Mesozoic

Distribution by Scientific Domains
Distribution within Earth and Environmental Science

Kinds of Mesozoic

  • late mesozoic

  • Terms modified by Mesozoic

  • mesozoic stratum

  • Selected Abstracts


    History of Deforestation and Reforestation in the Dinaric Karst

    GEOGRAPHICAL RESEARCH, Issue 1 2009
    ANDREJ KRANJC
    Abstract The term karst derives from the Kras plateau, which is in the northwestern part of the area now known as the Dinaric Karst. The landscape consists mostly of Mesozoic carbonate rocks and stretches along the Adriatic Sea coast for a distance of 600 km. Although the region lies parallel to the sea, the Mediterranean temperature influence is limited to a narrow coastal belt, except for the amount of precipitation, which can reach 5000 mm yr,1. Forests belonging to the Mediterranean and Euro-Siberian , North American region, covered the primary Dinaric Karst. Human deforestation of the Karst began during the Neolithic period, 6500,6000 BC. Throughout history there have been two main reasons for deforestation; economic (the requirements of new land, pastures, timber use and trade), and social (local increases in population, mass migration, wars, raids). Mankind's perception of forest protection and preservation can be traced through documents going back to the 12th century. Reforestation is mentioned in some of them, but successful reforestation did not begin until the 1850s. Nowadays dense natural forests, extensive forest plantations, dry karst shrublands and also completely barren karst areas can all be found on the Dinaric Karst. [source]


    Developments in research concerning Mesozoic,Tertiary Tethys and neotectonics in the Isparta Angle, SW Turkey

    GEOLOGICAL JOURNAL, Issue 3-4 2003
    Alastair H. F. Robertson
    Abstract The Isparta Angle has played a critical role in the development of concepts concerning the tectonic evolution of the Mesozoic,Tertiary Neotethys in the Eastern Mediterranean region. Following early regional studies, mainly by the Mineral Research and Exploration Institute of Turkey (MTA), during the 1960s and 1970s, a French team mapped the area and confirmed a regional tectonostratigraphy of three great allochthonous systems of mainly Mesozoic,Early Tertiary age, termed the Antalya, Lycian and Hoyran-Bey,ehir-Had,m nappes. During the 1970s and 1980s a British group studied the Neotethyan evolution of what they termed the Antalya Complex, utilizing knowledge of plate tectonic processes. Throughout the 1970s and 1980s MTA systematically remapped the area at 1:25,000 scale. The root zone of the Antalya allochthon was either a southerly Neotethys, within and to the south of the Isparta Angle, or a northerly Neotethys, many hundreds of kilometres to the north. The southerly origin is nowadays favoured but some questions remain. Attention focused in the 1990s until present to the post-collisional, neotectonic evolution of the Isparta Angle and its links with the neighbouring Mediterranean Sea. Here, we trace the development of research and the ongoing debates concerning alternative tectonic concepts used to explain the evolution of the Isparta Angle from Mesozoic to Recent time. We conclude by outlining several tectonic models for the evolution of the Antalya allochthon within a southerly Neotethys that require to be tested by future field studies. Copyright © 2003 John Wiley & Sons, Ltd. [source]


    Palaeomagnetic evidence for the Gondwanian origin of the Taurides and rotation of the Isparta Angle, southern Turkey

