Mediterranean Region (mediterranean + region)

Distribution by Scientific Domains
Distribution within Life Sciences

Kinds of Mediterranean Region

  • eastern mediterranean region


  • Selected Abstracts


    Dating methods for sediments of caves and rockshelters with examples from the Mediterranean Region

    GEOARCHAEOLOGY: AN INTERNATIONAL JOURNAL, Issue 4 2001
    H. P. Schwarcz
    A wide range of potential dating methods may be applied to archaeological deposits found in caves and rockshelters, depending on the nature of the deposit and age range of the deposit. Organic sediments, including faunal and floral material, can be dated by radiocarbon (AMS and high-sensitivity beta-counting). Many karstic features contain speleothems which can be dated with high accuracy by U-series. Wind-blown detritus, where it is the dominant constituent of the cave deposits, can be dated by luminescence (TL, OSL, or IRSL), taking care to avoid material fallen into the deposits from the shelter/cave walls. Fireplaces contain burned rocks (including stone artifacts) which can be dated by TL. Enamel from the teeth of mammals is present in most sites, representing either animal residents of the shelter, or residues from food brought to the shelter by human residents. Electron spin resonance (ESR) dating of enamel is applicable over a wide time range, with high accuracy and reasonable precision where uranium accumulation in teeth is low, but with lower precision where uranium content in teeth is high. In general, multiple dating methods applied to a site may resolve ambiguities arising from uncertain model assumptions in some dating methods. © 2001 John Wiley & Sons, Inc. [source]


    The Convergence of Civilizations: Constructing a Mediterranean Region , Edited by E. Adler, F. Bicchi, B. Crawford and R. Del Sarto

    JCMS: JOURNAL OF COMMON MARKET STUDIES, Issue 2 2007
    SHARON PARDO
    No abstract is available for this article. [source]


    Effect of Planting Date on Seed Yield, Oil Content and Fatty Acid Composition of Safflower (Carthamus tinctorius) Cultivars Grown in the Mediterranean Region of Turkey

    JOURNAL OF AGRONOMY AND CROP SCIENCE, Issue 5 2003
    B. Samanc
    Abstract A study was conducted to investigate the effect of different planting dates (25 April, 5 May and 15 May 1998 and 30 April, 15 May and 25 May 1999) on the seed yield, oil content and fatty acid composition of three safflower (Carthamus tinctorius) cultivars (Yenice 5-38, Dincer 5-118 and 5-154) grown in fields of the research facility of Akdeniz University in Antalya, Turkey. While seed yield, oil content, and palmitic acid, stearic acid, and oleic acid contents decreased, linoleic acid content increased from 50.86 to 55.72 % with delay in planting date. The effect of genotype on fatty acids was greater than that of environment. [source]


    Climatic stress, food availability and human activity as determinants of endemism patterns in the Mediterranean region: the case of dung beetles (Coleoptera, Scarabaeoidea) in the Iberian Peninsula

    DIVERSITY AND DISTRIBUTIONS, Issue 5 2002
    José R. Verdú
    Abstract. A study to assess the influence of abiotic (climatic conditions) and biotic factors (food resources, habitat preference and human activity) on endemism patterns of dung beetles in the Mediterranean region was conducted in the Iberian Peninsula and the Balearic Islands. The Thermicity Index (It), the Mediterraneity Index (Im3) and the Aridity Index (Ia) were used to assess the influence of abiotic factors. Relative rabbit density (DR), the proportion of landscape used historically for grazing by sheep and goats and the nature of the food resource were used to assess the influence of biotic factors. Relative endemism (EN) of dung beetle assemblages was positively and significantly related with all of the factors considered. However, the Aridity and Mediterraneity Indices are the best predictors of EN. The predicted endemism (EN = 0.017 Ia + 0.004 Im3 + 0.422) was highly positively and significantly related with the observed endemism. Dung beetle assemblages with the highest relative endemism were observed in the south-eastern part of the Iberian Peninsula. This distribution corresponded to the highest Aridity and Mediterraneity. In contrast, dung beetle assemblages with lower endemism were located in more humid and temperate areas. Assemblages of dung beetles with the highest endemism comprise many species adapted to aridity and the exploitation of dry dung pellets. Conservation of traditional grazing activity by pellet-dropping sheep and goats might benefit the maintenance of dung beetle biodiversity in Mediterranean ecosystems. [source]


    Tafoni development in a cryotic environment: an example from Northern Victoria Land, Antarctica

    EARTH SURFACE PROCESSES AND LANDFORMS, Issue 10 2008
    Andrea Strini
    Abstract Tafoni are a type of cavernous weathering widespread around the world. Despite the extensive distribution of the tafoni, their genesis is not clear and is still a matter of debate, also because they occur in such different climatic conditions and on so many different types of substrate. Geomorphological characterization of more than 60 tafoni in three different Antarctic sites (two coastal and one inland) between 74 and 76° S with sampling of weathering products and salt occurrences are described together with thermal data (on different surfaces) and wind speed recorded in different periods of the year in a selected tafone close to the Italian Antarctic station. The aim of this present study is to provide further information to help understand the processes involved in the growth of tafoni in a cryotic environment, and the relationship of these processes to climate, with particular attention to the thermal regime and the role of wind. The new data presented in this paper suggest that there is no single key factor that drives the tafoni development, although thermal stress seems the most efficient process, particularly if we consider the short-term fluctuations. The data also confirm that other thermal processes, such as freezing,thawing cycles and thermal shock, are not really effective for the development of tafoni in this area. The wind speed measured within the tafoni is half that recorded outside, thus favouring snow accumulation within the tafoni and therefore promoting salt crystallization. On the other hand, the wind effect on the thermal regime within the tafoni seems negligible. While both salt weathering and thermal stress appear active in this cryotic environment, these are azonal processes and are therefore active in other climatic areas where tafoni are widespread (such as the Mediterranean region). Copyright © 2007 John Wiley & Sons, Ltd. [source]


