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Ponderosa Pine Forest (ponderosa + pine_forest)
Selected AbstractsRecommendations for Integrating Restoration Ecology and Conservation Biology in Ponderosa Pine Forests of the Southwestern United StatesRESTORATION ECOLOGY, Issue 1 2006Reed F. Noss Abstract Over the past century, ponderosa pine,dominated landscapes of the southwestern United States have been altered by human activities such as grazing, timber harvest, road building, and fire exclusion. Most forested areas within these landscapes now show increased susceptibility to stand-replacing fires, insect outbreaks, and drought-related mortality. Recent large wildfires in the region have spurred public interest in large-scale fuel reduction and restoration programs, which create perceived and real conflicts with the conservation of biodiversity. Conservation concerns include the potential for larger road networks, soil and understory disturbance, exotic plant invasion, and the removal of large trees in treated areas. Pursuing prescribed burning, thinning, or other treatments on the broad scale that many scientists and managers envision requires the reconciliation of ecological restoration with biodiversity conservation. This study presents recommendations from a workshop for integrating the principles and practices of restoration ecology and conservation biology, toward the objective of restoring the composition, structure, and function of dry ponderosa pine forests. Planning on the scale of hundreds of thousands of hectares offers opportunities to achieve multiple objectives (e.g., rare species protection and restoration of ecological structures and processes) that cannot easily be addressed on a site-by-site basis. However, restoration must be coordinated with conservation planning to achieve mutual objectives and should include strict guidelines for protection of rare, declining, and sensitive habitats and species. [source] Short-Term Response of Land Birds to Ponderosa Pine RestorationRESTORATION ECOLOGY, Issue 4 2007William L. Gaines Abstract We monitored the short-term (>3 years) response of land birds to forest restoration treatments in Ponderosa pine forests located on the east slope of the North Cascade Range. Restoration treatments were designed to create stand structure and composition similar to pre-settlement forests, which were influenced by a frequent fire regime. Overall, avian community composition was changed as a result of the treatments. Cassin's Finch, Chipping Sparrow, and Yellow-rumped Warbler were found at higher densities in treated stands, whereas Mountain Chickadee, Western Tanager, and Red-breasted Nuthatch had higher densities in untreated stands. White-headed Woodpecker and Western Bluebird were only detected in the treated stands. Brown-headed Cowbird showed no response to treatments. We detected changes in the density of four of five foraging guilds in response to restoration treatments. Tree seedeaters, low understory and ground insectivores, and aerial insectivores all increased in density in treated stands. Overall, bark insectivores showed no density response to treatments. Tree foliage insectivore density was lower in treated than in untreated stands. Overall avian density, density of neotropical migrants, and density of some focal species were higher in treated stands. Monitoring should be continued to understand the longer-term (5,10 year) responses of land birds and to guide future forest restoration efforts. [source] A multi-trait test of the leaf-height-seed plant strategy scheme with 133 species from a pine forest floraFUNCTIONAL ECOLOGY, Issue 3 2010Daniel C. Laughlin Summary 1.,Westoby's [Plant and Soil (1998), 199, 213] Leaf-Height-Seed (LHS) plant strategy scheme quantifies the strategy of a plant based on its location in a three-dimensional space defined by three functional traits: specific leaf area (SLA), height, and seed mass. This scheme is based on aboveground traits and may neglect strategies of belowground resource capture if root functioning is not mirrored in any of the axes. How then do fine roots fit into the LHS scheme? 2.,We measured 10 functional traits on 133 plant species in a ponderosa pine forest in northern Arizona, USA. This data set was used to evaluate how well the LHS scheme accounts for the variation in above and belowground traits. 3.,The three most important plant strategies were composed of multiple correlated traits, but SLA, seed mass, and height loaded on separate principle components. The first axis reflected the widely observed ,leaf economics spectrum'. Species at the high end of this spectrum had high SLA, high leaf and fine root nitrogen (N) concentration, and low leaf dry matter content. The second axis reflected variation in seed mass and fine root morphology. Plants at the positive end of this spectrum were plants with large seeds and low specific root length (SRL). The third axis reflected variation in height and phenology. Plants at the positive end of this spectrum were tall species that flower late in the growing season. 