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Middle Eocene (middle + eocene)
Selected AbstractsA new Eocene swift-like bird with a peculiar featheringIBIS, Issue 3 2003Gerald Mayr A new taxon of swift-like birds is described from the Middle Eocene of Messel (Germany). It is tentatively assigned to the extinct family Jungornithidae and exhibits a completely unexpected feathering, which contrasts sharply with that of recent swifts. The short and rounded wings clearly show that it was not adapted to gliding, but might have caught its prey by sallying flights from a perch. The tail of the new taxon is very long and the tail feathers are broad and nearly symmetrical. The phylogenetic relationships between the Jungornithidae and other apodiform birds are still not convincingly resolved. The early Oligocene genus Jungornis itself shares unique derived characters with hummingbirds which are, however, absent in the Eocene genus Argornis and in the new taxon from Messel. [source] Metamorphic and cooling history of the Shimanto accretionary complex, Kyushu, Southwest Japan: Implications for the timing of out-of-sequence thrustingISLAND ARC, Issue 4 2008Hidetoshi Hara Abstract Illite crystallinity, K,Ar dating of illite, and fission-track dating of zircon are analyzed in the hanging wall (Sampodake unit) and footwall (Mikado unit) of a seismogenic out-of-sequence thrust (Nobeoka thrust) within the Shimanto accretionary complex of central Kyushu, southwest Japan. The obtained metamorphic temperatures, and timing of metamorphism and cooling, reveal the tectono-metamorphic evolution of the complex, and related development of the Nobeoka thrust. Illite crystallinity data indicate that the Late Cretaceous Sampodake unit was metamorphosed at temperatures of around 300 to 310°C, while the Middle Eocene Mikado unit was metamorphosed at 260 to 300°C. Illite K,Ar ages and zircon fission-track ages constrain the timing of metamorphism of the Sampodake unit to the early Middle Eocene (46 to 50 Ma, mean = 48 Ma). Metamorphism of the Mikado unit occurred no earlier than 40 Ma, which is the youngest depositional age of the unit. The Nobeoka thrust is inferred to have been active during about 40 to 48 Ma, as the Sampodake unit started its post metamorphic cooling after 48 Ma and was thrust over the Mikado unit at about 40 Ma along the Nobeoka thrust. These results indicate that the Nobeoka thrust was active for more than 10 million years. [source] Geological age of the Yokawa Formation of the Kobe Group (Japan) on the basis of terrestrial mammalian fossilsISLAND ARC, Issue 3 2007Takehisa Tsubamoto Abstract The age of the Yokawa Formation of the Cenozoic Kobe Group distributed in Hyogo Prefecture of western Japan based on mammalian fossils is discussed. Two fossil dental specimens of terrestrial mammals discovered from the lowest part of the Yokawa Formation in the Sanda area are described. These two fossils described here are: (i) a right mandibular fragment with p2,m3 of Bothriodon sandaensis sp. nov. (selenodont anthracotheriid artiodactyl), which appears to be the most primitive among the species of the genus; and (ii) right m1,m3 of cf. Hyrachyus sp. (primitive rhinocerotoid perissodactyl). In the lower part of the Yokawa Formation, Zaisanamynodon (amynodontid perissodactyl) was previously reported. The morphology (,evolutionary stage') of B. sandaensis is indicative of the latest Middle to Late Eocene, that of cf. Hyrachyus sp. is indicative of the Early to Middle Eocene, and Zaisanamynodon is indicative of the Late Middle to Late Eocene. Therefore, the fossil mammals of the Yokawa Formation indicate an latest Middle Eocene (ca. 38 Ma) correlation for the lower part of the formation, as a working hypothesis. Although the resolution of the geological age based on these mammalian fossils is relatively low compared to that based on marine index fossils, this result is concordant with the recent radiometric correlation of the lower part of the Yokawa Formation in the Sanda area. [source] Hindlimb adaptations in Ourayia and Chipetaia, relatively large-bodied omomyine primates from the Middle Eocene of UtahAMERICAN JOURNAL OF PHYSICAL ANTHROPOLOGY, Issue 3 2006Rachel H. Dunn Abstract North American omomyids represent a tremendous Eocene radiation of primates