneration was found up to 200 m higher in elevation than the mature aspen in the original undisturbed forests. Overall, there were 428 seedlings ha,1 established on these transects, and the age distribution indicates that aspen seedlings had established in each of the seven growing seasons since the disturbance. Nearly all of the seedlings (93%) were established on mineral soil microsites and virtually no seedlings were established on undisturbed forest floor layers. Significantly more seedlings were found in concave microtopographic positions. Main conclusions, This study indicates that aspen establishment from seed is currently not a stochastic event and demonstrates that aspen is rapidly expanding its range upslope in the Canadian Rocky Mountain region as a result of forest management practices that expose mineral soil substrates in conjunction with a warming climate. The change in canopy composition from conifer to deciduous forests at these higher elevations will have far-reaching implications for ecosystem processes and functions. [source] Late-glacial and Holocene palaeovegetation zonal reconstruction for central and north-central North AmericaJOURNAL OF BIOGEOGRAPHY, Issue 6 2005W. L Strong Abstract Aim, The purpose of this study is to develop palaeovegetation zonation models for central and north-central North America, based on late-Quaternary and Holocene pollen stratigraphic data (n = 246 sites). A secondary purpose was to evaluate an hypothesis (Strong & Hills, 2003) to explain the disjunct distribution of species in western Alberta. Location, Hudson Bay-Lake Michigan to the Rocky Mountains region, north of 36° N to the Arctic Ocean (c. 70° N). Methods, Pollen profiles spanning 40 years of palaeoecological research in North America were extracted from published and unpublished archival sources. Individual profiles were subdivided into 1000-year increments based on the assumption of a constant sedimentation rate between stratigraphic dates (e.g. surface sediments, radiocarbon 14C dates, tephra layers). The pollen composition among profiles was standardized to 54 commonly recognized taxa, with percentage composition within each stratigraphic sample prorated to 100% prior to analysis. Near-surface sediments from these profiles were included as analogues of modern vegetation. Cluster analysis was used as a guide to the classification of 2356 temporal stratigraphic samples, which resulted in the recognition of 16 pollen groups. These groups were summarized in terms of their pollen composition, mapped, and used in combination with terrain information and an ecological knowledge of the study area to construct six physiognomically-based palaeovegetation zonation models at 2000-year intervals from 14,000 to 4000 yr bp (radiocarbon years before present). Results, The 14,000 yr bp model placed Boreal and Cordilleran Forests proximal to the southern glacial front, whereas Arctic tundra dominated the Yukon Territory,Alaska ice-free zone. Pollen and macrofossil evidence suggests that this Boreal Forest zone contained a mixture of coniferous and deciduous tree species. Grassland was postulated immediately south of the forest zone, with its northern extreme near 49° N latitude in the Alberta,Montana border area. Separation of the Laurentide and Cordilleran glacial fronts about 12,000 yr bp initiated the northward advance of Boreal Forests into western Canada. By the end of the Hypsithermal at about 6000 yr bp, Boreal Forests occurred near the Arctic Ocean, and Grassland and Aspen Parkland zones may have extended to 54° N and 59° N latitude in Alberta, respectively. Between 6000 and 4000 yr bp, a 5° and 1° latitudinal southward shift of the northern Boreal Forest and Grassland/Aspen Parkland boundaries occurred, respectively, near their contemporary positions with corresponding expansions of the Subarctic and Arctic zones. Modern Canadian Cordilleran Forests along the eastern slopes of the Rocky Mountains were interpreted as originating from the north-central Montana,south-western Alberta area. Jack pine (Pinus banksiana Lamb.), a common Boreal Forest species, appears to have entered central Canada via the north side of Lake Superior after 11,000 yr bp. Main conclusions, Modern vegetation in central Canada evolved from biomes located in the northern USA during the late-Quaternary. The Boreal Forest biome contained the same arboreal taxa as the modern vegetation, except it lacked jack pine. The proposed regional palaeovegetation models support the hypothesis of Strong & Hills (2003), but new independent palaeoecological data will be needed for a proper evaluation. [source] Post-Hypsithermal plant disjunctions in western Alberta, CanadaJOURNAL OF BIOGEOGRAPHY, Issue 3 2003W. L Strong Abstract Aim, Evaluate the hypothesis that nine disjunct vascular plant species along the eastern slopes of the Rocky Mountains and in the Peace River District of west-central Alberta represent remnants of more southerly vegetation that occupied these areas during the Holocene Hypsithermal (9000,6000 yr bp). Alternatively, these plants represent populations that became established because of independent chance dispersal events. Location, This study focuses on the area east of the Rocky Mountain Continental Divide in the Province of Alberta and the State of Montana in western Canada and USA, respectively. Methods, Disjunct species were identified and their distributions mapped based on a review of occurrence maps and records, botanical floras and checklists, herbaria specimens, ecological and botanical studies, and field surveys of selected species. A disjunct species was defined as a plant population separated from its next nearest occurrence by a distance of > 300 km. Evaluation of the hypothesis was based on a review of published