4.,Leaf N concentration was positively correlated with fine root N concentration. SRL was weakly positively correlated with SLA. SRL was not correlated with fine root N concentration. Leaf litter decomposition rate was positively correlated with the leaf economics spectrum and was negatively correlated with the height and phenology spectrum. 5.,Leaf traits, seed mass, and height appear to be integrating properties of species that reflect much of the variation in plant function, including root function. Fine root N concentration was positively mirrored by the leaf economics spectrum, and SRL was inversely mirrored by seed mass. The leaf and height axes play a role in controlling leaf litter decomposability, indicating that these strategy axes have important consequences for ecosystem functioning. [source] Contrasting soil respiration in young and old-growth ponderosa pine forestsGLOBAL CHANGE BIOLOGY, Issue 12 2002J. IRVINE Abstract Three years of fully automated and manual measurements of soil CO2 efflux, soil moisture and temperature were used to explore the diel, seasonal and inter-annual patterns of soil efflux in an old-growth (250-year-old, O site) and recently regenerating (14-year-old, Y site) ponderosa pine forest in central Oregon. The data were used in conjunction with empirical models to determine which variables could be used to predict soil efflux in forests of contrasting ages and disturbance histories. Both stands experienced similar meteorological conditions with moderately cold wet winters and hot dry summers. Soil CO2 efflux at both sites showed large inter-annual variability that could be attributed to soil moisture availability in the deeper soil horizons (O site) and the quantity of summer rainfall (Y site). Seasonal patterns of soil CO2 efflux at the O site showed a strong positive correlation between diel mean soil CO2 efflux and soil temperature at 64 cm depth whereas diel mean soil efflux at the Y site declined before maximum soil temperature occurred during summer drought. The use of diel mean soil temperature and soil water potential inferred from predawn foliage water potential measurements could account for 80% of the variance of diel mean soil efflux across 3 years at both sites, however, the functional shape of the soil water potential constraint was site-specific. Based on the similarity of the decomposition rates of litter and fine roots between sites, but greater productivity and amount of fine litter detritus available for decomposition at the O site, we would expect higher rates of soil CO2 efflux at the O site. However, annual rates were only higher at the O site in one of the 3 years (597 ± 45 vs. 427 ± 80 g C m,2). Seasonal patterns of soil efflux at both sites showed influences of soil water limitations that were also reflected in patterns of canopy stomatal conductance, suggesting strong linkages between above and below ground processes. [source] Carbon storage and fluxes in ponderosa pine forests at different developmental stagesGLOBAL CHANGE BIOLOGY, Issue 7 2001B.E. Law Abstract We compared carbon storage and fluxes in young and old ponderosa pine stands in Oregon, including plant and soil storage, net primary productivity, respiration fluxes, eddy flux estimates of net ecosystem exchange (NEE), and Biome-BGC simulations of fluxes. The young forest (Y site) was previously an old-growth ponderosa pine forest that had been clearcut in 1978, and the old forest (O site), which has never been logged, consists of two primary age classes (50 and 250 years old). Total ecosystem carbon content (vegetation, detritus and soil) of the O forest was about twice that of the Y site (21 vs. 10 kg C m,2 ground), and significantly more of the total is stored in living vegetation at the O site (61% vs. 15%). Ecosystem respiration (Re) was higher at the O site (1014 vs. 835 g C m,2 year,1), and it was largely from soils at both sites (77% of Re). The biological data show that above-ground net primary productivity (ANPP), NPP and net ecosystem production (NEP) were greater at the O site than the Y site. Monte Carlo estimates of NEP show that the young site is a source of CO2 to the atmosphere, and is significantly lower than NEP(O) by c. 100 g C m,2 year,1. Eddy covariance measurements also show that the O site was a stronger sink for CO2 than the Y site. Across a 15-km swath in the region, ANPP ranged from 76 g C m,2 year,1 at the Y site to 236 g C m,2 year,1 (overall mean 158 ± 14 g C m,2 year,1). The lowest ANPP values were for the youngest and oldest stands, but there was a large range of ANPP for mature stands. Carbon, water and nitrogen cycle simulations with the Biome-BGC model suggest that disturbance type and frequency, time since disturbance, age-dependent changes in below-ground allocation, and increasing atmospheric concentration of CO2 all exert significant control on the net ecosystem exchange of carbon at the two sites. Model