exhibiting a wide range of body sizes and dietary patterns. Despite this adaptive diversity, relatively little is known of the postcranial specializations of the group. Here we describe hindlimb and foot bones of Ourayia uintensis and Chipetaia lamporea that were recovered from the Uinta B member (early Uintan Land Mammal Age), Uinta Formation, Utah. These specimens provide insights into the evolution of postcranial adaptations across different body sizes and dietary guilds within the Eocene primate radiation. Body mass estimates based on talar measurements indicate that Ourayia uintensis and Chipetaia lamporea weighed about 1,500,2,000 g and 500,700 g, respectively. Skeletal elements recovered for Ourayia include the talus, navicular, entocuneiform, first metatarsal, and proximal tibia; bones of Chipetaia include the talus, navicular, entocuneiform, and proximal femur. Both genera had opposable grasping big toes, as indicated by the saddle-shaped joint between the entocuneiform and first metatarsal. Both taxa were arboreal leapers, as indicated by a consistent assemblage of characters in all represented bones, most notably the somewhat elongated naviculars, the high and distinct trochlear crests of the talus, the posteriorly oriented tibial plateau (Ourayia), and the cylindrical head of the femur (Chipetaia). The closest resemblances to Ourayia and Chipetaia are found among the Bridger omomyines, Omomys and Hemiacodon. The results of our comparisons suggest that the later, larger, more herbivorous omomyines from Utah retained a skeletal structure characteristic of earlier, smaller North American omomyids. Am J Phys Anthropol, 2006. © 2006 Wiley-Liss, Inc. [source] Bird evolution in the Eocene: climate change in Europe and a Danish fossil faunaBIOLOGICAL REVIEWS, Issue 4 2006Bent E. K. Lindow ABSTRACT The pattern of the evolutionary radiation of modern birds (Neornithes) has been debated for more than 10 years. However, the early fossil record of birds from the Paleogene, in particular, the Lower Eocene, has only recently begun to be used in a phylogenetic context to address the dynamics of this major vertebrate radiation. The Cretaceous-Paleogene (K-P) extinction event dominates our understanding of early modern bird evolution, but climate change throughout the Eocene is known to have also played a major role. The Paleocene and Lower Eocene was a time of avian diversification as a result of favourable global climatic conditions. Deteriorations in climate beginning in the Middle Eocene appear to be responsible for the demise of previously widespread avian lineages like Lithornithiformes and Gastornithidae. Other groups, such as Galliformes display replacement of some lineages by others, probably related to adaptations to a drier climate. Finally, the combination of slowly deteriorating climatic conditions from the Middle Eocene onwards, appears to have slowed the evolutionary rate in Europe, as avian faunas did not differentiate markedly until the Oligocene. Taking biotic factors in tandem with the known Paleogene fossil record of Neornithes has recently begun to illuminate this evolutionary event. Well-preserved fossil taxa are required in combination with ever-improving phylogenetic hypotheses for the interrelationships of modern birds founded on morphological characters. One key avifauna of this age, synthesised for the first time herein, is the Lower Eocene Fur Formation of Denmark. The Fur birds represent some of the best preserved (often in three dimensions and with soft tissues) known fossil records for major clades of modern birds. Clear phylogenetic assessment of these fossils will prove critical for future calibration of the neornithine evolutionary timescale. Some early diverging clades were clearly present in the Paleocene as evidenced directly by new fossil material alongside the phylogenetically constrained Lower Eocene taxa. A later Oligocene radiation of clades other than Passeriformes is not supported by available fossil data. [source] CRETACEOUS CARBONATES IN THE ADIYAMAN REGION, SE TURKEY: AN ASSESSMENT OF BURIAL HISTORY AND SOURCE-ROCK POTENTIALJOURNAL OF PETROLEUM GEOLOGY, Issue 1 2000I. H. Demirel The burial history and source-rock potential of Cretaceous carbonates in the Adiyaman region of SE Turkey have been investigated. The