and unpublished pollen stratigraphy and palaeoecological studies. The potential geographical distribution of Hypsithermal vegetation was based on modern regional-based ecosystem mapping and associated monthly temperature summaries as well as future climatic warming models. Results, The hypothesis was compatible with Holocene pollen stratigraphy, Hypsithermal permafrost and fen occurrence, and palaeosol phytolith analyses; and future global climatic warming models. Modelled regional Hypsithermal vegetation based on a 1 °C increase in July temperatures relative to current conditions, indicated that much of the boreal forest zone in Alberta could have been grassland, which would explain the occurrence of Prairie species in the Peace River District. This amount of latitudinal vegetation shift (6.5°) was similar to an earlier Hypsithermal permafrost zone location study. An equivalent shift in vegetation along the eastern Cordillera would have placed south-western Montana-like vegetation and species such as Boykinia heucheriformis (Rydb.) Rosend. and Saxifraga odontoloma Piper within the northern half of the Rocky Mountains and foothills in Alberta, which represents the location of modern-day disjunct populations of these species. Main conclusions, Warmer and drier climatic conditions during the Holocene Hypsithermal resulted in the northward displacement of vegetation zones relative to their current distribution patterns. Most of Alberta was probably dominated by grasslands during this period, except the Rocky Mountains and northern highlands. Modern-day species disjunctions within the Rocky Mountains and Peace River District as well as more northerly areas such as the Yukon Territory occurred when the vegetation receded southward in response to climatic cooling after the Hypsithermal. Wind dispersal was considered an unlikely possibility to explain the occurrence of the disjunct species, as most of the plants lack morphological adaptations for long distance transport and the prevailing winds were from west to east rather than south to north. However, consumption and transport of seeds by northward migrating birds could not be excluded as a possibility. [source] DEVONIAN CARBONATES OF THE NIGEL PEAK AREA, ROCKY MOUNTAINS, CANADA: A FOSSIL PETROLEUM SYSTEM?JOURNAL OF PETROLEUM GEOLOGY, Issue 3 2008J. Köster In this study we report on Devonian (Frasnian , Famennian) limestones and dolostones exposed near Nigel Peak in the Main Ranges of the Canadian Rocky Mountains. These carbonates are a proximal facies of the Southesk-Cairn Carbonate Complex. The investigated strata are stratigraphically equivalent to the oil- and gas bearing Nisku Formation in the subsurface of the Western Canada Sedimentary Basin, about 300 km to the east. The rocks were investigated by polarisation and cathodoluminescence microscopy, total organic carbon analysis, Rock-Eval pyrolysis, solid bitumen reflectance measurements, gas chromatography and fluid inclusion analysis. Thin section analyses showed that silt-grade quartz and saddle dolomite increase upward from the base of the stratigraphic section, and that porosities are generally low. This is due to reduction of pore space due to early cementation and extensive dolomitization. Cathodoluminescence identified up to four generations of calcite cements. TOC values ranged from 0.2 to 2.4 %. Rock-Eval pyrolysis of carbonate samples resulted in measurable S1 peaks but not S2 peaks, indicating that there was no residual petroleum generation potential. Organic petrographic analyses identified dispersed kerogen and migrabitumen, and calculated vitrinite reflectance values were around 4 % on average which implies peak temperatures of 234,262 °C (due to deep burial) or 309,352 °C (due to short term hydrothermal heating). Fluid inclusion data indicates at least one pulse of hot fluids with elevated homogenization temperatures of > 300 °C, and this may explain the high thermal maturity of the studied rocks. [source] Aspen succession and nitrogen loading: a case for epiphytic lichens as bioindicators in the Rocky Mountains, USAJOURNAL OF VEGETATION SCIENCE, Issue 3 2009Paul C. Rogers Abstract Question: Can lichen communities be used to assess short- and long-term factors affecting seral quaking aspen (Populus tremuloides) communities at the landscape scale? Location: Bear River Range, within the Rocky Mountains, in northern Utah and southern Idaho, USA. Method: Forty-seven randomly selected mid-elevation aspen stands were sampled for lichens and stand conditions. Plots were characterized according to tree species cover, basal area, stand age, bole scarring, tree damage, and presence of lichen species. We also recorded ammonia emissions with passive sensors at 25 urban and agricultural sites throughout an adjacent populated valley upwind of the forest stands. Nonmetric multidimensional scaling (NMS) ordination was used to evaluate an array of 20 variables suspected to influence lichen communities. Results: In NMS, forest succession explained most variance in lichen composition and abundance, although atmospheric nitrogen from local agricultural and urban sources also significantly influenced the lichen communities. Abundance of nitrophilous lichen species decreased with distance from peak ammonia sources and the urban center in all aspen succession classes. One lichen, Phaeophyscia nigricans, was found to be an effective bioindicator of nitrogen loading. Conclusions: Lichen communities in this landscape assessment of aspen forests showed clear responses to long-term (stand succession) and short-term (nitrogen deposition) influences. At the same time, several environmental factors (e.g. tree damage and scarring, distance to valley, topography, and stand age) had little influence on these same lichen