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Ryegrass
Kinds of Ryegrass Selected AbstractsIsolates of Microdochium nivale and M. majus Differentiated by Pathogenicity on Perennial Ryegrass (Lolium perenne L.) and in vitro Growth at Low TemperatureJOURNAL OF PHYTOPATHOLOGY, Issue 5 2006I. S. Hofgaard Abstract Pink snow mould is a serious disease on grasses and winter cereals in cold and temperate zones during winter. To better understand the basis for the variation in pathogenicity between different isolates of Microdochium nivale and M. majus and to simplify selection of highly pathogenic isolates to use when screening for resistance to pink snow mould in perennial ryegrass, we sought traits correlated with pathogenicity. Isolates of M. nivale were more pathogenic on perennial ryegrass than isolates of M. majus, as measured by survival and regrowth of perennial ryegrass after infection and incubation under simulated snow cover. Pathogenicity as measured by relative regrowth was highly correlated with fungal growth rate on potato dextrose agar (PDA) at 2°C. Measuring fungal growth on PDA therefore seems to be a relatively simple method of screening for potentially highly pathogenic isolates. In a study of a limited number of isolates, highly pathogenic isolates showed an earlier increase and a higher total specific activity of , -glucosidase, a cell wall-degrading enzyme, compared with less pathogenic isolates. None of the M. majus isolates was highly pathogenic on perennial ryegrass. Our results indicate biological differences between M. nivale and M. majus and thus strengthen the recently published sequence-based evidence for the elevation of these former varieties to species status. [source] Liveweight gains of lambs from Caucasian clover/ryegrass and white clover/ryegrass swards on soils of high and low fertilityGRASS & FORAGE SCIENCE, Issue 2 2007A. D. Black Summary The high nutritive value and persistence under a wide range of climatic and soil fertility conditions make Caucasian clover a potentially useful forage legume but there is little information about the performance of livestock grazing Caucasian clover/grass swards. This study compared liveweight gains of lambs grazing Caucasian clover/perennial ryegrass and white clover/perennial ryegrass swards on high fertility (Olsen P 20 mg L,1, SO4 -S 12 mg kg,1) and low fertility (Olsen P 11 mg L,1, SO4 -S 7 mg kg,1) soils from 1998 to 2001 in the South Island of New Zealand. Mean annual liveweight gains were 1178 kg ha,1 for Caucasian clover/perennial ryegrass and 1069 kg ha,1 for white clover/perennial ryegrass swards at high fertility compared with 1094 kg ha,1 and 1015 kg ha,1, respectively, at low fertility. There was a higher mean proportion of clover in Caucasian clover/perennial ryegrass (0·19) than white clover/perennial ryegrass (0·11) swards, but there were no differences in total herbage production between the two clover/perennial ryegrass swards. The mean concentration of crude protein in the herbage of Caucasian clover (302 g kg DM,1) was higher than that in white clover (287 g kg DM,1) and grass herbage (227 g kg DM,1). Estimated mean metabolizable energy concentrations in the herbage were 12·5 MJ kg DM,1 for the two clovers and 11·6 MJ kg DM,1 for grass herbage. The difference in liveweight gain between swards on soils of high and low fertility was associated with an increase in total herbage production of similar composition and nutritive value, giving a greater number of grazing days for the swards on soils of high than low fertility. [source] Top-down and bottom-up regulation of herbivores: Spodoptera frugiperda turns tables on endophyte-mediated plant defence and virulence of an entomopathogenic nematodeECOLOGICAL ENTOMOLOGY, Issue 3 2004Douglas S. Richmond Abstract., 1. The fungus Neotyphodium lolii forms a symbiotic relationship with its grass host Lolium perenne (perennial ryegrass). The fungus benefits from access to plant nutrients and photosynthate, whereas the plant benefits from acquired chemical defence against herbivory. 2. This study examined the potential for endophyte-mediated plant defences to influence interactions between fall armyworm Spodoptera frugiperda, and the entomopathogenic nematode Steinernema carpocapsae and clarified biological mechanisms underlying the observations made. 3. In laboratory and greenhouse experiments, S. frugiperda larvae were fed endophytic or non-endophytic L. perenne then exposed to S. carpocapsae or injected with the nematodes' symbiotic bacteria Xenorhabdus nematophila. 4. In all instances, S. frugiperda larvae fed endophyte-infected grass suffered significantly lower mortality than those fed non-endophytic plants. Although larvae fed endophyte-infected grass often had significantly lower biomass than those fed uninfected grass, these differences did not account for altered susceptibility to S. carpocapsae. 5. Endophyte-mediated reductions in herbivore susceptibility to the nematode pathogen represent a herbivore adaptation that effectively turns the tables on both plant and natural enemy by reducing the virulence of the nematodes' symbiotic bacteria while expanding the temporal window of herbivory. [source] Fatty acid ruminal metabolism and digestibility in cows fed perennial ryegrassEUROPEAN JOURNAL OF LIPID SCIENCE AND TECHNOLOGY, Issue 8 2007Michel Doreau Abstract The ruminal metabolism and intestinal digestibility of fatty acids (FA) was studied in four mid-lactation dairy cows fitted with ruminal and duodenal cannulae. The cows received fresh perennial ryegrass as sole feed. Two grasses were compared, differing in the soluble carbohydrate/nitrogen ratio (SC+/N, and SC,/N+ ratios). This ratio was modified by the combination of three factors: age of regrowth, N,fertilisation and hour of cutting. The two grasses contained 12.5 and 20.7% of crude protein, and 24.6 and 13.7% of SC, respectively. The SC+/N, grass contained less FA [1.82 vs. 2.49% of dry matter (DM)] and a lower percentage of linolenic acid (60.9 vs. 65.6% of FA). As DM intake did not differ between treatments, FA intake was higher for the SC,/N+ treatment. The duodenal composition of FA revealed numerous isomers of 18:1 and conjugated and non-conjugated isomers of 18:2. SC,/N+ treatment resulted in a higher duodenal flow of stearic, linoleic, and linolenic acids and of several intermediates of linolenic acid hydrogenation. This was a consequence of differences in intake. Intestinal digestibility of FA from both grasses was high (around 90%). The proportions of FA in plasma did not reflect with accuracy the differences in duodenal FA resulting from differences in ruminal metabolism. [source] The influence of below-ground herbivory and defoliation of a legume on nitrogen transfer to neighbouring plantsFUNCTIONAL ECOLOGY, Issue 2 2007E. AYRES Summary 1Both foliar and root herbivory can alter the exudation of carbon from plant roots, which in turn can affect nitrogen availability in the soil. However, few studies have investigated the effects of herbivory on N fluxes from roots, which can directly increase N availability in the soil and uptake by neighbouring plants. Moreover, the combined effects of foliar and root herbivory on N fluxes remains unexplored. 