Open Areas (open + area)

Distribution by Scientific Domains
Distribution within Life Sciences


Selected Abstracts


Meteorological factors affecting the diversity of airborne algae in an urban atmosphere

ECOGRAPHY, Issue 5 2006
Naveen K. Sharma
Aeroalgal sampling of Varanasi City, India, was done using a Tilak Rotorod sampler and exposing agarised Bold basal medium Petri plates during March 2003 to February 2005. Amongst the 34 airborne algal genera recorded, cyanobacteria dominated the aero-algal flora, followed by green algae and diatoms. The generic diversity of airborne algae as well as the constituting groups exhibited seasonal variation. The most favored period for the appearance of cyanobacteria in the air was summer, while winter favored green algae. Presence of diatoms was almost uniform throughout the year. The presence of algal particles in the air depended upon the abundance and dynamics of algal source and their release and dispersal in the atmosphere. Best model selection with Akaike information criteria indicated temperature, relative humidity, rainfall, wind velocity as the most important climatic factors determining algal diversity. These factors exert their effect both directly by influencing entrainment and dispersal of algae from the source, and indirectly by regulating the dynamics of the possible algal source (soil, water, plant body, wall and roof of the building) by supporting or inhibiting the algal growth. In a closed environment and at low altitude sampling site characteristic is also an important factor. Open area near to the countryside had maximal aero-algal diversity. [source]


Woody plants modulate the temporal dynamics of soil moisture in a semi-arid mesquite savanna,

ECOHYDROLOGY, Issue 1 2010
Daniel L. Potts
Abstract Climate variability and human activities interact to increase the abundance of woody plants in arid and semi-arid ecosystems worldwide. How woody plants interact with rainfall to influence patterns of soil moisture through time, at different depths in the soil profile and between neighboring landscape patches is poorly known. In a semi-arid mesquite savanna, we deployed a paired array of sensors beneath a mesquite canopy and in an adjacent open area to measure volumetric soil water content (,) every 30 min at several depths between 2004 and 2007. In addition, to quantify temporally dynamic variation in soil moisture between the two microsites and across soil depths we analysed , time-series using fast Fourier transforms (FFT). FFT analyses were consistent with the prediction that by reducing evaporative losses through shade and reducing rainfall inputs through canopy interception of small rainfall events, the mesquite canopy was associated with a decline in high-frequency (hour-to-hour and day-to-day) variation in shallow ,. Finally, we found that, in both microsites, high-frequency , variation declined with increasing soil depth as the influence of evaporative losses and inputs associated with smaller rainfall events declined. In this case, we argue that the buffering of shallow soil moisture against high-frequency variations can enhance nutrient cycling and alter the carbon cycle in dryland ecosystems. Copyright © 2009 John Wiley & Sons, Ltd. [source]


Temporal and shrub adaptation effect on soil microbial functional diversity in a desert system

EUROPEAN JOURNAL OF SOIL SCIENCE, Issue 6 2009
V. Saul-Tcherkas
Summary The Negev Desert is characterized by spatial and temporal patterns of resource distribution, in which soil biota are considered to be among the most sensitive biological characteristics, easily influenced by changes related to soil and abiotic factors. Soil water availability and organic matter are among the most important factors, acting as triggers that determine the length of the period of activity. The main source of organic matter in this xeric environment is input from annual and perennial shrubs. In order to persist and propagate in this xeric environment, the plants have developed different ecophysiological adaptations (e.g. the excretion of salt (Reaumuria negevensis) and chemical compounds (Artemisia sieberi) via the leaves). We found that the values of soil moisture obtained for soil samples collected in the vicinity of R. negevensis were larger than for samples collected in the vicinity of Noaea mucronata and A. sieberi and in the open area. The maximum values of CO2 evolution, microbial biomass and Shannon index (H,) were obtained for the samples collected from the vicinity of N. mucronata. Therefore, we assume that the vicinity of N. mucronata afforded the best conditions for the soil bacterial community. In the Negev Desert, we also found that water availability and pulses of rain compared with frequent rainfall influenced CO2 evolution, microbial biomass, qCO2 and the Shannon index (H,). The differences in water amount and availability between the two rainy seasons caused larger values in most of the properties during the first four seasons (December 2005,November 2006) compared with the last four seasons (December 2006,November 2007) for most of the samples. [source]


Novel microfluidic platform for culturing neurons: Culturing and biochemical analysis of neuronal components

BIOTECHNOLOGY JOURNAL, Issue 11 2009
Jeong Won Park
Abstract Neurons, one of the most polarized types of cells, are typically composed of cell bodies (soma), dendrites, and axons. Many events such as electric signal transmission, axonal transport, and local protein synthesis occur in the axon, so that a method for isolating axons from somata and dendrites is required for systematically investigating these axonal events. Based on a previously developed neuron culture method for isolating and directing the growth of central nervous system axons without introducing neutrophins, we report three modified microfluidic platforms: (1) for performing biochemical analysis of the pure axonal fraction, (2) for culturing tissue explants, and (3) a design that allows high content assay on same group of cells. The key feature of these newly developed platforms is that the devices incorporate a number of microgrooves for isolating axons from the cell body. They utilize an open cellculture area, unlike the enclosed channels of the previous design. This design has extended the axonal channel so that a sufficient amount of pure axonal fraction can be obtained to perform biochemical analysis. The design also addresses the drawback of the previous neuron culture device, which was not adaptable for culturing thick neuronal tissues such as brain explants, neurospheres, and embryoid bodies, which are essential model tissues in neuroscience research. The design has an open cellculture area in the center and four enclosed channels around open area, and is suitable for multiple drug screening assays. [source]


Conservation value of degraded habitats for forest birds in southern Peninsular Malaysia