communities. We recommend further use of epiphytic lichens as bioindicators of dynamic forest conditions. [source] Biology and establishment of mountain shrubs on mining disturbances in the Rocky Mountains, USALAND DEGRADATION AND DEVELOPMENT, Issue 5 2003M. W. Paschke Abstract The Rocky Mountains of the western United States contain many economically important natural resources. Increasing development of these resources has lead to land degradation, which often requires restoration efforts. A common type of disturbance in this region is mineral extraction and these activities often occur in zones of vegetation dominated by shrubs. These mined lands have proven to be particularly challenging to restore to native shrub cover. Mountain shrubland species such as big sagebrush (Artemisia tridentata), antelope bitterbrush (Purshia tridentata), Saskatoon serviceberry (Amelanchier alnifolia), black chokecherry (Prunus virginiana), mountain mahogany (Cercocarpus montanus), and snowberry (Symphoricarpos sp.) are crucial species for wildlife in this region due to the seasonal dependence of many wildlife species on the mountain shrubland zones. The development of successful restoration techniques for these key shrub species is therefore important for the continued and sustainable utilization of natural resources in the Rocky Mountains. The purpose of this literature review is to provide a reference to land managers working on woody plant establishment on mining disturbances in mountain shrublands in the Rocky Mountains. This review covers the biology, ecology, and propagation of six shrub species that are of primary importance for meeting regulatory compliance in this region. Based upon this survey of the published literature, we conclude that successful establishment of these species has most often involved: (1) utilization of local shrub ecotypes, varieties or subspecies in reclamation efforts, (2) protection from browsing during the establishment phase, (3) strategies for avoiding herbaceous competition, and (4) providing a source of mutualistic soil organisms. Additional specific recommendations for each of these species are discussed. Copyright © 2003 John Wiley & Sons, Ltd. [source] Shell structure, ontogeny and affinities of the Lower Cambrian bivalved problematic fossil Mickwitzia muralensis Walcott, 1913LETHAIA, Issue 4 2004UWE BALTHASAR Exceptionally preserved carbonate- and shale-hosted Mickwitzia muralensis from the Lower Cambrian Mural Formation, southern Canadian Rocky Mountains, complement one another to yield an unusually complete account of its ontogeny, ecology and phylogenetic relationships. The shell of M. muralensis is composed of dense phosphatic layers interspersed with porous organic-rich layers. At the insertion of shell-penetrating tubes, shell layers deflect inwards to produce inwardly pointing cones. The tubes are interpreted as having hosted setae that were secreted by outer-epithelial follicles. Follicular setae also occurred at the mantle margin, where they were oriented within the plane of the shell as in modern brachiopods. During ontogeny, the initial setae oriented in the plane of the shell occurred before the first shell-penetrative setae. In the juvenile and early-mature stages of shell secretion, a posterior opening was present between both valves and was used for the protrusion of an attachment structure. In the late-mature shell, this opening became fixed in the ventral valve. Based on the posterior margin and the shell microstructure, a close relationship between Mickwitzia and the paterinids is proposed with differences interpreted as heterochronic. The shell-penetrative setal apparatus of M. muralensis is distinct from that previously described of Micrina, though both types are conceivably homologous to adult and juvenile setae of modern brachiopods. [source] Phylogeography of Douglas-fir based on mitochondrial and chloroplast DNA sequences: testing hypotheses from the fossil recordMOLECULAR ECOLOGY, Issue 9 2010PAUL F. GUGGER Abstract The integration of fossil and molecular data can provide a synthetic understanding of the ecological and evolutionary history of an organism. We analysed range-wide maternally inherited mitochondrial DNA and paternally inherited chloroplast DNA sequence data with coalescent simulations and traditional population genetic methods to test hypotheses of population divergence generated from the fossil record of Douglas-fir (Pseudotsuga menziesii), an ecologically and economically important western North American conifer. Specifically, we tested (i) the hypothesis that the Pliocene orogeny of the Cascades and Sierra Nevada caused the divergence of coastal and Rocky Mountain Douglas-fir varieties; and (ii) the hypothesis that multiple glacial refugia existed on the coast and in the Rocky Mountains. We found that Douglas-fir varieties diverged about 2.11 Ma (4.37 Ma,755 ka), which could be consistent with a Pliocene divergence. Rocky Mountain Douglas-fir probably resided in three or more glacial refugia. More variable molecular markers would be required to detect the two coastal refugia suggested in the fossil record. Comparison of mitochondrial DNA and chloroplast DNA variation revealed that gene flow via pollen linked populations isolated from seed exchange. Postglacial colonization of Canada from coastal and Rocky Mountain refugia near the ice margin at the Last Glacial Maximum produced a wide hybrid zone among varieties that formed almost exclusively by pollen exchange and chloroplast DNA introgression, not seed exchange. Postglacial migration rates were 50,165 m/year, insufficient to track projected 21st century warming in some regions. Although fossil and genetic data largely agree, each provides unique insights. [source] | ||||||||||||||||