2We subjected the legume white clover (Trifolium repens L.) to defoliation (through clipping) and root herbivory (by an obligate root-feeding nematode, Heterodera trifolii Goggart) to examine how these stresses individually, and simultaneously, affected the transfer of T. repens -derived N to neighbouring perennial ryegrass (Lolium perenne L.) plants using 15N stable-isotope techniques. We also examined the effects of defoliation and root herbivory on the size of the soil microbial community and the growth response of L. perenne. 3Neither defoliation nor root herbivory negatively affected T. repens biomass. On the contrary, defoliation increased root biomass (34%) and total shoot production by T. repens (100%). Furthermore, defoliation resulted in a fivefold increase in T. repens -derived 15N recovered in L. perenne roots, and increased the size of the soil microbial biomass (77%). In contrast, root herbivory by H. trifolii slightly reduced 15N transfer from T. repens to L. perenne when T. repens root 15N concentration was included as a covariate, and root herbivory did not affect microbial biomass. Growth of L. perenne was not affected by any of the treatments. 4Our findings demonstrate that defoliation of a common grassland legume can substantially increase the transfer of its N to neighbouring plants by directly affecting below-ground N fluxes. These finding require further examination under field conditions but, given the prevalence of N-limitation of plant productivity in terrestrial ecosystems, increased transfer of N from legumes to non-N-fixing species could alter competitive interactions, with implications for plant community structure. [source] Ten years of free-air CO2 enrichment altered the mobilization of N from soil in Lolium perenne L. swardsGLOBAL CHANGE BIOLOGY, Issue 8 2004Manuel K. Schneider Abstract Effects of free-air carbon dioxide enrichment (FACE, 60 Pa pCO2) on plant growth as compared with ambient pCO2 (36 Pa) were studied in swards of Lolium perenne L. (perennial ryegrass) at two levels of N fertilization (14 and 56 g m,2 a,1) from 1993 to 2002. The objectives were to determine how plant growth responded to the availability of C and N in the long term and how the supply of N to the plant from the two sources of N in the soil, soil organic matter (SOM) and mineral fertilizer, varied over time. In three field experiments, 15N-labelled fertilizer was used to distinguish the sources of available N. In 1993, harvestable biomass under elevated pCO2 was 7% higher than under ambient pCO2. This relative pCO2 response increased to 32% in 2002 at high N, but remained low at low N. Between 1993 and 2002, the proportions and amounts of N in harvestable biomass derived from SOM (excluding remobilized fertilizer) were, at high N, increasingly higher at elevated pCO2 than at ambient pCO2. Two factorial experiments confirmed that at high N, but not at low N, a higher proportion of N in harvestable biomass was derived from soil (including remobilized fertilizer) following 7 and 9 years of elevated pCO2, when compared with ambient pCO2. It is suggested that N availability in the soil initially limited the pCO2 response of harvestable biomass. At high N, the limitation of plant growth decreased over time as a result of the stimulated mobilization of N from soil, especially from SOM. Consequently, harvestable biomass increasingly responded to elevated pCO2. The underlying mechanisms which contributed to the increased mobilization of N from SOM under elevated pCO2 are discussed. This study demonstrated that there are feedback mechanisms in the soil which are only revealed during long-term field experiments. Such investigations are thus, a prerequisite for understanding the responses of ecosystems to elevated pCO2 and N supply. [source] Morphological development and nutritive value of herbage in five temperate grass species during primary growth: analysis of time dynamicsGRASS & FORAGE SCIENCE, Issue 2 2009Abstract In a 2-year field experiment, morphological development and measures of the nutritive value of herbage for livestock during primary growth in Meadow foxtail, Tall oatgrass, Cocksfoot, Perennial ryegrass and Yorkshire fog were investigated. All measured variables were affected significantly by both species and sampling date, and their interaction (P < 0·001), in the period of primary growth. Changes with time in mean stage weight for Meadow foxtail and Cocksfoot were different from the other species due to their indeterminate growth habits. Mean stage weight of Tall oatgrass and Yorkshire fog increased more rapidly than that of Perennial ryegrass with time. Changes in mean stage weight with time were described by linear, parabolic and sigmoid relationships. Crude protein (CP) concentration of herbage was higher for Cocksfoot and Meadow foxtail than for Perennial ryegrass. A parabolic relationship of CP concentration with time was typical for all the species. Concentrations of neutral-detergent fibre (NDF) and acid-detergent fibre (ADF) in herbage of the species differed most during the mid-period of primary growth. Their increases with time showed curvilinear (sigmoid and parabolic) relationships. Perennial ryegrass had lower concentrations of both NDF and ADF in herbage than the other species. Differences between the in vitro dry matter (DM) digestibility among the grasses increased in mid- and late periods of primary growth. Perennial ryegrass had higher values for in vitro DM digestibility but the difference from other species was small in the early period of primary growth and from cocksfoot in the late period of primary growth. In vitro DM digestibility showed, in most cases, a sigmoid and, in others, a linear decrease with time. Principal component analysis showed that perennial ryegrass and meadow foxtail were the most distinctive of the species in characteristics relating to morphological development and the nutritive value of herbage to livestock. [source] Effects of different rates and timing of application of nitrogen as slurry and mineral fertilizer on yield of herbage and nitrate-leaching potential of a maize/Italian ryegrass cropping system in north-west PortugalGRASS & FORAGE SCIENCE, Issue 1 2009H. Trindade Abstract Efficient use of cattle-slurry to avoid nitrogen (N) leaching and other losses is important in designing intensive dairy systems to minimize pollution of air and water. The response in dry-matter (DM) yield of herbage and nitrate-leaching potential to different rates and timing of application of N as cattle slurry and/or mineral fertilizer in a double-cropping system producing maize (Zea mays L.) silage and Italian ryegrass (Lolium multiflorum Lam.) was investigated in north-west Portugal. Nine treatments with different rates and combinations of cattle slurry, and with or without mineral-N fertilizer, applied at sowing and as a top-dressing to both crops, were tested and measurements were made of DM yield of herbage, N concentration of herbage, uptake of N by herbage and amounts of residual soil nitrate-N to a depth of 1 m, in a 3-year experiment. Regression analysis showed that the application of 150 