DIVERSITY AND DISTRIBUTIONS, Issue 5 2006
Kelvin S.-H.
ABSTRACT Clearance of tropical forest for agricultural purposes is generally assumed to seriously threaten the survival of forest species. In this study, we quantified the conservation value, for forest bird species, of three degraded habitat types in Peninsular Malaysia, namely rubber tree plantations, oil palm plantations, and open areas. We surveyed these degraded habitats using point counts to estimate their forest bird species richness and abundance. We assessed whether richness, abundance, and activities of different avian dietary groups (i.e. insectivores and frugivores) varied among the habitats. We identified the critical habitat elements that accounted for the distribution of forest avifauna in these degraded habitats. Our results showed that these habitats harboured a moderate fraction of forest avifauna (approximately 46,76 species) and their functions were complementary (i.e. rubber tree plantations for moving; open habitats for perching; shrubs in oil palm plantations for foraging). In terms of species richness and abundance, rubber tree plantations were more important than oil palm plantations and open habitats. The relatively high species richness of this agricultural landscape was partly due to the contiguity of our study areas with extensive forest areas. Forecasts of forest-species presence under various canopy cover scenarios suggest that leaving isolated trees among non-arboreal crops could greatly attract relatively tolerant species that require tree canopy. The conservation value of degraded habitats in agricultural landscapes seems to depend on factors such as the type of crops planted and distance to primary forest remnants. [source]


Combining information from range use and habitat selection: sex-specific spatial responses to habitat fragmentation in tawny owls Strix aluco

ECOGRAPHY, Issue 2 2006
Peter Sunde
How individuals respond to habitat heterogeneity is usually measured as variation in range size and by ranking the relative importance of habitat types (habitat selection). The combined effect of how individuals incorporate different habitat types in their home ranges and allocate their time budget between them is rarely derived. Additionally, when home range size varies between individuals, habitat selection analyses might be flawed if foraging decisions are based on variation in absolute rather than proportional availability. We investigated the suitability of standard analytical approaches by measuring the spatial responses of tawny owls to habitat fragmentation. These owls inhabited woodland of various sizes, representing a fragmentation gradient from open farmland with small, isolated woodland patches, to continuous woodland within their home ranges. In 17 territories within open farmland, the available area covered by woodland increased with the square root of the area of open land embraced in the home range. The owls did not display functional response in habitat selection, but females selected woodland more strongly than males. Females utilised woodland 10 times more intensively in farmland than in continuous woods, whereas males utilised farmland woods 3.2 times more intensively. Moreover, females in farmland exploited woodland 3.2 times as intensively as males, apparently because of higher travel costs in open areas. Since the extensive variation in intensity of use as a function of total availability was not indicated from the analysis of habitat selection, we suggest that information about intensity of use be more widely used as a supplementary measure of habitat use patterns than appears to be the practice at present. [source]


Post-fire recovery of ant communities in Submediterranean Pinus nigra forests

ECOGRAPHY, Issue 2 2006
Anselm Rodrigo
This study analyzes the variations in the structure and composition of ant communities in burned Pinus nigra forests in central Catalonia (NE Spain). Pinus nigra forests do not recover after fire, changing to shrublands and oak coppices. For this reason, we suggest that ant communities of burned P. nigra forests will change after fire, because the post-fire scenario, in particular with the increase of open areas, is different to the unburned one, and more favourable for some species than for others. In four locations previously occupied by P. nigra forests where different fires occurred 1, 5, 13 and 19 yr before the sampling, we sampled the structure and composition of ant communities with pitfall traps, tree traps and net sweeping in unburned plots and in plots affected by canopy and understory fire. The results obtained suggest that canopy and understory fire had little effect on the structure of ant communities. Thus, many variables concerning ant communities were not modified either by fire type (understory or canopy fire) or by time since fire. However, a number of particular species were affected, either positively or negatively, by canopy fire: three species characteristic of forest habitats decreased after fire, while eight species characteristic of open habitats increased in areas affected by canopy fire, especially in the first few years after fire. These differences in ant community composition between burned and unburned plots imply that the maximum richness is achieved when there is a mixture of unburned forests and areas burned with canopy fire. Moreover, as canopy cover in P. nigra forests burned with canopy fire is not completed in the period of time studied, the presence of the species that are characteristic of burned areas remains along the chronosequence studied, while the species that disappear after fire do not recover in the period of time considered. Overall, the results obtained indicate that there is a persistent replacement of ant species in burned P. nigra forests, as is also the case with vegetation. [source]


Effects of temperature and elevation on habitat use by a rare mountain butterfly: implications for species responses to climate change

ECOLOGICAL ENTOMOLOGY, Issue 4 2009
SARAH ASHTON
Abstract 1.,The present study used the mountain specialist butterfly Parnassius apollo as a model system to investigate how climate change may alter habitat requirements for species at their warm range margins. 2.,Larval habitat use was recorded in six P. apollo populations over a 700 m elevation gradient in the Sierra de Guadarrama (central Spain). Larvae used four potential host species (Sedum spp.) growing in open areas amongst shrubs. 3.,Parnassius apollo host-plant and habitat use changed as elevation increased: the primary host shifted from Sedum amplexicaule to Sedum brevifolium, and larvae selected more open microhabitats (increased bare ground and dead vegetation, reduced vegetation height and shrub cover), suggesting that hotter microhabitats are used in cooler environments. 4.,Larval microhabitat selection was significantly related to ambient temperature. At temperatures lower than 27 °C, larvae occupied open microhabitats that were warmer than ambient temperature, versus more shaded microhabitats that were cooler than ambient conditions when temperature was higher than 27 °C. 5.,Elevational changes in phenology influenced the temperatures experienced by larvae, and could affect local host-plant favourability. 6.,Habitat heterogeneity appears to play an important role in P. apollo larval thermoregulation, and may become increasingly important in buffering populations of this and other insect species against climatic variation. [source]


New perspectives on Holocene landscape development in the southern English chalklands: The upper Allen valley, Cranborne Chase, Dorset