    Use of terrain variables for mapping gully erosion susceptibility in Lebanon

    EARTH SURFACE PROCESSES AND LANDFORMS, Issue 12 2007
    Rania Bou Kheir
    Abstract This paper predicts the geographic distribution and size of gullies across central Lebanon using a geographic information system (GIS) and terrain analysis. Eleven primary (elevation; upslope contributing area; aspect; slope; plan, profile and tangential curvature; flow direction; flow width; flow path length; rate of change of specific catchment area along the direction of flow) and three secondary (steady-state; quasi-dynamic topographic wetness; sediment transport capacity) topographic variables were generated and used along with digital data collected from other sources (soil, geology) to statistically explain gully erosion field measurements. Three tree-based regression models were developed using (1) all variables, (2) primary topographic variables only and (3) different pairs of variables. The best regression tree model combined the steady-state topographic wetness and sediment transport capacity indices and explained 80% of the variability in field gully measurements. This model proved to be simple, quick, realistic and practical, and it can be applied to other areas of the Mediterranean region with similar environmental conditions, thereby providing a tool to help with the implementation of plans for soil conservation and sustainable management. Copyright © 2007 John Wiley & Sons, Ltd. [source]


    Mismatch between the timing of oviposition and the seasonal optimum.

    ECOLOGICAL ENTOMOLOGY, Issue 3 2010
    The stochastic phenology of Mediterranean acorn weevils
    1. The timing of reproduction is predicted to match the period of maximum food availability. In this sense, the case of many phytophagous insects in temperate regions is very illustrative, as their larvae usually depend on a resource only available for a limited period of time each year. 2. For 3 years the interactions between the weevil Curculio elephas and the Mediterranean Holm oak Quercus ilex were studied. Weevil larvae grow within the acorns, feeding on the cotyledons. The timing of oviposition will determine food availability for the larvae, as acorns stop growing once they are attacked. 3. Acorn temporal growing patterns did not change between years and food availability for larvae was at its highest in October, when temperature was still suitable for larval development. However, oviposition phenology did change between years. In 2002 females oviposited later, larvae grew within larger acorns, and their body mass was significantly higher than in 2003 or 2004, when females oviposited into early acorns. 4. Thus, weevils do not always adjust oviposition to the best possible feeding conditions for their offspring. Rather, they seem to maximise their own lifetime fitness, ovipositing as soon as they emerge in late summer. Emergence, in turn, depends strongly on stochastic events such as summer storms in the Mediterranean region. 5. Under a climate change perspective, the trend towards higher August rainfall recorded in our study area may alter oviposition phenology, with the subsequent cascade effects on weevil body size and fitness [source]


    Environmental benchmarking for management of energy and water use: A study of SMEs in the Mediterranean region

    ENVIRONMENTAL QUALITY MANAGEMENT, Issue 3 2008
    Aikaterini Makrinou
    First page of article [source]


    Rudabánya: A late miocene subtropical swamp deposit with evidence of the origin of the African apes and humans

    EVOLUTIONARY ANTHROPOLOGY, Issue 2 2002
    László Kordos
    Abstract Rudabánya, a rich late Miocene fossil site in northern central Hungary, has yielded an abundant record of fossil primates, including the primitive catarrhine Anapithecus and the early great ape Dryopithecus. While the affinities of Anapithecus are not clear, Dryopithecus is clearly a great ape sharing numerous characteristics of its dental, cranial and postcranial anatomy with living great apes. Like all Miocene hominids (great apes and humans), Dryopithecus is more primitive in a number of ways than any living hominid, which is probably related to the passage of time since the divergence of the various lineages of living hominids, allowing for similar refinements in morphology and adaptation to take place independently. On the other hand, Dryopithecus (and Ouranopithecus) share derived characters with hominines (African apes and humans), and Sivapithecus (and Ankarapithecus) share derived characters with orangutans, thus dating the split between pongines and hominines to a time before the evolution of these fossil great apes. Pongines and hominines follow similar fates in the late Miocene, the pongines moving south into Southeast Asia from southern or eastern Asia and the hominines moving south into East Africa from the Mediterranean region, between 6 to 9 Ma. [source]


    Micromorphological studies of Greek prehistoric sites: New insights in the interpretation of the archaeological record

    GEOARCHAEOLOGY: AN INTERNATIONAL JOURNAL, Issue 3 2002
    Panagiotis Karkanas
    Ongoing micromorphological studies in several prehistoric sites of Greece (Theopetra cave, Boila rockshelter, Alonnisos, Drakaina cave, Kouveleiki cave, Lakonis cave complex, and Dispilio lake dwelling) provide new information on the relationship between environmental changes and the cultural history of the sites. The frequent climatic oscillations during the last glacial directly influenced the occupational mode of Theopetra cave and Boila rockshelter in northern Greece. Soil micromorphology may be a promising tool in unraveling differences in the occupational history due to climatic changes among diverse areas of Greece. Some preliminary observations from the Lakonis cave complex, in southern coastal Greece, support the existence of such differences. Evidence brought forward with the micromorphological study of Dispilio lake dwelling and Theopetra cave suggests that during the Holocene, aridification phases evident in the Mediterranean region might have also played a role in the Greek prehistoric settlement pattern. So far, evidence for aridification phases is present for the end of the Mesolithic and probably for part of the Final Neolithic. In several cases, micromorphology has revealed details of the cultural nature of the sites. Questions related to occupational intensity (Theopetra, Lakonis, Kouveleiki, and Drakaina caves), post-depositional changes and cultural modification of the sediments (Alonnisos, Theopetra, and Drakaina caves), constructions (Theopetra and Drakaina caves), and stratigraphic correlation (Boila) have been satisfactorily addressed along with the analysis of the microstructure of the sediments. © 2002 Wiley Periodicals, Inc. [source]