and 100 kg of available N ha,1 to maize and Italian ryegrass, respectively, resulted in 0·95 of maximum DM yields of herbage and 0·90 of maximum N uptake by herbage. Residual amounts of nitrate-N in soil after maize ranged from 48 to 278 kg N ha,1 with an exponential increase in response to the amount of N applied; there were higher values of nitrate-leaching potential when mineral-N fertilizer was applied. The results suggest that it is possible in highly productive maize/Italian ryegrass systems to obtain high DM yields of herbage for maize silage and Italian ryegrass herbage with minimal leaching losses by using slurry exclusively at annual rates of up to 250 kg available N ha,1 (equivalent to 480 kg total N ha,1) in three applications. [source] Dry matter production, nutritive value and efficiency of nutrient utilization of a complementary forage rotation compared to a grass pasture systemGRASS & FORAGE SCIENCE, Issue 3 2008S. C. Garcia Abstract In pasture-based dairy farming, new sustainable systems that involve the annual dry matter (DM) production of grazed and conserved forage beyond the potential of grazed pasture alone are being sought. The objective of this experiment conducted in Australia was to compare a complementary forage rotation (CFR) for conservation and grazing, comprising an annual sequence of three crops, namely maize (Zea mays L), forage rape (Brassica napus L) and a legume (Persian clover, Trifolium repesinatum L or maple pea, Pisum sativum L), with a pasture [kikuyu grass (Pennisetum clandestinum) over-sown with short-rotation ryegrass (Lolium multiflorum L)] as a pasture control treatment. The experiment was a complete randomized block design with four replicates (,0·7 ha each). Annual dry-matter (DM) yield over the 3 years averaged >42 t ha,1 year,1 for the CFR treatment and >17 t ha,1 year,1 for the pasture treatment. The high DM yield of the CFR treatment resulted from >27 t ha,1 year,1 from maize harvested for silage and >15 t DM ha,1 year,1 utilized by grazing the forage rape and legumes. Total input of nitrogen (N) and water were similar for both treatments, resulting in higher N- and water-use efficiency for the CFR treatment, which was more than twice that for the pasture treatment. Overall, the nutritive value of the pasture treatment was slightly higher than the mean for that of the CFR treatment. The implications of these results are that a highly productive system based on the CFR treatment in conjunction with the use of pasture is achievable. Such a dairy production system in Australia could increase the total supply of feed resources grown on-farm and the efficiency of use of key resources such as N and water. [source] Modelling the concentrations of nitrogen and water-soluble carbohydrates in grass herbage ingested by cattle under strip-grazing managementGRASS & FORAGE SCIENCE, Issue 1 2008N. J. Hoekstra Abstract There is scope of increasing the nitrogen (N) efficiency of grazing cattle through manipulation of the energy and N concentrations in the herbage ingested. Because of asymmetric grazing by cattle between individual plant parts, it has not yet been established how this translates into the concentrations of N and water-soluble carbohydrates (WSC) in the herbage ingested. A model is described with the objective of assessing the efficacy of individual tools in grassland management in manipulating the WSC and N concentrations of the herbage ingested by cattle under strip-grazing management throughout the growing season. The model was calibrated and independently evaluated for early (April), mid- (June, regrowth phase) and late (September) parts of the growing season. There was a high correlation between predicted and observed WSC concentrations in the ingested herbage (R2 = 0·78, P < 0·001). The correlation between predicted and observed neutral-detergent fibre (NDF) concentrations in the ingested herbage was lower (R2 = 0·49, P < 0·05) with a small absolute bias. Differences in the N concentration between laminae and sheaths, and between clean patches and fouled patches, were adequately simulated and it was concluded that the model could be used to assess the efficacy of grassland management tools for manipulating the WSC and N concentrations in the ingested herbage. Model application showed that reduced rates of application of N fertilizer and longer rotation lengths were effective tools for manipulating herbage quality in early and mid-season. During the later part of the growing season, the large proportion of area affected by dung and urine reduced the effect of application rate of N fertilizer on herbage quality. In contrast, relative differences between high-sugar and low-sugar cultivars of perennial ryegrass were largest during this period. This suggests that high-sugar cultivars may be an important tool in increasing N efficiency by cattle when risks of N losses to water bodies are largest. The model output showed that defoliation height affects the chemical composition of the ingested herbage of both the current and the subsequent grazing period. [source] Effects of continuous or rotational grazing of two perennial ryegrass varieties on the chemical composition of the herbage and the performance of finishing lambsGRASS & FORAGE SCIENCE, Issue 3 2007C. L. Marley Abstract Plant breeding has developed perennial ryegrass varieties with increased concentrations of water-soluble carbohydrates (WSCs) compared with conventional varieties. Water-soluble carbohydrates are major metabolic and storage components in ryegrass. Therefore, if perennial ryegrass herbage is allowed to grow to greater heights it should contain higher water-soluble carbohydrates concentrations, for example as under rotational grazing rather than continuous grazing by livestock. This study investigated this hypothesis and measured the performance of lambs grazed rotationally and continuously. Replicated plots of the variety AberDart (bred to express high WSC concentrations) or the variety Fennema were grazed by a core group of ten male Cheviot lambs for 10 weeks. Lambs were weighed and replicate forage samples were taken every 7 d. Concentrations of WSC in AberDart herbage were significantly (P < 0·05), but not substantially, higher than those in Fennema herbage. Rotational grazing did not increase the differential in WSC concentration between the AberDart and Fennema varieties. However, there was a tendency (P = 0·07) for lambs rotationally grazing the AberDart swards to have a higher final live weight than lambs grazing the Fennema swards. Overall, lamb performance was increased when either perennial ryegrass variety was rotationally rather than continuously grazed (P < 0·001). [source] Genotypic variation in patterns of root distribution, nitrate interception and response to moisture stress of a perennial ryegrass (Lolium perenne L.) mapping populationGRASS & FORAGE SCIENCE, Issue 3 2007J. R. Crush Abstract Genotypic variation in patterns of root distribution, nitrate interception and response to moisture stress were assessed in both parents and 198 progeny of a perennial ryegrass (Lolium perenne L.) full-sibling mapping population. This was carried out in metre-deep tubes of