GEOARCHAEOLOGY: AN INTERNATIONAL JOURNAL, Issue 2 2005
C. French
A combination of on- and off-site paleoenvironmental and archaeological investigations of the upper Allen valley of Dorset, conducted from 1998,2002, has begun to indicate a different model of prehistoric landscape development to those previously put forward for this part of the southern English chalk downlands. Woodland growth in the earlier Holocene appears to have been slower and patchier than the presumed model of full climax deciduous woodland rapidly attained in a warming environment. With open areas still strongly present in the Mesolithic, the area witnessed its first exploitation, thus slowing and altering soil development. Consequently, many areas perhaps never developed thick, well-structured, brown forest earths, but more probably thin brown earths. By the later Neolithic period, these soils had become thin rendzinas, largely as a consequence of human exploitation and the predominance of pastoral land use. The early presence of thinner and less well-developed soils over large areas of downland removes the necessity for envisaging extensive soil erosion and the accumulation of thick colluvial and alluvial deposits in the dry valleys and valley floor as often postulated. If there were major changes in the vegetation and soil complexes in this area of chalk downland, these had already occurred by the Neolithic rather than the Bronze Age as often suggested, and the area has remained relatively stable ever since. This has major implications for models of prehistoric land use in the southern chalkland region, such as a much greater degree of stability in prehistoric and historic times, variability within sub-regions, and differences between different parts of the chalk downlands than had previously been envisaged. © 2005 Wiley Periodicals, Inc. [source]


Long-term investigations of the snow cover in a subalpine semi-forested catchment

HYDROLOGICAL PROCESSES, Issue 2 2006
Manfred Stähli
Abstract To improve spring runoff forecasts from subalpine catchments, detailed spatial simulations of the snow cover in this landscape is obligatory. For more than 30 years, the Swiss Federal Research Institute WSL has been conducting extensive snow cover observations in the subalpine watershed Alptal (central Switzerland). This paper summarizes the conclusions from past snow studies in the Alptal valley and presents an analysis of 14 snow courses located at different exposures and altitudes, partly in open areas and partly in forest. The long-term performance of a physically based numerical snow,vegetation,atmosphere model (COUP) was tested with these snow-course measurements. One single parameter set with meteorological input variables corrected to the prevailing local conditions resulted in a convincing snow water equivalent (SWE) simulation at most sites and for various winters with a wide range of snow conditions. The snow interception approach used in this study was able to explain the forest effect on the SWE as observed on paired snow courses. Finally, we demonstrated for a meadow and a forest site that a successful simulation of the snowpack yields appropriate melt rates. Copyright © 2006 John Wiley & Sons, Ltd. [source]


Estimating winter evaporation in boreal forests with operational snow course data

HYDROLOGICAL PROCESSES, Issue 8 2003
Angela Lundberg
Abstract Snow course measurements from 11 sites located in eastern and northern Finland were used to estimate the total interception evaporation of a winter season for different forest categories. We categorized the sites based on forest density and tree species. Results showed that interception loss from gross precipitation increased with forest density and approached 30% for a forest with the highest density class. Interception loss for the most common forest density class was 11%. Interception losses were slightly larger in spruce forests than in pine, deciduous, or mixed forests. We provide suggestions as to how to design snow surveys to estimate wintertime interception evaporation better. Rough terrain and transition zones between forest and open areas should be avoided. Since evaporation fraction was strongly dependent on tree crown characteristics, snow course data should include direct estimates of canopy closure. Qualitative observations made by different observers should be given a reference frame to ensure comparability of records from different sites. Copyright © 2003 John Wiley & Sons, Ltd. [source]


Habitat-specific demography and source,sink dynamics in a population of Siberian jays

JOURNAL OF ANIMAL ECOLOGY, Issue 1 2010
Magdalena Nystrand
Summary 1.,There are a number of models describing population structure, many of which have the capacity to incorporate spatial habitat effects. One such model is the source,sink model, that describes a system where some habitats have a natality that is higher than mortality (source) and others have a mortality that exceeds natality (sink). A source can be maintained in the absence of migration, whereas a sink will go extinct. 2.,However, the interaction between population dynamics and habitat quality is complex, and concerns have been raised about the validity of published empirical studies addressing source,sink dynamics. In particular, some of these studies fail to provide data on survival, a significant component in disentangling a sink from a low quality source. Moreover, failing to account for a density-dependent increase in mortality, or decrease in fecundity, can result in a territory being falsely assigned as a sink, when in fact, this density-dependent suppression only decreases the population size to a lower level, hence indicating a ,pseudo-sink'. 3.,In this study, we investigate a long-term data set for key components of territory-specific demography (mortality and reproduction) and their relationship to habitat characteristics in the territorial, group-living Siberian jay (Perisoreus infaustus). We also assess territory-specific population growth rates (r), to test whether spatial population dynamics are consistent with the ideas of source,sink dynamics. 4.,Although average mortality did not differ between sexes, habitat-specific mortality did. Female mortality was higher in older forests, a pattern not observed in males. Male mortality only increased with an increasing amount of open areas. Moreover, reproductive success was higher further away from human settlement, indicating a strong effect of human-associated nest predators. 5.,Averaged over all years, 76% of the territories were sources. These territories generally consisted of less open areas, and were located further away from human settlement. 6.,The source,sink model provides a tool for modelling demography in distinct habitat patches of different quality, which can aid in identifying key habitats within the landscape, and thus, reduce the risk of implementing unsound management decisions. [source]


Elevation and forest clearing effects on foraging differ between surface , and subterranean , foraging army ants (Formicidae: Ecitoninae)