    The sedimentary records in Mediterranean rockshelters and caves: Archives of environmental change

    GEOARCHAEOLOGY: AN INTERNATIONAL JOURNAL, Issue 4 2001
    Jamie C. Woodward
    It is important to develop rigorous methods and robust conceptual models for the interpretation of rockshelter and cave sediment records so that the cultural sequences they contain can be considered in their proper environmental context. Much of what we know about the prehistory of the Mediterranean region and adjacent areas has largely been pieced together from materials excavated from sedimentary sequences in these environments. The rockshelters and caves of the region form important environmental and sedimentary archives. Recent work has begun to consider if the remarkable climatic variability evident in the high resolution lacustrine and ice core records is manifest in the rockshelter and cave sediment records of the area. In this context, the two main characteristics of a rockshelter or cave site which control its usefulness as an archive of environmental change are the temporal resolution of the sedimentary record and the environmental sensitivity of the site. Many rockshelters and caves can be described as either Active Karst Settings (AKS) or Passive Karst Settings (PKS) and site type is an important influence on climatic sensitivity with a direct influence upon the usefulness of the sedimentary sequence as a proxy record of climate change. It is now clear that some sites may preserve detailed paleoclimatic records and the climatic signal may be represented by distinctive suites of micromorphological features, by variations in the input of allogenic sediment, or by fluctuations in the mineral magnetic properties of the fine sediment fraction. It can be argued that data derived from the analysis of bulk coarse-grained samples often lacks the stratigraphic resolution and environmental sensitivity that can be obtained from other approaches. The most favorable sites for detailed paleoclimatic reconstruction appear to be in active karst settings such as Theopetra Cave (Greece) and Pigeon Cave (Morocco) where micromorphological analyses offer insights into the stratigraphic record that are not otherwise obtainable. The temporal resolution of a site can only be established through a rigorous stratigraphic analysis and a comprehensive dating program. These are fundamental considerations in the study of rockshelter sediment records, especially when attempting to correlate between sites and draw comparisons with other proxy records of environmental change derived from sedimentary environments with rather different characteristics. Rockshelters and caves are part of a wider sediment system, and their investigation must be accompanied by detailed geomorphological, sedimentological, paleoecological, and geochronological studies of the off-site Quaternary record. © 2001 John Wiley & Sons, Inc. [source]


    Developments in research concerning Mesozoic,Tertiary Tethys and neotectonics in the Isparta Angle, SW Turkey

    GEOLOGICAL JOURNAL, Issue 3-4 2003
    Alastair H. F. Robertson
    Abstract The Isparta Angle has played a critical role in the development of concepts concerning the tectonic evolution of the Mesozoic,Tertiary Neotethys in the Eastern Mediterranean region. Following early regional studies, mainly by the Mineral Research and Exploration Institute of Turkey (MTA), during the 1960s and 1970s, a French team mapped the area and confirmed a regional tectonostratigraphy of three great allochthonous systems of mainly Mesozoic,Early Tertiary age, termed the Antalya, Lycian and Hoyran-Bey,ehir-Had,m nappes. During the 1970s and 1980s a British group studied the Neotethyan evolution of what they termed the Antalya Complex, utilizing knowledge of plate tectonic processes. Throughout the 1970s and 1980s MTA systematically remapped the area at 1:25,000 scale. The root zone of the Antalya allochthon was either a southerly Neotethys, within and to the south of the Isparta Angle, or a northerly Neotethys, many hundreds of kilometres to the north. The southerly origin is nowadays favoured but some questions remain. Attention focused in the 1990s until present to the post-collisional, neotectonic evolution of the Isparta Angle and its links with the neighbouring Mediterranean Sea. Here, we trace the development of research and the ongoing debates concerning alternative tectonic concepts used to explain the evolution of the Isparta Angle from Mesozoic to Recent time. We conclude by outlining several tectonic models for the evolution of the Antalya allochthon within a southerly Neotethys that require to be tested by future field studies. Copyright © 2003 John Wiley & Sons, Ltd. [source]


    Seismotectonics of the Sinai subplate , the eastern Mediterranean region

    GEOPHYSICAL JOURNAL INTERNATIONAL, Issue 1 2003
    Amos Salamon
    SUMMARY We define the Sinai subplate, from a seismotectonic perspective, as a distinct component in the plate tectonics of the eastern Mediterranean region. This is based on the tectonic characteristics of a comprehensive listing of all ML, 4 recorded seismicity in the region during the 20th century, on newly calculated and recalculated fault plane mechanisms of first P -wave arrivals and on published solutions based on waveform inversion of broad-band data. The low seismicity level and scarcity of strong events in the region required a thorough search for useful data and a careful examination of the reliability of the focal solutions. We gathered all available records of first P -wave onsets from the ISS and ISC Bulletins and the local seismic networks. Altogether, we were able to calculate 48 new focal mechanisms and 33 recalculated ones of events that occurred during the years 1940,1992. With the increasing number of teleseismic and regional broad-band stations in the later years, we added 37 solutions based on teleseismic and regional waveform inversions of events that occurred during 1977,2001. These mechanisms enabled us to examine the seismotectonic character of the Sinai subplate. The strike and rake directions of the calculated mechanisms usually reflect the geometry and the large-scale type of deformation observed along the boundaries of the Sinai subplate,the Dead Sea Transform, the Cypriot Arc convergent zone and the Suez Rift. Nevertheless, along each of these boundaries we found anomalous solutions that attest to the complexity of the deformation processes along plate margins. Earthquakes along the Dead Sea Transform exhibit mainly sinistral transtension and transpression, reflecting its leaky manner and local change in the transform geometry. The presence of other unexpected mechanisms near the transform, however, reflects the heterogeneous deformation it induces around. As expected, thrust mechanisms along the Cypriot Arc mirror its convergent nature and typical curved geometry. Transtension and transpressional solutions in the eastern segment of the arc reflect the sinistral shear motion between Anatolia and Sinai there. However, shear mechanisms found between Cyprus and the Eratosthenes Seamount pose a problem regarding its collision process. Most intriguing of all are ML, 4 thrust and shear solutions found in the Gulf of Suez. They are associated with predominantly normal mechanisms within a rift zone and therefore constitute a unique phenomenon, yet to be deciphered. [source]