    GEOLOGICAL JOURNAL, Issue 4 2002
    John D. A. Piper
    Abstract The Taurides, the southernmost of the three major tectonic domains that constitute present-day Turkey, were emplaced following consumption of the Tethyan Ocean in Late Mesozoic to mid-Tertiary times. They are generally assigned an origin at the northern perimeter of Gondwana. To refine palaeogeographic control we have investigated the palaeomagnetism of a range of Jurassic rocks. Forty-nine samples of Upper Jurassic limestones preserve a dual polarity remanence (D/I=303/,9°, ,95=6°) interpreted as a primary magnetization acquired close to the equator and rotated during emplacement of the Taurides. Result from mid-Jurassic dolerites confirm a low palaeolatitude for the Tauride Platform during Jurassic times at the Afro,Arabian sector of Gondwana. Approximately 4000,km of Tethyan closure subsequently occurred between Late Jurassic and Eocene times. Although related Upper Jurassic limestones and Liassic redbeds preserve a sporadic record of similar remanence, the dominant signature in these latter rocks is an overprint of probable mid-Miocene age, probably acquired during a single polarity chron and imparted by migration of a fluid front during nappe loading. This is now rotated consistently anticlockwise by c. 30° and conforms to results of previous studies recording bulk Neogene rotation of the Isparta region following Lycian nappe emplacement. The regional distribution of this overprint implies that the Isparta Angle (IA) has been subject to only small additional closure (<10°) since Late Miocene time. A smaller amount (c. 6°) of clockwise rotation within the IA since Early Pliocene times is associated with an ongoing extensional regime and reflects an expanding curvature of the Tauride arc produced by southwestward extrusion of the Anatolian collage as a result of continuing northward motion of Afro,Arabia. Copyright © 2002 John Wiley & Sons, Ltd. [source]


    Mesozoic,Paleogene sedimentary facies and paleogeography of Tibet, western China: tectonic implications

    GEOLOGICAL JOURNAL, Issue 3 2002
    Kai-Jun Zhang
    Abstract In Early,Middle Triassic time, an abyssal sea covered most of the Songpan,Ganzi area, whereas a Central Tibetan Landmass, up to 400,km wide, may have stretched across the Lhasa and Western Qiangtang terrains. In Late Triassic time, the Songpan,Ganzi sea closed, the Central Tibetan Landmass receded westwards away from southern Western Qiangtang, a littoral environment dominated Eastern Qiangtang, middle Western Qiangtang, and southeastern Lhasa, a shelf environment existed only in northern and southeastern Western Qiangtang and northwestern Eastern Qiangtang, and abyssal flysch was spread along the eastern Bangonghu,Nüjiang zone. In Early,Middle Jurassic time, Songpan,Ganzi had become part of the Eurasian continent, abyssal flysch sediments stretched throughout the Bangonghu,Nüjiang zone, the Central Tibetan Landmass was only locally present in southwestern Lhasa, and the Tethyan epicontinental sea nearly covered all Tibet southwest of the Jinsajiang suture. In Late Jurassic time, oceanic flysch deposition existed only along the westernmost Bangonghu,Nüjiang zone, nearly all of Tibet was covered by coastal deposits, and shelf deposits existed only in northern Western Qiangtang and westernmost Lhasa. In the early stage of Early Cretaceous time, the majority of Qiangtang had become dry land, and a supralittoral environment dominated across the entire Lhasa terrain. However, during the late stage of the Early Cretaceous time, platform,shelf carbonates prevailed on southern Western Qiangtang and northern Lhasa. In Late Cretaceous time, the majority of Qiangtang had become emergent land, and a supratidal environment dominated Lhasa, the western rim of Western Qiangtang, and Tarim. In Paleogene time, the majority of Tibet became emergent land, and a supratidal environment existed only on the southern and western rims. The dominance of Upper Triassic,Jurassic shelf carbonates on the northwestern Eastern Qiangtang corner and the northern Western Qiangtang rim suggests a diachronous closing of the Jinsajiang paleo-Tethys ocean, first during latest Triassic time when the Eastern Qiangtang terrain collided with Asia and finally in Jurassic time when the Western Qiangtang terrain was amalgamated to Asia. Rich picotites in Upper Triassic sandstones of middle Qiangtang suggest that the Shuanghu suture could have extended along the middle of Qiangtang, and stable shelf sedimentation during Late Triassic,Middle Jurassic time in the Western Qiangtang terrain shows that the suture probably could not have formed until Middle Jurassic time. The opening time of the Bangonghu,Nüjiang mid-Tethys ocean could be Late Triassic time due to the existence of the Central Tibetan Landmass across Western Qiangtang and Lhasa during Early,Middle Triassic time. However, its opening was diachronous, at Late Triassic time in the east and at Early,Middle Jurassic time in the west. Furthermore, its closing was also diachronous, first in the east at the beginning of Late Jurassic time and later in the west in latest Jurassic to earliest Cretaceous time. Widespread upper Lower Cretaceous limestone up to 5,km thick over the northern half of Lhasa indicates that southern Tibet could have undergone an extensive backarc subsidence during late Early Cretaceous time. Continuous shallow marine sedimentation through the entire Cretaceous time over much of southern Tibet indicates that southern Tibet was intensely elevated only after the end of Paleogene time, its high topography being the product of the Indo-Asian collision. Copyright © 2002 John Wiley & Sons, Ltd. [source]