JOURNAL OF ANIMAL ECOLOGY, Issue 1 2009
Anjali Kumar
Summary 1Forest fragmentation often results in a matrix of open areas mixed with patches of forest. Both biotic and abiotic factors can affect consumer species' ability to utilize the altered habitat, especially for species that range over large areas searching for prey. 2Army ants (Formicidae: Ecitoninae) are highly mobile top predators in terrestrial Neotropical ecosystems. Army ant foraging behaviour is influenced by forest clearing at lowland sites, and clearing can reduce army ant population persistence. 3Because high temperatures are implicated in hindering above-ground army ant foraging, we predicted that forest clearing effects on army ant foraging would be reduced at higher (cooler) elevations in montane forest. We also predicted that subterranean foraging, employed by some army ant species, would buffer them from the negative effects of forest clearing. 4We quantified the foraging rates of above-ground and underground foraging army ants at eight sites along an elevational gradient from 1090 to 1540 m a.s.l. We asked whether these two foraging strategies cause a difference in the ability of army ants to forage in open matrix areas relative to elevationally matched forested habitats, and whether elevation predicts open area vs. forest foraging rate differences. 5As predicted, army ants that forage above-ground had lower foraging rates in open areas, but the open area vs. forest difference declined with elevation. In contrast, underground foragers were not affected by habitat type, and underground foraging rates increased with elevation. Ground surface temperatures were higher in open areas than forested areas. Temperatures declined with elevation, and temperature differences between open and forested areas decreased with elevation. 6We conclude that army ants that forage above-ground may be restricted to forested areas due to a thermal tolerance threshold, but that they are released from this limitation at higher elevations. We further suggest that underground foraging permits some army ants to persist within modified landscapes. Our findings have implications for the effects of habitat modification and climate change on these top predators. [source]


Spatial patterns of tree recruitment in a relict population of Pinus uncinata: forest expansion through stratified diffusion

JOURNAL OF BIOGEOGRAPHY, Issue 11 2005
J. Julio Camarero
Abstract Aim, To infer future changes in the distribution of isolated relict tree populations at the limit of a species' geographical range, a deep understanding of the regeneration niche and the spatial pattern of tree recruitment is needed. Location, A relict Pinus uncinata population located at the south-western limit of distribution of the species in the Iberian System of north-eastern Spain. Methods,Pinus uncinata individuals were mapped within a 50 × 40-m plot, and their size, age and reproductive status were estimated. Data on seed dispersal were obtained from a seed-release experiment. The regeneration niche of the species was assessed based on the associations of seedling density with substrate and understorey cover. The spatial pattern of seedlings was described using point-pattern (Ripley's K) and surface-pattern (correlograms, Moran's I) analyses. Statistical and inverse modelling were used to characterize seedling clustering. Results, Pine seedlings appeared aggregated in 6-m patches. Inverse modelling estimated a longer mean dispersal distance (27 m), which corresponded to the size of a large cluster along the north to north-eastward direction paralleled by an eastward trend of increasing seedling age. The two spatial scales of recruitment were related to two dispersal processes. The small-scale clustering of seedlings was due to local seed dispersal in open areas near the edge of Calluna vulgaris mats: the regeneration niche. The long-range expansion might be caused by less frequent medium-distance dispersal events due to the dominant north-westerly winds. Main conclusions, To understand future range shifts of marginal tree populations, data on seed dispersal, regeneration niche and spatial pattern of recruitment at local scales should be obtained. The monitoring of understorey communities should be a priority in order to predict correctly shifts in tree species range in response to global warming. [source]


Scattered trees as modifiers of agricultural landscapes: the role of waddeessa (Cordia africana Lam.) trees in Bako area, Oromia, Ethiopia

AFRICAN JOURNAL OF ECOLOGY, Issue 2009
Abebe Yadessa
Abstract Scattered trees in general and scattered waddeessa (Cordia africana Lam.) trees in particular are very common across the agricultural landscapes in Oromia, Ethiopia. A study on this scattered waddeessa trees commonly growing on farmers' agricultural fields was conducted at Bako in western Oromia, Ethiopia with the objective of assessing their role in modifying the soil properties in the agricultural landscape. Soil samples from surface layers (0,10 cm) were taken at three concentric transects (0.5, 2 and 4 m) around the tree and compared with soil samples from the adjacent open areas (15 m distance from the tree), and then analysed following the standard procedures. Results showed that scattered waddeessa trees significantly modified the overall properties of the soil in the agricultural landscape of Bako area. But soil texture was not affected, indicating that it is more related to parent material than the tree influence. Hence, the soil patches observed under these waddeessa trees can be important local nutrient reserves that may influence the rural agricultural landscape. They also play an important role in generating local household income from the sale of products and conserving biodiversity by providing habitats and resources that are otherwise absent or scarce in agricultural landscape. [source]


The role of water abundance, thermoregulation, perceived predation risk and interference competition in water access by African herbivores

AFRICAN JOURNAL OF ECOLOGY, Issue 3 2008
Marion Valeix
Abstract In African savannas, surface water can become limiting and an understanding of how animals address the trade-offs between different constraints to access this resource is needed. Here, we describe water access by ten African herbivore species in Hwange National Park, Zimbabwe, and we explore four possible determinants of the observed behaviours: water abundance, thermoregulation, perceived predation risk and interference competition. On average, herbivores were observed to drink in 80% of visits to a waterhole. The probability of drinking was higher in 2003 (474 mm) than in 2004 (770 mm), and at the end of the dry season than at its beginning. For larger species, this probability may also be related to risks of interference competition with elephants or other herbivores. For smaller species, this probability may also be related to the perceived risk of predation. We also investigate the time spent accessing water to drink. The influence of herd size and the presence of young on the time spent accessing water for most species suggests that perceived predation risk plays a role. Thermoregulation also affects this time: during the hottest periods, herbivores spend less time in open areas, unless when wind is strong, probably owing to evapotranspired heat loss. Résumé Dans les savanes africaines, l'eau de surface disponible peut devenir un facteur limitant et il est nécessaire de comprendre comment les animaux agissent face aux différentes contraintes que pose l'accčs ŕ cette ressource. Nous décrivons ici l'accčs ŕ l'eau de dix herbivores africains du Parc National de Hwange, au Zimbabwe, et nous explorons quatre facteurs qui sont peut-ętre déterminants dans les comportements observés: l'abondance de l'eau, la thermorégulation, le risque de prédation ressenti et la compétition/ interférence. En moyenne, on a observé que les herbivores buvaient lors de 80% de leurs visites au point d'eau. La probabilité qu'ils boivent étai plus forte en 2003 (474 mm) qu'en 2004 (770 mm), et ŕ la fin de la saison sčche qu'au début. Pour les plus grandes espčces, cette probabilité pourrait aussi ętre liée aux risques de compétition par interférence avec les éléphants ou d'autres herbivores. Pour les plus petites espčces, cette probabilité pourrait aussi ętre liée au risque de prédation ressenti. Nous avons aussi étudié le temps passéŕ se rendre au point d'eau pour y boire. L'influence de la taille du groupe et de la présence de jeunes sur le temps pris par la plupart des espčces pour se rendre au point d'eau laisse penser que la perception du risque de prédation joue un rôle. La thermorégulation affecte aussi cette durée: pendant les périodes les plus chaudes, les herbivores passent moins de temps dans les espaces ouverts, sauf si le vent est fort, probablement ŕ cause de la perte de chaleur par évapotranspiration. [source]