    Crustal versus asthenospheric relaxation and post-seismic deformation for shallow normal faulting earthquakes:the Umbria,Marche (central Italy) case

    GEOPHYSICAL JOURNAL INTERNATIONAL, Issue 3 2000
    R. Riva
    Summary Following a normal mode approach for a stratified viscoelastic earth, we investigate the effects induced by shallow normal faulting earthquakes, on surface post-seismic vertical displacement and velocity at the surface, when stress relaxation occurs in the crust or in the asthenosphere. The modelled earthquake is a moderate one characteristic of some slowly deforming plate boundaries in the central Mediterranean region. We focus on the Umbria,Marche (central Italy) region where deep seismic reflection studies (CROP03) and the 1997 earthquake sequence clearly show a seismogenic layer decoupled from the lower crust by a sizeable transition zone. Accordingly, the crust is subdivided into three layers: an elastic upper crust, a transition zone and a low-viscosity lower crust. The fault is embedded in the upper crust. The layered viscoelastic structure of the crust and mantle imposes a pattern and scale on the modelled coseismic and post-seismic deformation with a major contribution from the transition crustal zone and low-viscosity lower crust, stress relaxation in the mantle being negligible. We show that significant vertical deformation rates of the order of 1 mm yr, 1 could be expected for a shallow and moderate event such as the recent Umbria,Marche earthquake for viscosity values of 1019 and 1018 Pa s in the crustal transition zone and lower crust, respectively. [source]


    Mass mortality in Northwestern Mediterranean rocky benthic communities: effects of the 2003 heat wave

    GLOBAL CHANGE BIOLOGY, Issue 5 2009
    J. GARRABOU
    Abstract Late in summer 2003, extensive mass mortality of at least 25 rocky benthic macro-invertebrate species (mainly gorgonians and sponges) was observed in the entire Northwestern (NW) Mediterranean region, affecting several thousand kilometers of coastline. We were able to characterize the mortality event by studying six areas covering the main regions of the NW Mediterranean basin. The degree of impact on each study area was quantified at 49 sites by estimating the proportion of colonies affected in populations of several gorgonian species compared with reference data obtained in years without mortality signs. According to these data, the western areas (Catalan coast and Balearic Islands) were the least affected, while the central areas (Provence coast and Corsica-Sardinia) showed a moderate impact. The northernmost and eastern areas (Gulf of Genoa and Gulf of Naples) displayed the highest impact, with almost 80% of gorgonian colonies affected. The heat wave of 2003 in Europe caused an anomalous warming of seawater, which reached the highest temperatures ever recorded in the studied regions, between 1 and 3 °C above the climatic values (mean and maximum). Because this exceptional warming was observed in the depth ranges most affected by the mortality, it seems likely that the 2003 anomalous temperature played a key role in the observed mortality event. A correlation analysis between temperature conditions and degree of impact seems to support this hypothesis. Under the present climate warming trend, new mass mortality events may occur in the near future, possibly driving a major biodiversity crisis in the Mediterranean Sea. [source]


    Rapid climate change-related growth decline at the southern range edge of Fagus sylvatica

    GLOBAL CHANGE BIOLOGY, Issue 11 2006
    ALISTAIR S. JUMP
    Abstract Studies on Fagus sylvatica show that growth in populations toward the southern limit of this species' distribution is limited strongly by drought. Warming temperatures in the Mediterranean region are expected to exacerbate drought where they are not accompanied by increases in precipitation. We studied levels of annual growth in mature F. sylvatica trees over the last half-century in the Montseny Mountains in Catalonia (northeast Spain). Our results show significantly lower growth of mature trees at the lower limit of this species' distribution when compared with trees at higher altitudes. Growth at the lower Fagus limit is characterized by a rapid recent decline starting in approximately 1975. By 2003, growth of mature trees had fallen by 49% when compared with predecline levels. This is not an age-related phenomenon, nor is it seen in comparable populations at higher altitudes. Analysis of climate-growth relationships suggests that the observed decline in growth is a result of warming temperatures and that, as precipitation in the region has not increased, precipitation is now insufficient to ameliorate the negative effects of increased temperatures on tree growth. As the climate-response of the studied forest is comparable with that of F. sylvatica forests in other southern European regions, it is possible that this growth decline is a more widespread phenomenon. Warming temperatures may lead to a rapid decline in the growth of range-edge populations and a consequent retreat of the species distribution in southern Europe. Assessment of long-term growth trends across the southern range edge of F. sylvatica therefore merits further attention. [source]


    Comparing and evaluating process-based ecosystem model predictions of carbon and water fluxes in major European forest biomes

    GLOBAL CHANGE BIOLOGY, Issue 12 2005
    Pablo Morales
    Abstract Process-based models can be classified into: (a) terrestrial biogeochemical models (TBMs), which simulate fluxes of carbon, water and nitrogen coupled within terrestrial ecosystems, and (b) dynamic global vegetation models (DGVMs), which further couple these processes interactively with changes in slow ecosystem processes depending on resource competition, establishment, growth and mortality of different vegetation types. In this study, four models , RHESSys, GOTILWA+, LPJ-GUESS and ORCHIDEE , representing both modelling approaches were compared and evaluated against benchmarks provided by eddy-covariance measurements of carbon and water fluxes at 15 forest sites within the EUROFLUX project. Overall, model-measurement agreement varied greatly among sites. Both modelling approaches have somewhat different strengths, but there was no model among those tested that universally performed well on the two variables evaluated. Small biases and errors suggest that ORCHIDEE and GOTILWA+ performed better in simulating carbon fluxes while LPJ-GUESS and RHESSys did a better job in simulating water fluxes. In general, the models can be considered as useful tools for studies of climate change impacts on carbon and water cycling in forests. However, the various sources of variation among models simulations and between models simulations and observed data described in this study place some constraints on the results and to some extent reduce their reliability. For example, at most sites in the Mediterranean region all models generally performed poorly most likely because of problems in the representation of water stress effects on both carbon uptake by photosynthesis and carbon release by heterotrophic respiration (Rh). The use of flux data as a means of assessing key processes in models of this type is an important approach to improving model performance. Our results show that the models have value but that further model development is necessary with regard to the representation of the some of the key ecosystem processes. [source]