    The Nordic Volcanological Institute: understanding volcanoes at spreading centres

    GEOLOGY TODAY, Issue 2 2009
    Kent Brooks
    The Nordic countries (known as ,Norden') are not immediately associated with volcanoes: Norway with folded mountains cut by fjords and its offshore oil and gas deposits, Sweden and Finland by the western part of the Baltic shield, a huge area of Precambrian rocks, of which gneisses form a large part, and Denmark, a country of Mesozoic and Cenozoic rocks, where glacial, superficial deposits are of major importance. But Norden also includes Iceland, where everyone immediately thinks of volcanoes and glaciers. Clearly volcanological research would be expected to be a major priority for the Icelandic nation. However, in the other Nordic countries old volcanic and other igneous rocks play a significant role, comprising a large part of the Precambrian and Caledonian terrains and being a key to many of the commercial mineral deposits which play a major role in the economies of Norway, Sweden and Finland. Even Denmark, a country of sedimentary rocks has an impressive sequence of Paleogene volcanic ashes and the Faeroe Islands, made up almost entirely of basalts, are part of Denmark. [source]


    A constrained 2D gravity model of the Sebastián Vizcaíno Basin, Baja California Sur, Mexico

    GEOPHYSICAL PROSPECTING, Issue 6 2005
    J. García-Abdeslem
    ABSTRACT The subsurface geometry of the Sebastián Vizcaíno Basin is obtained from the 2D inversion of gravity data, constrained by a density-versus-depth relationship derived from an oil exploration deep hole. The basin accumulated a thick pile of marine sediments that evolved in the fore-arc region of the compressive margin prevalent along western North America during Mesozoic and Tertiary times. Our interpretation indicates that the sedimentary infill in the Sebastián Vizcaíno Basin reaches a maximum thickness of about 4 km at the centre of a relatively symmetric basin. At the location of the Suaro-1 hole, the depth to the basement derived from this work agrees with the drilled interface between calcareous and volcaniclastic members of the Alisitos Formation. A sensitivity analysis strongly suggests that the assumed density function leads to a nearly unique solution of the inverse problem. [source]


    Robust support for tardigrade clades and their ages from three protein-coding nuclear genes

    INVERTEBRATE BIOLOGY, Issue 2 2004
    Jerome C. Regier
    Abstract. Coding sequences (5,334 nt total) from elongation factor-1,, elongation factor-2, and the largest subunit of RNA polymerase II were determined for 6 species of Tardigrada, 2 of Arthropoda, and 2 of Onychophora. Parsimony and likelihood analyses of nucleotides and amino acids yielded strong support for Tardigrada and all internal nodes (i.e., 100% bootstrap support for Tardigrada, Eutardigrada, Parachela, Hypsibiidae, and Macrobiotidae). Results are in agreement with morphology and an earlier molecular study based on analysis of 18S ribosomal sequences. Divergence times have been estimated from amino acid sequence data using an empirical Bayesian statistical approach, which does not assume a strict molecular clock. Divergence time estimates are pre-Vendian for Tardigrada/Arthropoda, Vendian or earlier for Eutardigrada/Heterotardigrada, Silurian to Ordovician for Parachela/Apochela, Permian to Carboniferous for Hypsibiidae and Macrobiotidae, and Mesozoic for Isohypsibius/Thulinia (both within Hypsibiidae) and Macrobiotus/Richtersius (both within Macrobiotidae). [source]


    Late Cretaceous-Cenozoic exhumation of the Yanji area, northeast China: Constraints from fission-track thermochronology