estimates of major carbon flux components agree with budget-based observations to within ±,20%, with larger differences for NEP and for several storage terms. Simulations showed the period of regrowth required to replace carbon lost during and after a stand-replacing fire (O) or a clearcut (Y) to be between 50 and 100 years. In both cases, simulations showed a shift from net carbon source to net sink (on an annual basis) 10,20 years after disturbance. These results suggest that the net ecosystem production of young stands may be low because heterotrophic respiration, particularly from soils, is higher than the NPP of the regrowth. The amount of carbon stored in long-term pools (biomass and soils) in addition to short-term fluxes has important implications for management of forests in the Pacific North-west for carbon sequestration. [source] Understory vegetation response to mechanical mastication and other fuels treatments in a ponderosa pine forestAPPLIED VEGETATION SCIENCE, Issue 2 2010Jeffrey M. Kane Abstract Questions: What influence does mechanical mastication and other fuel treatments have on: (1) canopy and forest floor response variables that influence understory plant development; (2) initial understory vegetation cover, diversity, and composition; and (3) shrub and non-native species density in a second-growth ponderosa pine forest. Location: Challenge Experimental Forest, northern Sierra Nevada, California, USA. Methods: We compared the effects of mastication only, mastication with supplemental treatments (tilling and prescribed fire), hand removal, and a control on initial understory vegetation response using a randomized complete block experimental design. Each block (n=4) contained all five treatments and understory vegetation was surveyed within 0.04-ha plots for each treatment. Results: While mastication alone and hand removal dramatically reduced the midstory vegetation, these treatments had little effect on understory richness compared with control. Prescribed fire after mastication increased native species richness by 150% (+6.0 species m2) compared with control. However, this also increased non-native species richness (+0.8 species m2) and shrub seedling density (+24.7 stems m2). Mastication followed by tilling resulted in increased non-native forb density (+0.7 stems m2). Conclusions: Mechanical mastication and hand removal treatments aided in reducing midstory fuels but did not increase understory plant diversity. The subsequent treatment of prescribed burning not only further reduced fire hazard, but also exposed mineral soil, which likely promoted native plant diversity. Some potential drawbacks to this treatment include an increase of non-native species and stimulation of shrub seed germination, which could alter ecosystem functions and compromise fire hazard reduction in the long-term. [source] Forest Stand Dynamics and Livestock Grazing in Historical ContextCONSERVATION BIOLOGY, Issue 5 2005MICHAEL M. BORMAN clima; incendio forestal; pastoreo histórico; pino ponderosa; supresión de fuego Abstract:,Livestock grazing has been implicated as a cause of the unhealthy condition of ponderosa pine forest stands in the western United States. An evaluation of livestock grazing impacts on natural resources requires an understanding of the context in which grazing occurred. Context should include timing of grazing, duration of grazing, intensity of grazing, and species of grazing animal. Historical context, when and under what circumstances grazing occurred, is also an important consideration. Many of the dense ponderosa pine forests and less-than-desirable forest health conditions of today originated in the early 1900s. Contributing to that condition was a convergence of fire, climate, and grazing factors that were unique to that time. During that time period, substantially fewer low-intensity ground fires (those that thinned dense stands of younger trees) were the result of reduced fine fuels (grazing), a substantial reduction in fires initiated by Native Americans, and effective fire-suppression programs. Especially favorable climate years for tree reproduction occurred during the early 1900s. Exceptionally heavy, unregulated, unmanaged grazing by very large numbers of horses, cattle, and sheep during the late nineteenth and early twentieth centuries occurred in most of the U.S. West and beginning earlier in portions of the Southwest. Today, livestock numbers on public lands are substantially lower than they were during this time and grazing is generally managed. Grazing then and grazing now are not the same. Resumen:,El pastoreo de ganado ha sido implicado como una causa de la mala salud de los bosques de pino ponderosa en el occidente de Estados Unidos. La evaluación de los impactos del pastoreo sobre los recursos naturales requiere de conocimiento del contexto en que ocurrió el pastoreo. El contexto debe incluir al período de ocurrencia, la duración y la intensidad del pastoreo, así como la especie de animal que