Case study of a one-sided attack by multiple troop members on a nontroop adolescent male and the death of Japanese macaques (Macaca fuscata)

AGGRESSIVE BEHAVIOR, Issue 4 2009
Masaki Shimada
Abstract An adolescent wild male Japanese macaque (Macaca fuscata), following Kinkazan A troop, was attacked one-sidedly by multiple members of the troop. The victim was identified as PI, and was estimated to be seven±one year old. The aggressive interaction was recorded by video camera until the end. Although at least 16 troop members approached PI more than once, only three males (one adult, two adolescents) of A troop attacked him. PI kept crouching throughout the attack, then escaped to the shore and dived into the sea. The interaction continued for more than one hour. PI was found dead a few hours after the end of interaction. The damage caused by the assailants was not the direct cause of PI's death; it was due to hypothermia caused by drifting in the sea. PI's life history was reconstructed from past records. PI was a normal adolescent male who migrated from an all-male group around B1 troop and started ranging around A troop. The aggressive interaction is believed to be a typical example of conflict between troop males and a nontroop male. The interaction period was very long compared with previous reports on such conflicts among Japanese macaques. PI kept crouching in open areas, exposing himself as a potential competitor for the resources of the troop, and did not show any submissive or reconciliatory behavior toward the troop males. This may be why the troop males did not stop the attack. Aggr. Behav. 35:334,341, 2009. © 2009 Wiley-Liss, Inc. [source]


Seed dynamics of the mast seeding tussock grass Ampelodesmos mauritanica in Mediterranean shrublands

JOURNAL OF ECOLOGY, Issue 3 2000
Montserrat Vilŕ
Summary 1,The Mediterranean perennial grass Ampelodesmos mauritanica may have the potential to expand its range. We analysed temporal variability of its reproductive components (seedfall, seed bank, seed predation, seed germination, seedling emergence, survival and growth) in three microsites (open areas, beneath Ampelodesmos and beneath shrubs) at two sites. 2,Reproductive components prior to seedling emergence were both closely linked and very similar between microsites within a site. Seedling survival and growth differed between microsites, being lowest in open areas. Recruitment patterns cannot therefore be predicted from seedfall. 3,Abundant seed production in 1996 was followed by successful germination and high seedling survival. In the following (non-masting) year, although Ampelodesmos has a low-density persistent seed bank, recruitment was much lower because germination was low and post-dispersal seed predation was high. 4,Our results suggest that Ampelodesmos reproduction is episodic. Expansion of its distribution may be triggered by intermittent seedling recruitment following masting, but is otherwise constrained by seed limitation, post-dispersal seed predation and a loss of viability in the seed bank. [source]


Density and habitat associations of Henslow's Sparrows wintering in saline soil barrens in southern Arkansas

JOURNAL OF FIELD ORNITHOLOGY, Issue 4 2008
William C. Holimon
ABSTRACT Although the habitat requirements of breeding populations of Henslow's Sparrow (Ammodramus henslowii) have been examined, less is known about their habitat requirements and ecology during the nonbreeding season. We estimated population densities and quantified habitat associations of Henslow's Sparrows wintering in saline soil barrens in southern Arkansas. Densities of Henslow's Sparrows in the saline soil barrens were similar to those in the Longleaf Pine (Pinus palustris) Ecosystem of the southeastern United States, considered by many to be their primary wintering habitat. Henslow's Sparrows were closely associated with open areas with greater cover of Aristida spp. and globe beaksedge (Rhynchospora globularis), greater stem density at 11,20 cm above ground, more lichens, more herbaceous cover, more bare ground, greater occurrence of little bluestem (Schizacyrium scoparium) as the tallest vegetation, less moss, and less shrub cover than randomly selected sites. In contrast to the results of studies conducted in the Longleaf Pine Ecosystem, the presence of Henslow's Sparrows in our study was not correlated with the height of the tallest vegetation. Our results indicate that saline soil barrens of southern Arkansas support a high density of wintering Henslow's Sparrows and do so for longer postdisturbance periods than longleaf pine savanna. We also found that stem density near the ground was similar to that reported from longleaf pine savanna, but only about half that observed on their breeding grounds. Areas used by Henslow's Sparrows had more lichen and less moss cover, suggesting that those areas were drier than random sites within the barrens. Further research is needed to determine if large populations of Henslow's Sparrows winter in other saline soil barrens and if fire influences habitat associations and densities in the barrens. SINOPSIS Aunque los requisitos de hábitat para las poblaciones reproductivas de Ammodramus henslowii han sido determinadas, se conoce muy poco sobre su ecología y requerimientos de hábitat durante la temporada no-reproductiva. Estimamos la densidad poblacional y cuantificamos el hábitat asociado a Gorrión de Henslow que pasan el invierno en un salitral con suelo empobrecido en el sur de Arkansas. La densidad de las aves en el salitral resultó similar a lo encontrado en Ecosistemas de Pinos (Pinus palustris) en el sureste de los EUA, considerado por muchos como el principal hábitat invernal para la especie. Los gorriones estuvieron altamente asociados a áreas abiertas con covertura de Aristida spp. y Rhynchospora globularis, con mayor densidad de tallos, altura entre 11,20 cm sobre el suelo, mayor cantidad de líquenes, mayor cubierta herbácea, más suelo desnudo, mayor presencia de Schzacyrium scoparium (como la vegetación de mayor tamańo), menos musgos, y menos arbustos que localidades seleccionadas al azar. En contraste a los resultados de estudios conducidos en Ecosistemas de Pinos, la presencia del gorrión en nuestra área de estudio no estuvo correlacionada con la altura de la vegetación de mayor tamańo. Nuestros resultados indican que las salinas en Arkansas sostienen una alta densidad de aves invernales, y lo hacen por periodos más largos, después de disturbios, que en las savanas de pinos. También encontramos que la densidad de tallos, cerca del suelo, era similar a la informada en savanas de pinos, pero tan solo la mitad de lo indicado para lugares en donde las aves se reproducen. Las áreas utilizadas tienen más líquenes, pero menos musgos, lo que sugiere que dichas áreas son más secas que lugares con suelo empobrecido muestreados al azar. Se necesitan más trabajos para determinar si otras grandes poblaciones del gorrión de Henslow pasan el invierno en otras salinas con suelos empobrecidos y si eventos como fuegos incluyen en la asociación del hábitat y densidades en los lugares con suelo empobrecido. [source]