    Canopy recovery after drought dieback in holm-oak Mediterranean forests of Catalonia (NE Spain)

    GLOBAL CHANGE BIOLOGY, Issue 12 2004
    Francisco Lloret
    Abstract Climate change is likely to produce more frequent and longer droughts in the Mediterranean region, like that of 1994, which produced important changes in the Quercus ilex forests, with up to 76% of the trees showing complete canopy dieback. At the landscape level, a mosaic of responses to the drought was observed, linked to the distribution of lithological substrates. Damage to the dominant tree species (Q. ilex) and the most common understorey shrub (Erica arborea) was more noticeable on the compact substrates (breccia) than on the fissured ones (schist). This result was consistent with observations documenting deeper root penetration in schist than in breccia materials, allowing the plants growing on fissured substrates to use water from deeper soil levels. Smaller plants were more vulnerable to drought than larger plants in the trees, but not in the shrubs. Overall, Q. ilex was more affected than E. arborea. The resilience of the system was evaluated from the canopy recovery 1 year after the episode. Stump and crown resprouting was fairly extensive, but the damage pattern in relation to substrate, plant size, and species remained similar. The effect of recurrent drought episodes was studied on vegetation patches of Q. ilex located on mountain slopes and surrounded by bare rock. We observed that plants that resprouted weakly after a previous drought in 1985 were more likely to die or to produce poor regeneration in 1995 than plants that had resprouted vigorously. Vegetation patches located on the lower part of the slope were also less damaged than patches situated uphill. The study provides evidence of relevant changes in forest canopy as a consequence of extreme climate events. The distribution of this effect across the landscape is mediated by lithological substrate, causing patchy patterns. The results also support the hypothesis that recurrent droughts can produce a progressive loss of resilience, by depleting the ability of surviving plants to regenerate. [source]


    Changed plant and animal life cycles from 1952 to 2000 in the Mediterranean region

    GLOBAL CHANGE BIOLOGY, Issue 6 2002
    Josep Peńuelas
    Abstract The available data on climate over the past century indicate that the earth is warming. Important biological effects, including changes of plant and animal life cycle events, have already been reported. However, evidence of such effects is still scarce and has been mostly limited to northern latitudes. Here we provide the first long-term (1952,2000) evidence of altered life cycles for some of the most abundant Mediterranean plants and birds, and one butterfly species. Average annual temperatures in the study area (Cardedeu, NE Spain) have increased by 1.4 °C over the observation period while precipitation remained unchanged. A conservative linear treatment of the data shows that leaves unfold on average 16 days earlier, leaves fall on average 13 days later, and plants flower on average 6 days earlier than in 1952. Fruiting occurs on average 9 days earlier than in 1974. Butterflies appear 11 days earlier, but spring migratory birds arrive 15 days later than in 1952. The stronger changes both in temperature and in phenophases timing occurred in the last 25 years. There are no significant relationships among changes in phenophases and the average date for each phenophase and species. There are not either significant differences among species with different Raunkiaer life-forms or different origin (native, exotic or agricultural). However, there is a wide range of phenological alterations among the different species, which may alter their competitive ability, and thus, their ecology and conservation, and the structure and functioning of ecosystems. Moreover, the lengthening of plant growing season in this and other northern hemisphere regions may contribute to a global increase in biospheric activity. [source]


    Climate change and grasslands through the ages: an overview

    GRASS & FORAGE SCIENCE, Issue 2 2007
    L. 't Mannetje
    Summary Change from cool to warm temperatures and vice versa have occurred throughout geological time. During the Jurassic and Cretaceous periods (206,65 million years ago, Ma) the climate was more uniformly warm and moist than at present and tropical rainforests were widespread. Grasses evolved during the Jurassic period and they expanded greatly as the climate differentiated with reduced rainfall and temperatures. C4 -grasses probably arose during the Oligocene period (24,35 Ma). During the Miocene period (23·8,5·3 Ma) grasslands expanded into huge areas (e.g. prairies in the USA, steppe in Eurasia, and pampas and llanos in South America). During the Quaternary period (1·8 Ma till now) some twenty-two different ice ages with periodicities of about 100 000 years occurred. Eighteen-thousand years ago, north-western Europe had a polar climate with tundra vegetation and the Mediterranean region was covered by steppe. During that time Amazonia was so dry that it was covered in extensive areas of savanna and the Sahara expanded rapidly. Only in the last 10 000 years has a closed rainforest covered the Amazonian region again. However, 9000 years ago a brief period of global warming caused excessive rains, which caused the sea and river levels to rise in north-western Europe with tremendous loss of life. The present period of extreme dryness in the Sahara only started some 5000 years ago and then the desert expanded rapidly into the Sahel. Before that the Sahara was covered by steppe. Global warming took place between about ad 900 and about ad 1200 or 1300 just before the Little Ice Age (1550,1700 ad). The article concludes with a description of temperature and vegetation changes that are occurring in Europe at present. It is predicted that C4 -grasses, which are already present in southern Europe, will further expand but that, in the short term, land abandonment will have much more deleterious effects than temperature change due to increased wild fires, loss of biodiversity and desertification. [source]