sand culture in a glasshouse experiment. The proportion of root dry matter (DM) weight in the top 10 cm of sand ranged from 0·33 to 0·75 and values of log10(1 , K), where K is the constant for an exponential model relating root DM weight and root depth, also showed wide variation among genotypes. The proportion of a pulse of 15N recovered in whole plants ranged from 0·124 to 0·431. There was a positive linear correlation between the proportion of 15N recovered and plant total DM weight, but no relationship between nitrate interception and patterns of distribution of DM weight of roots. Some genotypes responded to moisture stress by increasing root growth, and in others root growth was inhibited. It is concluded that this below-ground variability in root variables may be an evolutionary adaptation by plant populations to survive heterogeneity in soil biotic and edaphic factors. [source] Liveweight gains of lambs from Caucasian clover/ryegrass and white clover/ryegrass swards on soils of high and low fertilityGRASS & FORAGE SCIENCE, Issue 2 2007A. D. Black Summary The high nutritive value and persistence under a wide range of climatic and soil fertility conditions make Caucasian clover a potentially useful forage legume but there is little information about the performance of livestock grazing Caucasian clover/grass swards. This study compared liveweight gains of lambs grazing Caucasian clover/perennial ryegrass and white clover/perennial ryegrass swards on high fertility (Olsen P 20 mg L,1, SO4 -S 12 mg kg,1) and low fertility (Olsen P 11 mg L,1, SO4 -S 7 mg kg,1) soils from 1998 to 2001 in the South Island of New Zealand. Mean annual liveweight gains were 1178 kg ha,1 for Caucasian clover/perennial ryegrass and 1069 kg ha,1 for white clover/perennial ryegrass swards at high fertility compared with 1094 kg ha,1 and 1015 kg ha,1, respectively, at low fertility. There was a higher mean proportion of clover in Caucasian clover/perennial ryegrass (0·19) than white clover/perennial ryegrass (0·11) swards, but there were no differences in total herbage production between the two clover/perennial ryegrass swards. The mean concentration of crude protein in the herbage of Caucasian clover (302 g kg DM,1) was higher than that in white clover (287 g kg DM,1) and grass herbage (227 g kg DM,1). Estimated mean metabolizable energy concentrations in the herbage were 12·5 MJ kg DM,1 for the two clovers and 11·6 MJ kg DM,1 for grass herbage. The difference in liveweight gain between swards on soils of high and low fertility was associated with an increase in total herbage production of similar composition and nutritive value, giving a greater number of grazing days for the swards on soils of high than low fertility. [source] Development and growth characteristics of Caucasian and white clover seedlings, compared with perennial ryegrassGRASS & FORAGE SCIENCE, Issue 4 2006A. D. Black Abstract Seedling competition for resources during establishment affects the potential success of individual species within a pasture. Germination, emergence and leaf expansion are key characteristics that contribute to the competitive ability of species. In this study, development and growth characteristics of Caucasian clover, white clover and perennial ryegrass (PRG) seedlings were quantified. A base temperature of <4°C and an optimum temperature of ,27°C were found for development in each species. Thermal time (Tt) requirements for 75% of final germination were lower for Caucasian clover (46°C d) and white clover (40°C d) than for PRG (76°C d), but Tt requirements for 50% of final emergence were similar (,110°C d). The phyllochron (°C d leaf,1) for primary stem leaves was slower for Caucasian clover (109°C d) than for white clover (94°C d) and PRG (101°C d). Appearance of the first PRG tiller, which indicates the initiation of secondary leaf development, occurred after 373°C d, compared with 532°C d for the first white clover stolon. Caucasian clover crown shoots did not develop until >1180°C d. Consequently, white clover and PRG had more leaves (,15 plant,1) and faster shoot relative growth rates (,0·062 mg mg,1 d,1) than Caucasian clover (5 leaves plant,1, 0·049 mg mg,1 d,1). [source] The relationship between tiller appearance in spring and contribution to dry-matter yield in perennial ryegrass (Lolium perenne L.) cultivars differing in heading dateGRASS & FORAGE SCIENCE, Issue 2 2005A. S. Laidlaw Abstract The relative contribution of tillers present in April and those appearing in consecutive periods in spring was assessed for perennial ryegrass cultivars in the three maturity groups (early, intermediate- and late-heading). Each group was represented by two diploid and one tetraploid cultivar each in plots in their third (2000) and fourth (2001) harvest years in three replicated blocks receiving an average of 325 kg N ha,1 and cut seven (in 2001) or eight (in 2000) times annually. ,Main' tillers and their daughters were marked with colour-coded PVC-covered wire loops in early April as were daughters which appeared in consecutive periods between harvests, the loop colour identifying the period of origin of the tiller. Tillers were harvested at cutting height (5 cm) before the plots were harvested and the herbage from tillers with the same colour code bulked per plot. Tillers were identified retrospectively as ,reproductive' if they had been decapitated at the previous harvest. Dry-matter yield was higher in the early than late-heading cultivars in April and early May but this was reversed in harvests in late May and June. The early heading group had a lower lamina content than the late-heading group during reproduction growth, both due to the reproductive tillers (mainly those which overwintered) having a lower leaf content and to their being fewer and smaller vegetative tillers during the reproductive phase than for the late-heading group. Turnover of tillers was high in spring due to decapitation of reproductive tillers and rapid post-flowering tillering. This was particularly pronounced in the early heading group which also had slightly more tillers marked in April which were subsequently decapitated than in the other maturity groups, i.e. 0·56 compared with 0·44 for the late-maturing group. Mean ratios of rate of death: rate of tillering for 3 years (1999,2001) for the early and late-heading groups were 0·8 and 0·4, respectively, for April,May and 1·1 and 2·4, respectively, for June indicating the different patterns in tiller turnover for the two extreme maturity groups. Information on tiller origin and contribution to yield can be used to refine tiller-based grass growth models. [source] Impact of seeding rate on annual ryegrass performanceGRASS & FORAGE SCIENCE, Issue 1 2004B. C. Venuto Abstract Annual ryegrass (Lolium multiflorum Lam.) is a primary forage resource for livestock producers throughout the south-eastern USA during the winter-growing season. It is important for livestock producers to begin grazing annual ryegrass as early as possible and any management practices maximizing early season production could be beneficial. To assess the impact of seeding rate on subsequent yield, yield distribution, quality, seedling