carbonates belong to the Aptian-Campanian Mardin Group and the overlying Karabogaz Formation. The stratigraphy of these carbonates at four well locations was recorded. At each well, the carbonate succession was found to be incomplete, and important unconformities were present indicating periods of non-deposition and/or erosion. These unconformities are of variable extent. When combined with the effects of rapid subsidence and sedimentation which took place in the SW of the Adiyaman region during end-Cretaceous foredeep development, they have resulted in variations in the carbonates' present-day burial depths, thereby influencing the regional pattern of source-rock maturation and the timing of oil generation. Burial history curves indicate that the carbonates' maturity increases from SW to NE, towards the Late Cretaceous thrust belt. Predicted levels of maturity for the Mardin Group are consistent with measured geochemical data from three of the wells in the study area (the exception being well Karadag-1). Three potential source-rock intervals of Cretaceous age have been identified. Two of these units , the Derdere and Karababa Formations of the Mardin Group , are composed of shallow-water carbonates which were deposited on the northern margin of the Arabian Platform. The third source-rock unit, the overlying Karabogaz Formation, is composed of pelagic carbonates which were deposited during a regional transgression. These potential source-rock intervals contain marine organic matter dominated by Type II kerogen. Total organic carbon contents range from 0.5 to 2.9 %. Time-temperature analyses indicate that the Mardin Group carbonates are immature to marginally mature at well locations in the SW of the study area, and are mature at western and NE well locations. The onset of oil generation in these Cretaceous source rocks took place between the middle Eocene (48 million yrs ago) and the Oligocene (28 million yrs ago). [source] A new genus of rodents (Remyidae, Mammalia) from the Iberian EocenePALAEONTOLOGY, Issue 1 2010RAMÓN QUER Abstract:, In this article, a new genus, Frontanyamys, is defined based on the new species F. russelli. The genus is recorded from the lower Upper Eocene (Bartonian) beds AT Sant Jaume de Frontanya (NE Spain). This genus shows clear affinities with the previously described genera Zamoramys, Remys and Pairomys. They are therefore assembled in the family Remyidae (new rank). The remyids are characterised by the precocious development of high-crowned molars and retain a morphologically primitive dental pattern. The oldest remyids are found in the middle Eocene of Spain (Zamoramys) and are probably derived from a morphologically primitive protrogomorph rodent such as Corbarimys paisi. The last representatives of the family such as Remys and Pairomys developed a fully lophodont dental pattern. [source] The phylogenetic affinities of the Pondaung taliAMERICAN JOURNAL OF PHYSICAL ANTHROPOLOGY, Issue 2 2010Marian Dagosto Abstract The phylogenetic affinities of the primates of the late middle Eocene Pondaung Formation of Myanmar have long been disputed. The discovery of the NMMP 39 talus (Marivaux et al.: Proc Natl Acad Sci USA 100 (2003) 13173,13178) provided the first clear evidence from the postcranium that a relatively large-bodied haplorhine primate is represented in the Pondaung fauna. Another talus (NMMP 82; Marivaux et al., 2010). Talar morphology, phylogenetic affinities and locomotor adaptation of a large-bodied amphipithecid primate from the late middle Eocene of Myanmar, Am J Phys Anthropol DOI: 10.1002/ajpa.21307) has been recently recovered which also pertains to Haplorhini. The metric and nonmetric features supporting the hypothesis of anthropoid affinities for NMMP 39 have been criticized by Gunnell and Ciochon (Gunnell GF, Ciochon RL. 2008. Revisiting primate postcrania from the Pondaung Formation of Myanmar. In: Fleagle JG, Gilbert CC, editors. Elwyn Simons: a search for origins. New York: Springer. p 211,228). Their analysis, however, was based on a very limited choice of variables, taxa, and individuals. Based on an extended sample, we are able to produce both principal components and discriminant functions that yield a rather clear separation of extant haplorhine and strepsirhine tali. Both principal