pastoreó. El contexto histórico, cuando y bajo que circunstancias ocurrió el pastoreo, también es una consideración importante. Muchos de los bosques densos de pino ponderosa y de las condiciones, menos que deseables, de salud de los bosques actuales se originaron al principio del siglo pasado. Contribuyó a esa condición una convergencia de factores, fuego, clima y pastoreo, que fueron únicos en ese tiempo. Durante ese período, hubo sustancialmente menos incendios superficiales de baja intensidad (que afectaron a grupos densos de árboles más jóvenes) como resultado de la reducción de combustibles finos (pastoreo), una reducción sustancial en los incendios iniciados por Americanos Nativos y programas efectivos de supresión de incendios. Al inicio del siglo pasado hubo años con clima especialmente favorable para la reproducción de árboles. Al final del siglo diecinueve y comienzo del veinte hubo pastoreo no regulado ni manejado, excepcionalmente intensivo, por una gran cantidad de caballos, reses y ovejas en la mayor parte del oeste de E.U.A. y aun antes en porciones del suroeste. En la actualidad, el número de semovientes en terrenos públicos es sustancialmente menor al de ese tiempo, y el pastoreo generalmente es manejado. El pastoreo entonces y el pastoreo ahora no son lo mismo. [source] Contrasting soil respiration in young and old-growth ponderosa pine forestsGLOBAL CHANGE BIOLOGY, Issue 12 2002J. IRVINE Abstract Three years of fully automated and manual measurements of soil CO2 efflux, soil moisture and temperature were used to explore the diel, seasonal and inter-annual patterns of soil efflux in an old-growth (250-year-old, O site) and recently regenerating (14-year-old, Y site) ponderosa pine forest in central Oregon. The data were used in conjunction with empirical models to determine which variables could be used to predict soil efflux in forests of contrasting ages and disturbance histories. Both stands experienced similar meteorological conditions with moderately cold wet winters and hot dry summers. Soil CO2 efflux at both sites showed large inter-annual variability that could be attributed to soil moisture availability in the deeper soil horizons (O site) and the quantity of summer rainfall (Y site). Seasonal patterns of soil CO2 efflux at the O site showed a strong positive correlation between diel mean soil CO2 efflux and soil temperature at 64 cm depth whereas diel mean soil efflux at the Y site declined before maximum soil temperature occurred during summer drought. The use of diel mean soil temperature and soil water potential inferred from predawn foliage water potential measurements could account for 80% of the variance of diel mean soil efflux across 3 years at both sites, however, the functional shape of the soil water potential constraint was site-specific. Based on the similarity of the decomposition rates of litter and fine roots between sites, but greater productivity and amount of fine litter detritus available for decomposition at the O site, we would expect higher rates of soil CO2 efflux at the O site. However, annual rates were only higher at the O site in one of the 3 years (597 ± 45 vs. 427 ± 80 g C m,2). Seasonal patterns of soil efflux at both sites showed influences of soil water limitations that were also reflected in patterns of canopy stomatal conductance, suggesting strong linkages between above and below ground processes. [source] Carbon storage and fluxes in ponderosa pine forests at different developmental stagesGLOBAL CHANGE BIOLOGY, Issue 7 2001B.E. Law Abstract We compared carbon storage and fluxes in young and old ponderosa pine stands in Oregon, including plant and soil storage, net primary productivity, respiration fluxes, eddy flux estimates of net ecosystem exchange (NEE), and Biome-BGC simulations of fluxes. The young forest (Y site) was previously an old-growth ponderosa pine forest that had been clearcut in 1978, and the old forest (O site), which has never been logged, consists of two primary age classes (50 and 250 years old). Total ecosystem carbon content (vegetation, detritus and soil) of the O forest was about twice that of the Y site (21 vs. 10 kg C m,2 ground), and significantly more of the total is stored in living vegetation at the O site (61% vs. 15%). Ecosystem respiration (Re) was higher at the O site (1014 vs. 835 g C m,2 year,1), and it was largely from soils at both sites (77% of Re). The biological data show that above-ground net primary productivity (ANPP), NPP and net ecosystem production (NEP) were greater at the O site than the Y site. Monte Carlo estimates of NEP show that the young site is a source of CO2 to the atmosphere, and is significantly lower than NEP(O) by c. 100 g C m,2 year,1. Eddy covariance measurements also show that the O site was a stronger sink for CO2 than the Y site. Across a 15-km swath in the region, ANPP ranged from 76 g C m,2 year,1 at the Y site to 236 g C m,2 year,1 (overall mean 158 ± 14 g C m,2 year,1). The lowest ANPP values were for the