density, and end-of-season plant and tiller density, a 2-year study was initiated at four locations in Louisiana. Three annual ryegrass cultivars, varying in seed size, were established at four seeding rates based on pure live seed (PLS) rates of 400, 800, 1200 and 1600 PLS m,2. There was no advantage in total yield from increasing seeding rates beyond 800 PLS m,2. However, first-harvest yields increased from 360 to 930 kg dry matter (DM) ha,1 as seeding rate increased from 400 to 1600 PLS m,2. Crude protein and neutral-detergent fibre concentrations, and in vitro DM digestibility, were not affected by seeding rate. Seedling density and end-of-season plant numbers increased as seeding rate increased. However, stems per plant decreased as seeding rate increased, indicating compensatory tillering for the reduced plant numbers observed at the lower seeding rates. These results indicate first-harvest yield can be increased by planting at higher seeding rates but total yields are not increased. [source] The influence of sward canopy structure on foraging decisions by grazing cattle.GRASS & FORAGE SCIENCE, Issue 2 2003Abstract Patch selection by grazing dairy cows in response to simultaneous variation in combinations of sward structural characteristics was examined in three experiments in which four mature dairy cows were offered a choice of patches (typically 0.9 m × 0.9 m) of perennial ryegrass (Lolium perenne) presented in a linear arrangement. Treatments involved combinations of variations in sward height, stubble height and/or depth of regrowth, prepared by preliminary cutting treatments. They were arranged in balanced sets of four to nine treatments, which were arranged in linear sequences of eighteen to twenty-seven patches. Within experiments, sequences were balanced across replicate sets of patches, which were grazed separately by individual cows. The number of bites removed and the residence time for each patch were highly correlated in all three experiments, and the results are reported using number of bites per patch as an estimator of foraging behaviour. In the first experiment, with vegetative swards, cows preferentially selected the tallest swards. When swards comprising reproductive stem were offered in Experiment 2, cows selectively grazed short-stubble swards rather than tall-stubble swards, although both offered a similar depth of regrowth. Cows did not exhibit preference for swards comprising the greatest quantity of leaf mass, indicating that the spatial distribution of plant components assumed greater importance. In the third experiment, the number of bites removed increased with increasing depth of regrowth, and was negatively correlated with sward height. The three patch-appraisal cues investigated were broadly ranked in order of importance as (i) depth of regrowth, (ii) sward maturity and (iii) sward height. There was no evidence, at least at a short temporal scale, that patch behaviour was influenced by conditions in adjacent patches, suggesting that the cows assessed grazing opportunities on a patch-by-patch basis. [source] Effects of white clover cultivar and companion grass on winter survival of seedlings in autumn-sown swardsGRASS & FORAGE SCIENCE, Issue 2 2003A. Elgersma Abstract The aim was to study the effects of white clover cultivar and combinations with perennial ryegrass cultivars on seedling establishment in autumn-sown swards and on winter survival of seedlings. Large-leaved white clover cv. Alice and small-leaved white clover cv. Gwenda, and an erect and a prostrate perennial ryegrass cultivar were sown in autumn in pure stands and as four binary grass-clover mixtures. Mixtures of white clover cv. Huia and Aberherald with perennial ryegrass were also sown. Companion grasses had no significant impact on the establishment of white clover. The number of seedlings of white clover cv. Alice in mixtures (335 m,2) was higher than cv. Gwenda (183 m,2) and pure swards had similar white clover population densities as mixed swards. White clover cv. Huia tended to have more seedlings than Aberherald (355 and 205 m,2 respectively). No stolons were produced prior to a severe winter, because of the late sowing date. Winter survival of clover seedlings was 0·56 in mixtures and 0·69 in pure stands, irrespective of white clover or companion grass cultivar. Stolon development of white clover in autumn is often considered essential for overwintering survival and spring growth. In this study, there was considerable survival of the non-stoloniferous tap-rooted seedlings of all four clover cultivars despite a severe winter. [source] The effect of sward structure as influenced by ryegrass genotype on bite dimensions and short-term intake rate by dairy cowsGRASS & FORAGE SCIENCE, Issue 1 2003P. D. Barrett Abstract The effects of genotypic variation in ryegrasses on sward structure, bite dimensions and intake rate by dairy cows were investigated. Two experiments were conducted. In Experiment 1, swards were in a vegetative state whereas, in Experiment 2, they were partly reproductive and were taller with higher herbage mass but lower leaf proportion than in Experiment 1. Applicability of relationships between sward structure and bite characteristics, previously established from artificial or hand-constructed swards, to field conditions were tested. Additional short-term intake rates and/or sward structural characteristics were considered as indicators of potential intake for use in protocols for the evaluation of grass varieties. Four cultivars were studied: AberElan, Twins (diploid and tetraploid perennial ryegrasses respectively), Polly, a hybrid ryegrass (perennial × Italian ryegrass) and Multimo (Italian ryegrass), each established in 200-m2 plots in four replicated blocks. Herbage intake rate was determined by short-term liveweight change (taking account of insensible weight loss) using 16 dairy cows allocated to four balanced groups with each plot grazed by one group for a 1-h assessment period. One block was grazed per day, over a 4-d experimental period, with each group grazing each variety in a complete crossover design. Sward characteristics and bite rate were also measured in both experiments. Bite dimensions were subsequently estimated, with bite depth being determined as a function of extended tiller height (ETH) in both experiments. Within both experiments, bite mass and intake rate did not differ significantly between swards of different cultivars despite swards containing Multimo generally having a higher ETH and water-soluble carbohydrate concentration and lower green leaf mass, sward bulk density and neutral-detergent fibre concentration than the other swards. However, bite depth was significantly higher (P < 0·01) in swards containing Multimo swards than in the others and, in Experiment 1, bite depth, as a proportion of ETH, was higher in swards containing Multimo and lower in those containing Twins than in the other two cultivars, whereas there was no difference in Experiment 2. Taking both experiments together, the mean bite depth was 0·5 of ETH with sward bulk density accounting for almost half the variance in the relationship between bite depth and ETH. The bulk density