components and discriminant function scores of the Pondaung tali fall with those of haplorhine primates. In addition, the Pondaung tali lack all the derived nonmetric features characteristic of strepsirhine primates, but exhibit all the features characteristic of haplorhine primates. We dispute the features Gunnell and Ciochon (2008) claim are uniquely shared by the Pondaung tali and adapiforms. Their rejection of the phylogenetic significance of the features shared by these tali and haplorhines is unwarranted by the evidence. Based on both metric and nonmetric features, the Pondaung tali are structurally most similar to the tali of haplorhines, particularly anthropoids. Am J Phys Anthropol 143:223,234, 2010. © 2010 Wiley-Liss, Inc. [source] A new species of Pseudoloris (Omomyidae, Primates) from the middle Eocene of Sant Jaume de Frontanyŕ (Eastern Pyrenees, Spain)AMERICAN JOURNAL OF PHYSICAL ANTHROPOLOGY, Issue 1 2010Raef Minwer-Barakat Abstract In this article we describe a new species of Pseudoloris (Omomyidae, Primates) from the Robiacian (middle Eocene) locality of Sant Jaume de Frontanyŕ (Eastern Pyrenees, Spain). Pseudoloris pyrenaicus is characterized by its medium size, thickened paracristid, absence of a distinct paraconid, and well-developed buccal cingulid in the lower molars, large hypoconulid in the M3, paraconule and hypocone reduced in the M1 and M2 and absent in the M3. The material from Sant Jaume de Frontanyŕ constitutes the most abundant sample of the genus Pseudoloris found until now in the Iberian Peninsula. Almost all the dental elements have been recovered, including those teeth hardly known for other species of the genus, such as lower and upper incisors. The new species shows intermediate features between Pseudoloris isabenae from Capella and Pseudoloris parvulus, present in different Spanish and French sites. Therefore, we consider that Pseudoloris pyrenaicus is an intermediate form between these two species. Am J Phys Anthropol 143:92,99, 2010. © 2010 Wiley-Liss, Inc. [source] Talar morphology, phylogenetic affinities, and locomotor adaptation of a large-bodied amphipithecid primate from the late middle eocene of MyanmarAMERICAN JOURNAL OF PHYSICAL ANTHROPOLOGY, Issue 2 2010Laurent Marivaux Abstract A well-preserved fossil talus [National Museum of Myanmar Primates (NMMP) 82] of a large-bodied primate is described from the late middle Eocene Pondaung Formation of central Myanmar. The specimen was collected at Thandaung Kyitchaung, a well-known amphipithecid primate,bearing locality near the village of Mogaung. NMMP 82 adds to a meager but growing sample of postcranial remains documenting the large-bodied primates of the Pondaung Formation. This new talus exhibits a suite of features that resemble conditions found in living and fossil haplorhine primates, notably anthropoids. As such, the phylogenetic signal deriving from the morphology of NMMP 82 conflicts with that provided by NMMP 20, a partial skeleton (including a fragmentary calcaneus) of a second large-bodied Pondaung primate showing undoubted adapiform affinities. Analysis subtalar joint compatibility in a hypothetical NMMP 82/NMMP 20 combination (talus/calcaneus) reveals a substantial degree of functional mismatch between these two tarsal bones. The functional incongruence in subtalar joint morphology between NMMP 20 and NMMP 82 is consistent with the seemingly divergent phylogenetic affinities of these specimens, indicating that two higher level taxa of relatively large-bodied primates are documented in the Pondaung Formation. On the basis of its size and morphology, we refer the NMMP 82 talus to the large-bodied amphipithecid Pondaungia. The occurrence of anthropoid-like tali in the Pondaung Formation obviates the need to invoke homoplasy to explain the shared, derived dental characters that are common to amphipithecids and undoubted anthropoids. Functionally, the NMMP 82 talus appears to have pertained to a primate that is engaged in active quadrupedalism in an arboreal environment along broad and subhorizontal branches. The primate taxon represented by NMMP 82 was capable of climbing and leaping, although it was not particularly specialized for either of these activities. Am J Phys Anthropol 143:208,222, 2010. © 2010 Wiley-Liss, Inc. [source] | |||||||||||||