youngest and oldest stands, but there was a large range of ANPP for mature stands. Carbon, water and nitrogen cycle simulations with the Biome-BGC model suggest that disturbance type and frequency, time since disturbance, age-dependent changes in below-ground allocation, and increasing atmospheric concentration of CO2 all exert significant control on the net ecosystem exchange of carbon at the two sites. Model estimates of major carbon flux components agree with budget-based observations to within ±,20%, with larger differences for NEP and for several storage terms. Simulations showed the period of regrowth required to replace carbon lost during and after a stand-replacing fire (O) or a clearcut (Y) to be between 50 and 100 years. In both cases, simulations showed a shift from net carbon source to net sink (on an annual basis) 10,20 years after disturbance. These results suggest that the net ecosystem production of young stands may be low because heterotrophic respiration, particularly from soils, is higher than the NPP of the regrowth. The amount of carbon stored in long-term pools (biomass and soils) in addition to short-term fluxes has important implications for management of forests in the Pacific North-west for carbon sequestration. [source] Arbuscular mycorrhizal propagule densities respond rapidly to ponderosa pine restoration treatmentsJOURNAL OF APPLIED ECOLOGY, Issue 1 2003Julie E. Korb Summary 1Mycorrhizae form a critical link between above-ground plants and the soil system by influencing plant nutrition, nutrient cycling and soil structure. Understanding how mycorrhizae respond to disturbances may lead to important advances in interpreting above-ground plant recovery. 2The inoculum potential for arbuscular mycorrhizae (AM) and ectomycorrhizal (EM) fungi was investigated in thinned-only, thinned and prescribed burned (both restoration treatments) and unthinned and unburned control stands in northern Arizona ponderosa pine forests. The relationships between mycorrhizal fungal propagule densities and plant community and soil properties were quantified. 3The relative amount of infective propagules of AM fungi was significantly higher in samples collected from both restoration treatments than their paired controls (unthinned and unburned stands). In contrast, the same restoration treatments had no significant effect on the relative amount of infective propagules of EM fungi. 4The relative amount of infective propagules of AM fungi was significantly positively correlated with graminoid cover and herbaceous understorey species richness and negatively correlated with overstorey tree canopy cover and litter cover. 5Synthesis and applications. These results indicate that population densities of AM fungi can rapidly increase following restoration treatments in northern Arizona ponderosa pine forests. This has important implications for restoring the herbaceous understorey of these forests because most understorey plants depend on AM associations for normal growth. These results also can be applied to other ecosystems that are in a state of restoration or where the role of fire is just beginning to be understood. [source] Recommendations for Integrating Restoration Ecology and Conservation Biology in Ponderosa Pine Forests of the Southwestern United StatesRESTORATION ECOLOGY, Issue 1 2006Reed F. Noss Abstract Over the past century, ponderosa pine,dominated landscapes of the southwestern United States have been altered by human activities such as grazing, timber harvest, road building, and fire exclusion. Most forested areas within these landscapes now show increased susceptibility to stand-replacing fires, insect outbreaks, and drought-related mortality. Recent large wildfires in the region have spurred public interest in large-scale fuel reduction and restoration programs, which create perceived and real conflicts with the conservation of biodiversity. Conservation concerns include the potential for larger road networks, soil and understory disturbance, exotic plant invasion, and the removal of large trees in treated areas. Pursuing prescribed burning, thinning, or other treatments on the broad scale that many scientists and managers envision requires the reconciliation of ecological restoration with biodiversity conservation. This study presents recommendations from a workshop for integrating the principles and practices of restoration ecology and conservation biology, toward the objective of restoring the composition, structure, and function of dry ponderosa pine forests. Planning on the scale of hundreds of thousands of hectares offers opportunities to achieve multiple objectives (e.g., rare species protection and restoration of ecological structures and processes) that cannot easily be addressed on a site-by-site basis. However, restoration must be coordinated with conservation planning to achieve mutual objectives and should include strict guidelines for protection of rare, declining, and sensitive habitats and species. [source] |