of the bite (bite mass per unit bite volume), measured in Experiment 2, followed a similar pattern to sward bulk density, increasing in the order Multimo, Polly, AberElan and Twins. It is concluded that the relationships between sward characteristics and bite dimensions, derived from artificial swards, are applicable to field swards, although the range in natural ryegrass sward characteristics is usually not as wide as in experiments using artificial swards. Lack of precision in the measurement of short-term intake and in sward-based measurements is likely to preclude their use in the evaluation of grass varieties. [source] Performance of white clover varieties combined in blends and alone when grown with perennial ryegrass under sheep and cattle grazingGRASS & FORAGE SCIENCE, Issue 1 2003T. A. Williams Abstract Two experiments were carried out to evaluate the performance of blends of three white clover (Trifolium repens L.) varieties in comparison with the component varieties and three other varieties sown individually in a mixture with perennial ryegrass (Lolium perenne L.). The plots were grazed rotationally in Experiment 1 by cattle and sheep and in Experiment 2 by sheep alone. In both experiments, the blend was composed of three medium-leaved varieties (AberDai, AberVantage and AberHerald), but with different relative contributions of the three varieties in the two experiments. Dry matter (DM) yields of white clover and perennial ryegrass were assessed in replicate plots for two years (1999 and 2000) after the establishment year. In Experiment 1, there was no significant difference between the DM yields of white clover or perennial ryegrass in either year. The decline in DM yield of white clover between years that was observed for some varieties was not found for the blend. In Experiment 2, significant differences were found in DM yields of white clover in both years. In 1999, AberDai had the highest DM yield. In 2000, AberHerald and AberVantage had the highest DM yields, and AberDai showed a decline in DM yield that was mirrored by the mean for all the white clover varieties. In both experiments, the blend did not show significantly higher DM yield than one or more of its components; indeed, in Experiment 2, it was significantly lower yielding than AberDai in 1999. However, where one component of the blend declined in DM yield between years, this was compensated for by an increase in the yield of another component. These preliminary findings suggest that the yield stability of blends may give them a potential role in agricultural practice. [source] Seed variation among annual ryegrass cultivars in south-eastern USA and the relationship with seedling vigour and forage productionGRASS & FORAGE SCIENCE, Issue 4 2002B. C. Venuto Abstract Annual ryegrass (Lolium multiflorum Lam.) is grown on more than one million ha in the south-eastern USA each year. Recommended and actual seeding rates vary substantially within the region. The objective of this study was to evaluate variation in seed weight, germination, seedling vigour and seasonal yield performance among annual ryegrass cultivars. During 1997, 1998 and 1999, seed from fourteen commercial cultivars was weighed and germinated to determine numbers of pure live seed (PLS) m,2 before yield evaluation at four locations. Seed from ten cultivars was planted at 0·7 and 2·0 cm depth in a greenhouse study to evaluate relative seedling vigour. Cultivar mean single-seed weight ranged from 2·4 to 4·8 mg in 1997, 1·8 to 4·5 mg in 1998, and 2·6 to 4·6 mg in 1999. Seed germination ranged from 78·8% to 98·0% in 1997, 82·3 to 98·3% in 1998 and 77·8 to 98·3% in 1999. Seed number, PLS m,2, ranged from 675 to 1289 in 1997, 710 to 1550 in 1998, and 717 to 1179 in 1999. Among the ten cultivars evaluated for seedling vigour, seedling weight differed between planting depths and a significant cultivar by year interaction was observed. Seedling weight was highly correlated with seed weight at each seeding depth. The effect of increasing number of PLS m,2 on subsequent yield performance, although small, was consistently negative. These results indicate that target plant populations may be obtained more economically by adjusting seeding rates for seed size differences among cultivars and seed lots of annual ryegrass. [source] Intake, liveweight gain and feed preference by steers fed combinations of lucerne and Westerwolds ryegrass silagesGRASS & FORAGE SCIENCE, Issue 1 2002E. Charmley Nutritive value and voluntary intake of legumes are generally considered to be higher than those of grasses when ensiled at similar digestibility, although high levels of soluble protein can result in low N utilization by animals and high losses to the environment. The objectives of this experiment were to describe the optimum combination of Westerwolds ryegrass (Lolium multiflorum Lam. cv. Aubade) and lucerne (Medicago sativa L. cv. AC Caribou) silages to maximize liveweight gain of steers fed silage, determine chemical components that are important and ascertain whether steers selected the optimum mixture when given a choice. Both silages contained similar concentrations of dry matter (DM), acid-detergent fibre (ADF) and organic acids, but lucerne silage had higher concentrations of N, soluble-N and ammonia-N. Westerwolds ryegrass silage contained more neutral-detergent fibre (NDF). In a 12-week experiment, voluntary intake by Hereford steers was not influenced when the proportion of the two silages was changed from 1 to 0 in 0·25 increments. However, liveweight gain and feed efficiency increased linearly (P < 0·001) as the proportion of ryegrass silage fed was increased. When preconditioned to either of the two silages, steers showed a significant preference for ryegrass over lucerne (P < 0·05). When conditioned to a mixture of both silages, no preference was elicited. It is suggested that extensive solubilization and deamination of protein in the lucerne silage may have caused the preference for Westerwolds ryegrass silage and the higher liveweight gains on diets containing higher proportions of Westerwolds ryegrass silage. [source] Fermentation characteristics and aerobic stability of grass silage inoculated with Lactobacillus buchneri, with or without homofermentative lactic acid bacteriaGRASS & FORAGE SCIENCE, Issue 4 2001F. Driehuis Aerobic spoilage by yeasts and moulds is a major cause of reduced nutritional value of silage and increases the risk of potential pathogenic microorganisms. Recent studies have shown that inoculation with Lactobacillus buchneri inhibits yeast growth and reduces the susceptibility to aerobic spoilage of various ensiled forages. The aim of this study was to determine whether these effects are retained when L. buchneri is added in combination with homofermentative lactic acid bacteria. In three experiments, silages were produced from perennial ryegrass [240,421 g kg,1 dry matter (DM)] inoculated with L. buchneri or L. buchneri plus a mixture of Pediococcus pentosaceus and Lactobacillus plantarum (inoculant PL). Uninoculated silage and silage inoculated with PL alone served as controls. Silages were examined for pH and DM loss in the course of ensilage