    Interdialytic blood pressure obtained by ambulatory blood pressure measurement and left ventricular structure in hypertensive hemodialysis patients

    HEMODIALYSIS INTERNATIONAL, Issue 3 2008
    Siddig MOMINADAM
    Abstract Unlike in subjects with normal renal function, the relationship between hypertension and cardiovascular morbidity and mortality in dialysis patients is still being debated. In order to clarify this issue, we performed 44-hour ambulatory blood pressure measurements (ABPM) during the interdialytic period in a group of 164 hypertensive patients, the blood pressure (BP) control based on conventional antihypertensive strategy previously, on chronic hemodialysis treatment in the Mediterranean region of Turkey. These results were then compared with their echocardiographic data. This is a cross-sectional analysis. The mean ABPM during 44 hours was close to the manually measured predialysis value, but there was a gradual increase in the ABPM values in the interdialytic period. When divided into a group with mild or no left ventricular hypertrophy (LVH) (45 patients) and severe LVH (119 patients), the latter had significantly higher BP levels in all separate periods, while the difference in predialysis BP was not significant. Patients with severe LVH had larger left atrium and left ventricular diameters, and consumed more antihypertensive drugs. Systolic BP during the night before dialysis showed the strongest relation to LVH, but interdialytic weight gain was also independently related to LVH. Yet, 56% of the patients with systolic BP <135 had severe LVH. There is not only an association between BP and presence of LVH, but it is shown that volume expansion is also an important independent determinant of LVH. This may explain the difficulty in identifying hypertension as a cardiac risk factor in these patients. [source]


    Direct and indirect methods to simulate the actual evapotranspiration of an irrigated overhead table grape vineyard under Mediterranean conditions

    HYDROLOGICAL PROCESSES, Issue 2 2008
    Gianfranco Rana
    Abstract Two methods, indirect and direct, for simulating the actual evapotranspiration (E) were applied to an irrigated overhead table grape vineyard during summer, situated in the Mediterranean region (south Italy), over two successive years. The first method, indirect but more practical, uses the crop coefficient (Kc) approach and requires determination of the reference evapotranspiration E0 (FAO (Food and Agriculture Organization) method). This method underestimated on average by 17% the daily values of the actual evapotranspiration E. The analysis in this paper shows that the values of Kc for the table grapes determined by the FAO method seem to not be valid in our experimental conditions. Similar conclusions can be found in the literature for the table grape cultivated under different experimental conditions and using different training systems. The second method, is a direct method for estimating the evapotranspiration. It requires development of a model for the overhead table grape vineyard E, following the Penman,Monteith one-step approach, and using standard meteorological variables as inputs for the determination of the canopy resistance. This method, which needs a particularly simple calibration, provided a better simulation of the hourly and daily evapotranspiration than the indirect method. In additon, the standard error of the daily values for the direct method ( ± 0 · 41 mm) was about 50% lower than that obtained for the indirect method, also when the indirect method used a locally calibrated coefficient Kc instead of a generic Kc. Both, for practical application and theoretical issues, the advantages and disadvantages linked to the use of each tested method are discussed in detail. Copyright © 2007 John Wiley & Sons, Ltd. [source]


    Spatial and temporal 850 hPa air temperature and sea-surface temperature covariances in the Mediterranean region and their connection to atmospheric circulation

    INTERNATIONAL JOURNAL OF CLIMATOLOGY, Issue 6 2002
    C. J. Lolis
    Abstract The spatial and temporal covariability between the lower troposphere and sea surface temperatures (SSTs) are studied in the Mediterranean basin for the period 1958,98. Monthly air temperature anomalies for the 850 hPa pressure level (T-850hPa) at 2.5° × 2.5° grid points and SST anomalies in 5° × 5° grid boxes are utilized. As a first step, factor analysis is applied on both sets of data in order to reduce their dimensionality. Then, canonical correlation analysis is applied and this leads to one statistically significant pair of canonical variates for winter and to two pairs for summer. In winter, a teleconnection (see-saw) between western Europe and the eastern Mediterranean at the 850 hPa level is revealed, and a corresponding weaker one between the areas of central-west and eastern Mediterranean for SST. The correlation between T-850hPa and SST appears higher over the eastern Mediterranean. In summer, the first pair of canonical variates reveals a covariability between T-850hPa and SST in the western Mediterranean, and the second one shows a covariability in the eastern Mediterranean, without the existence of any strong spatial teleconnection. The analysis is repeated, using time lags of 1 month, or longer, in order to detect any possible non-synchronous relation. Statistically significant results are found only when T-850hPa leads SST with a time lag of 1 month. In particular, the results are statistically significant for winter only, and the findings are similar to those of the first analysis. Therefore, the existence of a 1 month time scale SST persistence is detected for winter months. Copyright © 2002 Royal Meteorological Society. [source]


    Floristic patterns and plant traits of Mediterranean communities in fragmented habitats

    JOURNAL OF BIOGEOGRAPHY, Issue 7 2006
    Guillem Chust
    Abstract Aim, To contrast floristic spatial patterns and the importance of habitat fragmentation in two plant communities (grassland and scrubland) in the context of ecological succession. We ask whether plant assemblages are affected by habitat fragmentation and, if so, at what spatial scale? Does the relative importance of the niche differentiation and dispersal-limitation mechanisms change throughout secondary succession? Is the dispersal-limitation mechanism related to plant functional traits? Location, A Mediterranean region, the massif of Albera (Spain). Methods, Using a SPOT satellite image to describe the landscape, we tested the effect of habitat fragmentation on species composition, determining the spatial scale of the assemblage response. We then assessed the relative importance of dispersal-related factors (habitat fragmentation and geographical distance) and environmental constraints (climate-related variables) influencing species similarity. We tested the association between dispersal-related factors and plant traits (dispersal mode and life form). Results, In both community types, plant composition was partially affected by the surrounding vegetation. In scrublands, animal-dispersed and woody plants were abundant in landscapes dominated by closed forests, whereas wind-dispersed annual herbs were poorly represented in those landscapes. Scrubby assemblages were more dependent on geographical distance, habitat fragmentation and climate conditions (temperature, rainfall and solar radiation); grasslands were described only by habitat fragmentation and rainfall. Plant traits did not explain variation in spatial structuring of assemblages. Main conclusions, Plant establishment in early Mediterranean communities may be driven primarily by migration from neighbouring established communities, whereas the importance of habitat specialization and community drift increases over time. Plant life forms and dispersal modes did not explain the spatial variation of species distribution, but species richness within the community with differing plant traits was affected by habitat patchiness. [source]