    ISLAND ARC, Issue 1 2010
    Xiaoming Li
    Abstract The Yanji area, located at the border of China, Russia, and Korea, where the Phanerozoic granitoids have been widely exposed, was considered part of the orogenic collage between the North China Block in the south and the Jiamusi,Khanka Massifs in the northeast. In this study, the cooling and inferred uplift and denudation history since the late Mesozoic are intensively studied by carrying out apatite and zircon fission-track analyses, together with electron microprobe analyses (EMPA) of chemical compositions of apatite from the granitoid samples in the Yanji area. The results show that: (i) zircon and apatite fission-track ages range 91.7,99.6 Ma and 76.5,85.4 Ma, respectively; (ii) all apatite fission-track length distributions are unimodal and yield mean lengths of 12,13.2 µm, and the apatites are attributed to chlorine-bearing fluorapatite as revealed by EMPA results; and (iii) the thermal history modeling results based on apatite fission-track grain ages and length distributions indicate that the time,temperature paths display similar patterns and the cooling has been accelerated for each sample since ca 15 Ma. Thus, we conclude that sequential cooling, involving two rapid (95,80 Ma and ca 15,0 Ma) and one slow (80,15 Ma) cooling, has taken place through the exhumation of the Yanji area since the late Cretaceous. The maximum exhumation is more than 5 km under a steady-state geothermal gradient of 35°C/km. Combined with the tectonic setting, this exhumation is possibly related to the subduction of the Pacific Plate beneath the Eurasian Plate since the late Cretaceous. [source]


    Structure of Sumatra and its implications for the tectonic assembly of Southeast Asia and the destruction of Paleotethys

    ISLAND ARC, Issue 1 2009
    Anthony J. Barber
    Abstract It is now generally accepted that Southeast Asia is composed of continental blocks which separated from Gondwana with the formation of oceanic crust during the Paleozoic, and were accreted to Asia in the Late Paleozoic or Early Mesozoic, with the subduction of the intervening oceanic crust. From east to west the Malay peninsula and Sumatra are composed of three continental blocks: East Malaya with a Cathaysian Permian flora and fauna; Sibumasu, including the western part of the Malay peninsula and East Sumatra, with Late Carboniferous,Early Permian ,pebbly mudstones' interpreted as glaciogenic diamictites; and West Sumatra, again with Cathaysian fauna and flora. A further unit, the Woyla nappe, is interpreted as an intraoceanic arc thrust over the West Sumatra block in the mid Cretaceous. There are varied opinions concerning the age of collision of Sibumasu with East Malaya and the destruction of Paleotethys. In Thailand, radiolarites have been used as evidence that Paleotethys survived until after the Middle Triassic. In the Malay peninsula, structural evidence and the ages of granitic intrusions are used to support a Middle Permian to Early Triassic age for the destruction of Paleotethys. It is suggested that the West Sumatra block was derived from Cathaysia and emplaced against the western margin of Sibumasu by dextral transcurrent faulting along a zone of high deformation, the Medial Sumatra Tectonic Zone. These structural units can be traced northwards in Southeast Asia. The East Malaya block is considered to be part of the Indochina block, Sibumasu can be traced through Thailand into southern China, the Medial Sumatra Tectonic Zone is correlated with the Mogok Belt of Myanmar, the West Burma block is the extension of the West Sumatra block, from which it was separated by the formation of the Andaman Sea in the Miocene, and the Woyla nappe is correlated with the Mawgyi nappe of Myanmar. [source]


    Zircon U,Pb ages and tectonic implications of ,Early Paleozoic' granitoids at Yanbian, Jilin Province, northeast China