Spatial patterns of association at local and regional scales in coastal sand dune communities

JOURNAL OF VEGETATION SCIENCE, Issue 5 2009
Estelle Forey
Abstract Questions: Are positive understorey-dominant associations important in physically severe dune communities and does the strength of positive associations vary with disturbance at the local scale and with stress at the regional scale? Do associational patterns observed at the neighbourhood scale predict diversity at higher scales? Location: Coastal sand dunes, Aquitaine (France). Methods: Associational patterns with five dominant species were recorded along a local gradient of disturbance and a 240-km long regional gradient. Density, richness, cover and variance ratio of understorey species were recorded in quadrats located in dominant and in open areas. Spatial pattern of dominant plant species was recorded using a distance-based method. Results: Positive understorey-dominant associations were most frequent at both regional and local scale, although negative associations with understorey species were observed for one of the five dominants. At the regional scale, there was a shift in the magnitude of spatial associations, with higher positive associations in the most stressful sites, whereas spatial associations where not affected by the local disturbance gradient. Positive associations were not related to the size of the dominants but rather influenced by the identity of the dominant species. Conclusions: Our study highlights the potential crucial role of facilitation together with the importance of turnover of the dominants in explaining large-scale variation in diversity. However, because positive associations may also be attributed to environmental heterogeneity or co-occurrence of microhabitat preferences of species, experiments are needed to fully assess the relative importance of facilitation versus other drivers of community diversity. [source]


Facilitation of tree saplings by nurse plants: Microhabitat amelioration or protection against herbivores?

JOURNAL OF VEGETATION SCIENCE, Issue 2 2008
Lorena Gómez-Aparicio
Abstract Question: Positive interactions are predicted to be common in communities developing under high physical stress or high herbivory pressure due to neighbour amelioration of limiting physical and consumer stresses, respectively. However, when both stress sources meet in the same community, the relative importance of the two facilitation mechanisms is poorly understood. We ask: What is the relative importance of abiotic vs. biotic mechanisms of facilitation of tree saplings by shrubs in Mediterranean mountain forests? Location: Sierra Nevada, SE Spain (1800,1850 m a.s.l.) Methods: Saplings of four tree taxa (Acer opalus ssp. grana-tense, Quercus ilex, Pinus nigra ssp. salzmanii and P. sylvestris var. nevadensis) were planted following a 2 × 2 factorial design: two levels of herbivory (control and ungulate exclusion) and two microhabitats (under shrubs and in open areas). Sapling survival and growth were monitored for five years. Results: Shrubs had positive effects on sapling survival both in control and ungulate excluded plots. This effect was species-specific, with shrubs increasing the survival of Acer opalus and Quercus ilex three and twofold, respectively, but having a minor effect on the Pinus species. Herbivory damage was also species-specific, being much higher for Acer opalus than for any other species. Shrubs did not protect saplings of any species against ungulates. Thus, all Acer saplings (the most damaged species) suffered herbivory outside the exclosures, which largely reduced sapling height. Conclusions: Protection from abiotic stress (summer drought and winter frost) was much more relevant than protection from biotic stress (herbivory). However, we propose that the final balance between the two mechanisms can be expected to vary strongly between sites, depending on the relative magnitude of the different sources of stress and the intrinsic traits (e.g. palatability) of the species interacting. [source]


The influence of Prosopis canopies on understorey vegetation: Effects of landscape position

JOURNAL OF VEGETATION SCIENCE, Issue 5 2003
J.D. Schade
Abstract. The influence of canopy trees and shrubs on under-storey plants is complex and context-dependent. Canopy plants can exert positive, negative or neutral effects on production, composition and diversity of understorey plant communities, depending on local environmental conditions and position in the landscape. We studied the influence of Prosopis velutina (mesquite) on soil moisture and nitrogen availability, and understorey vegetation along a topographic gradient in the Sonoran Desert. We found significant increases in both soil moisture and N along the gradient from desert to riparian zone. In addition, P. velutina canopies had positive effects, relative to open areas, on soil moisture in the desert, and soil N in both desert and intermediate terrace. Biomass of understorey vegetation was highest and species richness was lowest in the riparian zone. Canopies had a positive effect on biomass in both desert and terrace, and a negative effect on species richness in the terrace. The effect of the canopy depended on landscape position, with desert canopies more strongly influencing soil moisture and biomass and terrace canopies more strongly influencing soil N and species richness. Individual species distributions suggested interspecific variation in response to water- vs. N-availability; they strongly influence species composition at both patch and landscape position levels. [source]