    Mid-Holocene and glacial-maximum vegetation geography of the northern continents and Africa

    JOURNAL OF BIOGEOGRAPHY, Issue 3 2000
    I. Colin Prentice
    Abstract BIOME 6000 is an international project to map vegetation globally at mid-Holocene (6000 14C yr bp) and last glacial maximum (LGM, 18,000 14C yr bp), with a view to evaluating coupled climate-biosphere model results. Primary palaeoecological data are assigned to biomes using an explicit algorithm based on plant functional types. This paper introduces the second Special Feature on BIOME 6000. Site-based global biome maps are shown with data from North America, Eurasia (except South and Southeast Asia) and Africa at both time periods. A map based on surface samples shows the method's skill in reconstructing present-day biomes. Cold and dry conditions at LGM favoured extensive tundra and steppe. These biomes intergraded in northern Eurasia. Northern hemisphere forest biomes were displaced southward. Boreal evergreen forests (taiga) and temperate deciduous forests were fragmented, while European and East Asian steppes were greatly extended. Tropical moist forests (i.e. tropical rain forest and tropical seasonal forest) in Africa were reduced. In south-western North America, desert and steppe were replaced by open conifer woodland, opposite to the general arid trend but consistent with modelled southward displacement of the jet stream. The Arctic forest limit was shifted slighly north at 6000 14C yr bp in some sectors, but not in all. Northern temperate forest zones were generally shifted greater distances north. Warmer winters as well as summers in several regions are required to explain these shifts. Temperate deciduous forests in Europe were greatly extended, into the Mediterranean region as well as to the north. Steppe encroached on forest biomes in interior North America, but not in central Asia. Enhanced monsoons extended forest biomes in China inland and Sahelian vegetation into the Sahara while the African tropical rain forest was also reduced, consistent with a modelled northward shift of the ITCZ and a more seasonal climate in the equatorial zone. Palaeobiome maps show the outcome of separate, independent migrations of plant taxa in response to climate change. The average composition of biomes at LGM was often markedly different from today. Refugia for the temperate deciduous and tropical rain forest biomes may have existed offshore at LGM, but their characteristic taxa also persisted as components of other biomes. Examples include temperate deciduous trees that survived in cool mixed forest in eastern Europe, and tropical evergreen trees that survived in tropical seasonal forest in Africa. The sequence of biome shifts during a glacial-interglacial cycle may help account for some disjunct distributions of plant taxa. For example, the now-arid Saharan mountains may have linked Mediterranean and African tropical montane floras during enhanced monsoon regimes. Major changes in physical land-surface conditions, shown by the palaeobiome data, have implications for the global climate. The data can be used directly to evaluate the output of coupled atmosphere-biosphere models. The data could also be objectively generalized to yield realistic gridded land-surface maps, for use in sensitivity experiments with atmospheric models. Recent analyses of vegetation-climate feedbacks have focused on the hypothesized positive feedback effects of climate-induced vegetation changes in the Sahara/Sahel region and the Arctic during the mid-Holocene. However, a far wider spectrum of interactions potentially exists and could be investigated, using these data, both for 6000 14C yr bp and for the LGM. [source]


    Investigating the presence of pesticide transformation products in water by using liquid chromatography-mass spectrometry with different mass analyzers,

    JOURNAL OF MASS SPECTROMETRY (INCORP BIOLOGICAL MASS SPECTROMETRY), Issue 2 2008
    Félix Hernández
    Abstract Many pesticide transformation products (TPs) can reach environmental waters as a consequence of their normally having a higher polarity than their parent pesticides. This makes the development of analytical methodology for reliable identification and subsequent quantification at the sub-microgram per liter levels necessary, as required under current legislation. In this paper we report the photodegradation of several pesticides frequently detected in environmental waters from the Spanish Mediterranean region using the high-resolution and exact-mass capabilities of hybrid quadrupole time-of-flight mass spectrometry (QTOF MS) hyphenated to liquid chromatography (LC). Once the main photodegradation/hydrolysis products formed in aqueous media were identified, analytical methodology for their simultaneous quantification and reliable identification in real water samples was developed using on-line solid-phase extraction (SPE)-LC-tandem MS with a triple-quadrupole (QqQ) analyzer. The methodology was validated in both ground and surface water samples spiked at the limit of quantification (LOQ) and 10 × LOQ levels, i.e. 50 and 500 ng/l, obtaining satisfactory recoveries and precision for all compounds. Subsequent analysis of ground and surface water samples resulted in the detection of a number of TPs higher than parent pesticides. Additionally, several soil-interstitial water samples collected from the unsaturated zone were analyzed to explore the degradation/transformation of some pesticides in the field using experimental plots equipped with lisimeters. Several TPs were found in these samples, with most of them having also been detected in ground and surface water from the same area. This paper illustrates the extraordinary potential of LC-MS(/MS) with QTOF and QqQ analyzers for qualitative/structural and quantitative analysis, respectively, offering analytical chemists one of the most powerful tools available at present to investigate the presence of pesticide TPs in water. Copyright © 2007 John Wiley & Sons, Ltd. [source]