and chemical and microbiological composition and aerobic stability after 3,4 months. L. buchneri plus PL and PL alone increased the initial rate of pH decline. L. buchneri alone and L. buchneri plus PL enhanced aerobic stability and, in general, reduced yeast and mould counts. In addition, these inoculants increased the final pH and DM loss and the concentrations of acetic acid and 1,2-propanediol (or propionic acid and 1-propanol instead of 1,2-propanediol), and decreased the concentration of lactic acid. The effects of L. buchneri on fermentation products increased with decreasing DM content. In silages of less than 270 g kg,1 DM, L. buchneri increased the ammonia-N concentration. It is suggested that this was associated with the relatively high final pH resulting from the high metabolic activity of L. buchneri in these silages. [source] Relationships between the yield of perennial ryegrass and of small-leaved white clover under cutting or continuous grazing by sheepGRASS & FORAGE SCIENCE, Issue 3 2001T. A. Williams Seven varieties or advanced breeding lines of white clover (Trifolium repens L.), all of small leaf size, were grown separately in mixtures with perennial ryegrass (Lolium perenne L.) in an experiment encompassing three harvest years. Harvestable dry-matter (DM) yield measurements were taken of these mixtures and of perennial ryegrass monocultures under two management regimes: cutting and continuous sheep grazing. Considerable differences were observed in the harvestable DM yields of white clover, perennial ryegrass and total yields of the mixtures between plots containing different white clover varieties. White clover yields were generally higher under cutting, and perennial ryegrass yields were higher under grazing. The difference between perennial ryegrass yield in monoculture and in mixture was variable. In the second harvest year, a significant interaction effect was seen between management and white clover variety for white clover yield but not for perennial ryegrass yield. The relationship between clover yield and grass yield differed between the two management regimes. Under cutting, a negative correlation was observed, indicative of competitive effects. However, under grazing, no such correlation was seen. Possible mechanisms underlying these outcomes are discussed. [source] How does the vertical and horizontal structure of a perennial ryegrass and white clover sward influence grazing?GRASS & FORAGE SCIENCE, Issue 2 2001P. Carrère Mixtures of perennial ryegrass (Lolium perenne L.) and white clover (Trifolium repens L.) sown in alternate rows or in a thoroughly mixed matrix were grazed by sheep, either continuously or during short grazing tests, and were used to investigate the influence of the vertical and horizontal components of the sward structure on defoliation by sheep. In an experiment under continuous grazing, the defoliation intensity was greater for white clover compared with perennial ryegrass leaves (0·80 and 0·58 respectively). In spring, perennial ryegrass leaves were more defoliated than white clover leaves, whereas the reverse was observed in summer. The ratio of the proportion of white clover to perennial ryegrass leaves grazed was negatively correlated with the difference between the surface height of the perennial ryegrass and white clover rows in spring. In both spring and summer, white clover leaves of the same extended leaf length had a higher proportion of them grazed than perennial ryegrass leaves. In another experiment, during short grazing tests with perennial ryegrass,white clover swards that were grazed at the same sward surface height and at the same white clover content as in the previous experiment, there were no significant differences in the proportion of white clover and perennial ryegrass leaves grazed between strips of the two species and thoroughly mixed structures. The proportion of white clover leaves grazed was higher than that of perennial ryegrass leaves. These results show that the differential defoliation by sheep of perennial ryegrass and white clover leaves varies according to their vertical distribution in the mixed canopy, but is little affected by their horizontal distribution. Even small differences in sward surface height between mixed perennial ryegrass and white clover can affect diet selection by sheep to a rather large extent. [source] The effect of accumulation period and harvest date in spring on dry-matter yield and forage quality in mixed swards containing Lolium spp. and Trifolium subterraneum in Western AustraliaGRASS & FORAGE SCIENCE, Issue 1 2001M. N. Callow The object of this study was to determine the effect of closing date and date of harvest for conservation (accumulation period), on dry-matter (DM) yield and forage quality of annual pasture in Western Australia. The field study comprised 48 plots, 2 m × 2 m, sown with either annual ryegrass (Lolium rigidum Gaud.) or Italian ryegrass (L. multiflorum Lam.), and mixed with subterranean clover (Trifolium subterraneum L.). Defoliation of swards until the end of winter was at the three leaves tiller,1 stage. In spring, once stem nodal development had commenced, swards were defoliated every 3,4 weeks. Swards were defoliated either twice with three leaves tiller,1 (accumulation period 1 commenced on 15 August); twice with three leaves tiller,1 and then once after 4 weeks (accumulation period 2 commenced on 11 September); twice with three leaves tiller,1 and then twice after 4-week intervals (accumulation period 3 commenced on 9 October) or; twice with 3 leaves tiller,1 and then twice after 4-week intervals and then once after 3 weeks (accumulation period 4 commenced on 30 October). From the commencement of the accumulation period, tiller density, DM yield and forage quality were determined weekly for up to 10 weeks. There was a positive quadratic association between DM yield and days after the commencement of the accumulation period. Yields were maximized from accumulation period 1 with 5·3, 6·6 and 9·5 t DM ha,1, and growth rates were 140, 128 and 145 kg DM ha,1 d,1, for Wimmera annual ryegrass and Richmond and Concord cultivars of Italian ryegrass respectively. In contrast, in vitro dry-matter digestibility (IVDMD) and crude protein (CP) content were negatively associated with days after the commencement of the accumulation period, and initial values were greater than 0·80 and 180 g kg DM,1 for IVDMD and CP content respectively. The rate of decline in IVDMD d,1 for Wimmera annual ryegrass was 0·005, 0·019 and 0·012 d,1 for accumulation periods 1, 2 and 3, respectively, while for Italian ryegrass cultivars Richmond was 0·015, 0·011, 0·02 and 0·012 d,1 and Concord was 0·014, 0·009, 0·013 and 0·01 d,1, for the 4 accumulation periods respectively. It is recommended that annual and Italian ryegrass pastures be harvested between 10% and 20% inflorescence emergence when IVDMD will exceed 0·70 regardless of cultivar and/or defoliation practice prior to the commencement of the accumulation period. [source] Effect of seed coating on plant growth and soil conditions: A preliminary study for restoration of degraded rangeland in the Qinghai,Tibetan Plateau, ChinaGRASSLAND