Nucleation and facilitation in salt pans in Mediterranean salt marshes

JOURNAL OF VEGETATION SCIENCE, Issue 6 2001
A.E. Rubio-Casal
Tutin et al. (1992) Abstract. Arthrocnemum macrostachyum is a perennial species acting as a primary colonizer of salt pans in Mediterranean high salt marshes. Salicornia ramosissima, an annual, occurs in salt pans under Arthrocnemum canopies and in open areas. The aim of this study was to analyse, in wild populations and a transplant experiment, how S. ramosissima population dynamics and growth are affected by A. macrostachyum. The environmental conditions within the patches of Arthrocnemum were less stressful than in the open areas, with lower radiation levels and salinity concentrations. In the inner areas of A. macrostachyum patches, density-dependent mortality processes of S. ramosissima seedlings led to low densities of adult individuals with greater morphological development and reproductive success than in open areas. However, at the edges of Arthrocnemum patches facilitation of seedling survival favoured high densities. Environmental stress hindered development, decreased reproduction and premature death. These results are in agreement with the general theory of factors controlling vegetation distribution that biotic interactions dominate in low stress environments, while abiotic interactions dominate under harsher environmental conditions. A. macrostachyum plays an essential role in the succession in these salt pans, facilitating seed production and stimulating nucleation processes in S. ramosissima. [source]


Anti-predator behaviour, space use and habitat selection in female roe deer during the fawning season in a wolf area

JOURNAL OF ZOOLOGY, Issue 3 2008
P. Bongi
Abstract This study investigated the anti-predator strategies adopted by 19 radio-collared female roe deer during the fawning season by monitoring their spatial behaviour and habitat selection by means of radio-tracking. The study was carried out in a forest area of the Apennine Mountains (central Italy), where wolves are natural predators of roe deer and in summer fawns are more frequently predated than adult roe deer. The presence of fawns was monitored by means of direct observations. Roe deer fawns are known to adopt the hiding strategy during the lactation period, when they lie concealed for long periods waiting for their mothers' milk. As a consequence of this, the home-range sizes of mothers were significantly smaller than those of non-mothers during the summer lactation only. In contrast, no significant difference was found in spring, when fawns were absent, or autumn, when they were already weaned. In order to increase the success of the hiding strategy adopted by their fawns against wolf predation, mothers selected denser habitats (deciduous coppice forests) that provided denser undergrowth vegetation and limited visibility. In doing so, mothers traded open areas for forests throughout the fawning season. Indeed, mothers made significant habitat selection throughout the monitored period, and this was marked after the birth of fawns. In contrast, non-mothers generally used habitat types according to their availability. During lactation, the correlation between habitat use by mothers and habitat visibility (assessed using the form of a standard-sized roe deer) was inversely significant. These results taken together highlight the importance of anti-predator strategies adopted by roe deer mothers during the critical phase for fawn survival. [source]


Post-breeding migrations of newts (Triturus cristatus and T. marmoratus) with contrasting ecological requirements

JOURNAL OF ZOOLOGY, Issue 3 2000
R. Jehle
Abstract Two newt species with contrasting ecological requirements (Triturus cristatus and T. marmoratus) were radio-tracked after leaving a shared breeding pond in western France. Movements of 30 individuals were recorded over 491 days. During the first terrestrial night, the newts migrated with high directionality up to 137 m to their refuges. Burrows of small mammals were among the favourite refuges. Movements after the first night were mostly underground and over shorter distances (< 6.8 m). Sixty-four per cent of all tracked newts stayed within 20 m of the pond edge, and migrations were recorded up to 146 m away from the pond. Both species preferred areas with bushes, hedgerows and trees, and avoided pastures and open areas. Migration in the direction of a habitat type characterized by trees and underground shelters was favoured over migrations in other directions. A clear terrestrial niche separation between the two newt species was not observed. The results are discussed in relation to previous findings that T. cristatus supersedes T. marmoratus as a consequence of anthropogenic change to the terrestrial environment, affecting the species differentially. [source]


Seed removal in two coexisting oak species: ecological consequences of seed size, plant cover and seed-drop timing

OIKOS, Issue 9 2008
Ignacio M. Pérez-Ramos
Seed predation and dispersal can critically influence plant community structure and dynamics. Inter-specific differences arising at these early stages play a crucial role on tree recruitment patterns, which in turn could influence forest dynamics and species segregation in heterogeneous environments such as Mediterranean forests. We investigated removal rates from acorns set onto the ground in two coexisting Mediterranean oak species ,Quercus canariensis and Q. suber, in southern Spain. We developed maximum likelihood estimators to investigate the main factors controlling probabilities of seed removal and to describe species-specific functional responses. To account for inter-specific differences in seed-drop timing, two experiments were established: a simultaneous exposure of acorns of the two species (synchronous experiments) and a seed exposure following their natural seed-drop phenology (diachronic experiments). A total of 1536 acorns were experimentally distributed along a wide and natural gradient of plant cover, and removal was periodically monitored for three months at two consecutive years (with contrasting differences in seed production and thus seed availability on the ground). The probability of seed removal increased with plant cover (leaf area index, LAI) for the two oak species. Inter-specific differences in acorn removal were higher in open areas and disappeared in closed microhabitats, especially during a non-mast year. Despite later seed-drop, Q. suber acorns were removed faster and at a higher proportion than those of Q. canariensis. The higher probability of seed removal for this species could be attributed to its larger seed size compared to Q. canariensis, as inter-specific differences were less pronounced when similar sized acorns were exposed. Inter-specific differences in seed removal, arising from seed size variability and microsite heterogeneity, could be of paramount importance in oak species niche separation, driving stand dynamics and composition along environmental gradients. [source]


Relationship between seed and clonal growth in the reproduction of Carex rugulosa Kük. in riverside meadows