    EFFECT OF COPPER ON ALGAL COMMUNITIES FROM OLIGOTROPHIC CALCAREOUS STREAMS1

    JOURNAL OF PHYCOLOGY, Issue 2 2002
    Helena Guasch
    Two sets of experiments were done to quantify the effects of chronic copper exposure on natural peri- phyton in a nonpolluted calcareous river. The results of short-term (up to 6 h exposure) experiments corroborated the significance of pH on copper toxicity. Copper toxicity increased when pH was reduced from 8.6 to 7.7, and this was related to the effect of pH on copper speciation (free copper concentration increased from 0.2% to 2.3% of total copper). Longer term experiments demonstrated that periphyton communities exposed to copper under pH variation (8.2,8.6) were already affected at 10 ,g·L,1 (20,80 ng·L,1 Cu2+) after 12 days of exposure. Copper exposure caused stronger effects on structural (algal biomass and community structure) than on functional (photosynthetic efficiency) parameters of peri- phyton. Changes in community composition included the enhancement of some taxa (Gomphonema gracile), the inhibition of others (Fragilaria capucina and Phormidium sp.), and the appearance of filament malformations (Mougeotia sp.). The results of our study demonstrated that several weeks of exposure to copper (10,20 ,g·L,1) were sufficient to cause chronic changes in the periphyton of oligotrophic calcareous rivers. This degree of copper pollution can be commonly found in the Mediterranean region as a result of agricultural practices and farming activities. [source]


    Level of trace elements in Pteridophytes growing on serpentine and metalliferous soils

    JOURNAL OF PLANT NUTRITION AND SOIL SCIENCE, Issue 6 2007
    Laura Cornara
    Abstract A screening of Pteridophytes growing on serpentine and metalliferous soils in Northern Italy was carried out to assess the ability of these plants to tolerate or accumulate trace elements of toxicological interest. Few data are available on metal tolerance/accumulation of terrestrial ferns growing in the Mediterranean region, and several species presented here have never been investigated for this purpose. The trace-element composition (As, Cd, Cr, Cu, Fe, Ni, Pb, Zn) and P content of aerial parts of plants (12 fern and 2 horsetail species) and of their associated soils were measured. An analysis of the relationship between element concentrations in soil and in shoot revealed a significant correlation only for Zn (p < 0.05). Hierarchical cluster analysis based on element concentrations in plant aerial parts showed two outliers, viz. Equisetum ramosissimum Desf., showing the highest levels of Cr, Fe, Cu, Ni, and As, and Nephrolepis cordifolia C. Presl., showing the highest Pb value. The bioaccumulation factor exceeded 1 only for Cd in two species, Athyrium filix-femina and Dryopteris filix-mas. However, also in these cases the corresponding values of the metal in the shoots were below the thresholds for hyperaccumulators. The examined Pteridophytes seem to have developed their adaptation prevalently through mechanisms of tolerance based on metal exclusion. None of these plant species seem suitable for phytoextraction, but N. cordifolia, Pteridium aquilinum ssp. aquilinum, and E. ramosissimum have potential to be used for stabilization and restoration of soils rich in heavy metals. [source]


    A multicentury perspective on the summer North Atlantic Oscillation (SNAO) and drought in the eastern Atlantic Region,

    JOURNAL OF QUATERNARY SCIENCE, Issue 5 2009
    Hans W. Linderholm
    Abstract The summer North Atlantic Oscillation (SNAO) is strongly associated with July,August climate variability over Europe, especially in northern regions. This association includes drought, where a positive SNAO corresponds to dry conditions over much of northern Europe and wet conditions in southern Europe, but the SNAO/climate association is weaker and less homogeneous in the south. Here we use a dendroclimatological reconstruction of the SNAO for the last 550 a to investigate the SNAO/drought relationship in the past. An association between the SNAO and a regional summer drought index from Sweden suggests that the northern European drought relationship holds back to 1700. In the last 550 a, the relationship between SNAO and drought in the Mediterranean region as a whole is weak, but over the Eastern Mediterranean the relationship is clearer and statistically significant (P,<,0.05 level). The Mediterranean relationship is clearest at century scales. An association between the SNAO and Sahel rainfall can clearly be seen on interannual as well as longer timescales in the 20th century. Past droughts in the Sahel, as inferred from historical data, correspond quite well with positive phases of the SNAO on multidecadal timescales back to 1500, the phase expected from instrumental data. The physical reasons for the relationship between Sahel rainfall and the SNAO are, however, not yet understood. This research is a first step towards understanding how the atmospheric circulation over the North Atlantic region affects drought, necessary for forecast future droughts. Copyright © 2009 John Wiley & Sons, Ltd. and © Crown Copyright 2009. [source]


    Sub-millennial climate shifts in the western Mediterranean during the last glacial period recorded in a speleothem from Mallorca, Spain,

    JOURNAL OF QUATERNARY SCIENCE, Issue 8 2008
    Edward J. Hodge
    Abstract Very few high-resolution and directly dated terrestrial archives of the last glacial period exist for the western Mediterranean region, yet this is a key locality for recording sub-millennial North Atlantic and Mediterranean climate change. Here, we present evidence of effective precipitation changes based on growth history and ,13C of calcite in a Mallorcan stalagmite that grew between 112 and 48,ka. Effective precipitation in Mallorca appears to have been sensitive to proximal sea surface temperature variations and at certain times, ca. 76,ka for example, changed rapidly from moist to arid conditions in only a few centuries. A sea-level highstand during Marine Isotope Stage (MIS) 5a interrupted growth. Regrowth started promptly after this, but effective precipitation decreased markedly for much of the later part of MIS 5a, and also for shorter periods correlative with Heinrich events H8 (ca. 90,ka) and H6 (ca. 65,ka), with growth ceasing during H5 (ca. 48,ka). Arid episodes in Mallorca appear to be expressions of extremely cold periods recorded further north in Europe and occur contemporaneously with rapid decreases in Greenland temperature. Copyright © 2008 John Wiley & Sons, Ltd. [source]