SCIENCE, Issue 3 2010Yingchun Liu Abstract Rangeland degradation is a significant problem in the Qinghai,Tibetan Plateau, China. Restoration of the degraded rangelands through reseeding is being undermined by poor seedling growth under adverse soil moisture and nutrition conditions. Accordingly, seeds of Italian ryegrass (Lolium multiflorum) and Chinese milk vetch (Astragalus sinicus) were coated with a mixture of hygroscopic and plant-derived polysaccharide gums and alga powder (Phaeocystis sp. necolon-1), and inoculated with spores of microorganisms (Aspergillus sp. and Streptomyces sp.), serving as coating mixture decomposers, in order to improve rhizosphere moisture and nutrition. In a growth chamber simulating spring climate on the Plateau, seedling emergence of the coated seeds was 60,80 and 48,82%, respectively, for the two plant species in comparison to 38 and 24% for the uncoated seeds, which was due to moisture being supplied by the coat. In the outdoor pots with soil texture and moisture conditions similar to those of the degraded rangelands on the Plateau, dry weight of the plants from the coated seeds was 109,184 and 118,156 mg plant,1 for the respective plant species, while that of the plants from uncoated seeds was 18,20 and 10,11 mg plant,1. The number of Rhizobium sp. nodules on Chinese milk vetch plants from the coated seeds was 21,25 plant,1 while 0,2 in plants from uncoated seeds. Enhancement of plant growth was caused by increased activity of soil microbes. Plant growth on the soil after sampling plants from the coated seeds was also enhanced, indicating sustainability of improvement of soil conditions. Thus seed coating with hygroscopic nutrient-source materials and inoculation with microorganism spores as slow decomposers of the coating materials is a promising method for restoring degraded rangelands. [source] Contrasting infection frequencies of Neotyphodium endophyte in naturalized Italian ryegrass populations in Japanese farmlandsGRASSLAND SCIENCE, Issue 2 2010Masayuki Yamashita Abstract Neotyphodium endophytes often confer benefits to their host grasses and may enhance invasiveness of some grasses. The knowledge of infection frequencies of endophytes among invading weed populations is necessary to understand the relationships between endophyte infection and invasiveness. Here we present data on infection frequencies of Italian ryegrass (Lolium multiflorum Lam.), an important weed in some farmlands in Japan, persisting in contrasting farmlands: a terraced paddy field and a wheat-soybean double-cropped field in the western region of Shizuoka prefecture, Japan. The terraced paddy site is a mosaic of several landscape elements such as paddy fields, levees, fallow and abandoned fields, with a high percentage of non-crop area. Rice (Oryza sativa L.) has been cultivated for more than a decade with no application of chemical fertilizers, pesticides and fungicides. The wheat-soybean field is characterized by the aggregation of large-scaled fields that were originally reconstructed paddy fields, showing a low percentage of non-crop area. Wheat and soybean have been grown as winter and summer crops, respectively, using chemical fertilizers and herbicides. We examined the presence or absence of endophytes in a total of 1200 seeds sampled from the two Italian ryegrass populations. The terraced paddy population exhibited a markedly high infection frequency (91.0%), due possibly to selective feeding of non-infected seeds by insects. In contrast, the wheat-soybean farmland population showed almost no infection (1.1%), whereas the putative source of the invasion in the proximity exhibited a relatively high infection rate (64.4%). Such a micro-scale variation in infection frequencies may be attributable to a loss in endophyte viability within the wheat-soybean field. The findings suggest that endophyte infection frequency may markedly differ among the Italian ryegrass populations even within the same region, presumably depending on the abundance of the seed-eating insects, farmland management regimes and/or environmental conditions such as soil humidity. [source] Hybridization between perennial ryegrass and Italian ryegrass in naturalized Japanese populationsGRASSLAND SCIENCE, Issue 2 2008Hiroyuki Tobina Abstract Introduced Lolium species, including perennial ryegrass (Lolium perenne) and Italian ryegrass (Lolium multiflorum), have been widely utilized in Japan for forage, turf and soil conservation. These ryegrasses have escaped from cultivated areas and become naturalized, and this has become a serious issue in recent years. Interspecific hybrids between perennial ryegrass and Italian ryegrass have often been found in naturalized populations. It has also been suggested that hybridization between plant species might serve as a stimulus for the evolution of invasiveness. We surveyed the genetic structure of naturalized ryegrass populations in Japan using genetic markers that distinguished perennial ryegrass and Italian ryegrass. Of the 55 naturalized populations surveyed, 41 exhibited morphological traits of Italian ryegrass. DNA analysis using simple sequence repeat and chloroplast DNA markers characterized 20 of these 41 populations as Italian ryegrass, with the remaining populations as interspecific hybrid derivatives. Approximately half of the naturalized ryegrasses populations in Japan were inferred to include interspecific hybrids. [source] Seasonal differences in the adaptability of herbage species to environmental variations in a long-term grazing experimentGRASSLAND SCIENCE, Issue 1 2007Yiruhan Abstract Mixtures of orchardgrass, perennial ryegrass, tall fescue, redtop, Kentucky bluegrass, and white clover were sown in autumn 1973. Two 8-year grazing experiments were conducted at the National Grassland Research Institute (Nasushiobara, Japan) to determine the effects of grazing intensity and nitrogen levels on pasture ecosystems. These experiments involved two different grazing intensities (1974,1981) and two different nitrogen levels (1982,1989). Large spatiotemporal variations in phytomass due to environmental variations were observed in both experiments. Finlay,Wilkinson analysis was applied to clarify seasonal (monthly) differences in the adaptability of the herbage species, as measured by phytomass, to environmental variations by year and treatments in the two experiments. Seasonality in the adaptability to environments differed greatly among species. In this paper, we examined from livestock farmers' standpoint whether seasonality in adaptability of herbage species in the grazing pasture could be satisfied. A significantly high adaptability was shown for: orchardgrass from May to July and November; tall fescue in April, June and July, and November; redtop and Kentucky bluegrass in April; and Zoysia japonica in September and October. In contrast, perennial ryegrass and white clover exhibited very low adaptability in any season. Z. japonica and weeds such as Pennisetum alopecuroides, Eragrostis ferruginea and Digitaria adscendens, which had invaded from surrounding areas, showed low adaptability, except in autumn, when they showed moderate adaptability. [source] | ||||||||||||||||