PLANT SPECIES BIOLOGY, Issue 2 2008
SATORU ARAKI
Abstract Carex rugulosa Kük. develops riverside vegetation in brackish marshes along the lower parts of rivers. Because the rhizomatous ramets grow densely (590,950/m2) in its developed population, it is expected that seedlings cannot get enough light to grow if seeds germinate in crowded ramets. We studied the reproductive contribution of seed in a marshy meadow of the Ohashi River, Japan. The number of seeds produced was 6900,14 200/m2 in 2002. The number of buried seeds in the following germination season differed among sites from 1190 to 2690/m2. No seedlings were observed at plots where the ramets from rhizomes grew densely. In contrast, seedlings appeared in plots where all rhizomatous ramets were artificially cut. The number of seedlings corresponded to 17.5,39.5% of that of buried seeds. All these seedlings were submerged and died when it rained heavily. In the genotype analysis using allozyme detection, one of the phosphoglucoisomerase loci (Pgi-2) indicated a pair of alleles. Among eight plots (each plot was 50 or 60-cm square), four showed an excess of heterozygotes and two showed homozygote domination in the Pgi-2 genotype. These results suggest that: (i) seed germination is suppressed in crowded ramets; (ii) seedling survival is severely reduced by inundation; and (iii) the population is usually maintained by clonal growth. As germination is induced in open areas, it is hypothesized that the main role of seed is recovery after vegetation decline caused by environmental stresses or colonization to other areas without dense vegetation. [source]


Use of Shrubs as Nurse Plants: A New Technique for Reforestation in Mediterranean Mountains

RESTORATION ECOLOGY, Issue 2 2002
Jorge Castro
Abstract Common techniques currently used for afforestation in the Mediterranean basin consider the pre-existing vegetation (mainly shrubs) as a source of competition for trees, and consequently it is generally eliminated before planting. Nevertheless, it has been demonstrated that woody plants can facilitate the establishment of understory seedlings in environments that, like the Mediterranean area, are characterized by a pronounced dry season. In this study, we experimentally analyze the usefulness of shrubs as nurse plants for afforestation of two native conifers, Pinus sylvestris L. (Scots pine) and Pinus nigra Arnold (black pine). Two-year-old seedlings were planted in four microhabitats: (1) open interspaces without vegetation (which is the usual method used in afforestation programs), (2) under individuals of Salvia lavandulifolia, (3) under the north side of spiny shrubs, and (4) under the south side of spiny shrubs. Pine survival was remarkably higher when planted under individuals of the shrub S. lavandulifolia (54.8% for Scots pine, 81.9% for black pine) compared with open areas (21.5% for Scots pine, 56.8% for black pine; chi square, p < 0.05). The survival of both pines was also higher when planted on the north side of spiny shrubs, although the survival on the south side was similar to that found in open areas. In addition, pine growth was not inhibited when planted in association with shrubs. This pattern appears to result from the combination of abiotic conditions imposed by the presence of a nurse shrub, which leads to improvement in seedling water status and therefore reduced summer mortality by drought. The results show that the use of shrubs as nurse plants is a technique that offers both economic and ecological advantages, in terms of savings in labor and plant material and reduced and even negligible impact on the pre-existing vegetation. [source]


Effect of agro-forestry and landscape changes on common buzzards (Buteo buteo) in the Alps: implications for conservation

ANIMAL CONSERVATION, Issue 1 2005
Fabrizio Sergio
In Italy, pre-Alpine forests, once managed through coppice silviculture, are being converted to mature woodland, while land abandonment is causing woodland expansion and erosion of open habitats. Based on habitat-selection analyses, we predicted the impact of such changes on common buzzards (Buteo buteo), which depend on forested and open areas for nesting and foraging. Compared to availability, at a micro-scale buzzards selected nests higher above ground and on trees frequently covered by ivy. At the landscape-scale, buzzards avoided roads and conspecifics, while selecting rugged areas with high habitat heterogeneity, probably related to a varied food supply. Productivity was related to the availability of arid habitats, probably because of their richness in main prey species. Finally, population density was negatively related to the abundance of eagle owls (Bubo bubo), a potential predator of adults and nestlings, and positively related to the availability of woodland, a low predation-risk habitat rich in food and nest-sites. Therefore, buzzard settlement, density and productivity depended on the complex interplay of food availability, human persecution and predation risk. Thus, the current landscape changes would benefit buzzards by providing more nest-sites, but would be detrimental because of the lower productivity associated with the disappearance of dry open areas. Proposed conservation guidelines focus on conversion of coppice woodland to mature forests and active management of dry heath, a conservation sensitive habitat, through controlled burning. [source]


What helps Opuntia stricta invade Kruger National Park, South Africa: Baboons or elephants?

APPLIED VEGETATION SCIENCE, Issue 2 2007
L.C. Foxcroft
Germishuizen & Meyer (2003) for plant species Abstract Question: Is Opuntia stricta more frequent, and its patches larger, under trees suitable for baboon roosting? If so, does it mean that baboons are major dispersal agents and that plants established under these trees are important foci of Opuntia stricta spread? Location: Skukuza, Kruger National Park, South Africa. Method: We surveyed an area invaded by Opuntia stricta in the Skukuza region of KNP. The survey included plots under potential baboon roosting trees,plots under trees unlikely to support baboons,and paired randomly located open sites. Results: The null hypothesis -tree- Opuntia spatial independence , can be rejected for Acacia nilotica, but not for Spirostachys africana. Opuntia plants are positively associated with Acacia trees suitable for baboon roosting. However, there is no significant difference between frequency of Opuntia under Acacia trees suitable and unsuitable for baboon roosting. It appears that all Acacia trees can serve as nurse trees for Opuntia. Compared to plots under Acacia trees, frequencies of old and robust Opuntia plants are significantly higher in open areas and under dead trees. Conclusions: While baboons may be responsible for long distance Opuntia dispersal (over km),their role is not detectable at a local scale. On the other hand, elephants seem to contribute substantially to the local vegetative propagation of this species. Opuntia establishment and growth are more influenced by micro-habitat than previously thought. [source]