    ISLAND ARC, Issue 4 2004
    Yanbin Zhang
    Abstract The Yanbian area is located in the eastern part of the Central Asian Orogenic Belt (CAOB) of China and is characterized by widespread Phanerozoic granitic intrusions. It was previously thought that the Yanbian granitoids were mainly emplaced in the Early Paleozoic (so-called ,Caledonian' granitoids), extending east,west along the northern margin of the North China craton. However, few of them have been precisely dated; therefore, five typical ,Caledonian' granitic intrusions (the Huangniling, Dakai, Mengshan, Gaoling and Bailiping batholiths) were selected for U,Pb zircon isotopic study. New-age data show that emplacement of these granitoids extended from the Late Paleozoic to Late Mesozoic (285,116 Ma). This indicates that no ,Caledonian' granitic belt exists along the northern margin of the North China craton. The granitoids can be subdivided into four episodes based on our new data: Early Permian (285 ± 9 Ma), Early Triassic (249,245 Ma), Jurassic (192,168 Ma) and Cretaceous (119,116 Ma). The 285 ± 9 Ma tonalite was most likely related to subduction of the Paleo-Asian Oceanic Plate beneath the North China craton, followed by Triassic (249,245 Ma) syn-collisional monzogranites, representing the collision of the CAOB orogenic collage with the North China craton and final closure of the Paleo-Asian Ocean. The Jurassic granitoids resulted from subduction of the Paleo-Pacific plate and subsequent collision of the Jiamusi,Khanka Massif with the existing continent, assembled in the Triassic. The Early Cretaceous granitoids formed in an extensional setting along the eastern Asian continental margin. [source]


    Mesozoic to Cenozoic igneous activity and tectonism in the Ryoke Belt, Southwest Japan

    ISLAND ARC, Issue 1 2000
    Y. Hayama
    No abstract is available for this article. [source]


    Historical biogeography of scarabaeine dung beetles

    JOURNAL OF BIOGEOGRAPHY, Issue 9 2002
    Adrian L. V. Davis
    Abstract Aim, (1) To review briefly global biogeographical patterns in dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae), a group whose evolutionary history has been dominated by ecological specialization to vertebrate dung in warmer climates. (2) To develop hypotheses accounting for the evolution of these patterns. Location, Six principal biogeographical regions: Palaearctic, Oriental, Afrotropical, Australasia, Neotropical, Nearctic and five outlying islands or island groups harbouring endemic genera: Caribbean, Madagascar, Mauritius, New Caledonia, New Zealand. Methods, Major patterns of tribal, generic and species distribution are investigated using cluster analysis, ordination, parsimony analysis of endemism and track analysis. Attempts are made to resolve biogeographical patterns with findings in the fields of plate tectonics, fossil and evolutionary history, plus phylogeny of both mammals and dung beetles. Results, Because of conflict between published findings, it is uncertain at what point in time density of dinosaur dung, mammal dung or both became sufficiently great to select for specialized habits in dung beetles. However, biogeographical evidence would suggest a Mesozoic origin followed by further taxonomic radiation during the Cenozoic, possibly in response to the increasing size and diversity of mammalian dung types in South America and Afro-Eurasia. Proportional generic distribution in fourteen tribes and subtribes showed four principal biogeographical patterns: (1) southerly biased Gondwanaland distribution, (2) Americas or (3) Madagascar endemism, and (4) northerly biased, Afro-Eurasian-centred distribution with limited numbers of genera also widespread in other regions. Proportional composition of faunas in eleven geographical regions indicated three principal distributional centres, East Gondwanaland fragments, Afro-Eurasia and the Americas. These patterns probably result from three principal long-term range expansion and vicariance events (Mesozoic: Gondwanaland interchange and fragmentation, Cenozoic: Afro-Eurasian/Nearctic interchange and the Great American interchange). It is suggested that old vicariance caused by the Mesozoic fragmentation of Gondwanaland leads to a high degree of regional endemism at generic or tribal level across one or more Gondwanaland tracks. In contrast, it is suggested that the more recent Cenozoic range expansions occurred primarily towards northern regions leading to endemism primarily at species level. These Cenozoic radiations were facilitated by the re-linking of continents, either because of tectonic plate movements (Africa to Eurasia in Miocene), climatically induced sea-level change (Afro-Eurasia to Nearctic in Miocene and Pleistocene), or similar coupled with orogenics (Nearctic to Neotropical in Pliocene). Speciation has followed vicariance either because of climatic change or physical barrier development. These recent range expansions probably occurred principally along an Afro-Eurasian land track to the Nearctic and Neotropical and an Americas land track northwards from the Neotropics to the Nearctic, with limited dispersal from Eurasia to Australia, probably across a sea barrier. This accounts for the overall, spatially constrained, biogeographical pattern comprising large numbers of species-poor genera endemic to a single biogeographical region and fewer more species-rich genera, many of which show wider biogeographical distributions. In most southerly regions (Australasia, Madagascar, Neotropical), faunal composition and generic endemism is primarily dominated by elements with Gondwanaland ancestry, which is consistent with the Gondwanaland origin claimed for Scarabaeinae. In Afro-Eurasia (Palaearctic, Oriental, Afrotropical), generic endemism of monophyletically derived Afro-Eurasian and widespread lineages is centred in the Afrotropical region and faunal composition is numerically dominated by Afro-Eurasian and widespread elements. In the Nearctic region, the fauna is jointly dominated by widespread elements, derived from Afro-Eurasia, and Gondwanaland and Americas elements derived from the Neotropical region. Main conclusions, Global biogeographical patterns in scarabaeine dung beetles primarily result from Mesozoic and Cenozoic range expansion events followed by vicariance, although recent dispersal to Australia may have occurred across sea barriers. Detailed phylogenetics research is required to provide data to support dispersal/vicariance hypotheses. [source]


    Reconciling fossils and molecules: Cenozoic divergence of cichlid fishes and the biogeography of Madagascar

    JOURNAL OF BIOGEOGRAPHY, Issue 9 2001
    M. Vences
    Aim The biogeographical origins of the extant vertebrates endemic to Madagascar are largely unsolved, but have often been related to vicariance in the context of fragmentation of the supercontinent Gondwana in the Mesozoic. Such hypotheses are especially appealing in the case of cichlid fishes, which show phylogenetic relationships reflecting the temporal successions of the breakup of Gondwana. We used molecular clock data to test this assumption. Location Fragments of the 16S rRNA gene and of the nuclear Tmo-4C4 locus, partly obtained from Genbank from South American, African, Malagasy and Indian cichlids were analysed. Methods Based on monophyletic cichlid radiations in African lakes, we calibrated a molecular clock. The obtained rates were used to estimate the age of divergence of the major cichlid clades. Results The results agreed better with a Cenozoic than with a Mesozoic divergence, and were in accordance with the fossil record. Sequence divergences of the 16S and 12S rRNA genes of most lineages of Malagasy terrestrial and freshwater vertebrates from their non-Malagasy sister groups were below saturation and many were relatively similar to those of cichlids. Main conclusions A Cenozoic dispersal from continental landmasses may explain the origin of most extant Malagasy vertebrate groups better than a Jurassic/Cretaceous vicariance. [source]


    Skull shape and feeding strategy in Sphenodon and other Rhynchocephalia (Diapsida: Lepidosauria),

    JOURNAL OF MORPHOLOGY, Issue 8 2008
    Marc E.H. Jones
    Abstract The Rhynchocephalia are a group of small diapsid reptiles that were globally distributed during the early Mesozoic. By contrast, the only extant representatives, Sphenodon punctatus and S. guntheri (Tuatara), are restricted to New Zealand off-shore islands. The Rhynchocephalia are widely considered to be morphologically uniform but research over the past 30 years has revealed unexpected phenotypic and taxonomic diversity. Phylogenetically "basal taxa" generally possess relatively simple conical or columnar teeth whereas more derived taxa possessed stouter flanged teeth and sophisticated shearing mechanisms: orthal in some (e.g., Clevosaurus hudsoni) and propalinal in others (e.g., S. punctatus). This variation in feeding apparatus suggests a wide range of feeding niches were exploited by rhynchocephalians. The relationship of skull shape to skull length, phylogenetic grouping, habit, and characters relating to the feeding apparatus are explored here with geometric morphometric analysis on two-dimensional landmarks. Principle components analysis demonstrates that there are significant differences between phylogenetic groups. In particular, Sphenodon differs significantly from all well known fossil taxa including the most phylogenetically basal forms. Therefore, it is not justifiable to use Sphenodon as a solitary outgroup when studying skull shape and feeding strategy in squamates; rhynchocephalian fossil taxa also need to be considered. There are also significant differences between the skull shapes of aquatic taxa and those of terrestrial taxa. Of the observed variation in skull shape, most variation is subsumed by variation in dentary tooth base shape, the type of jaw movement employed (e.g., orthal vs. propalinal) and the number of palatal tooth rows. By comparison, the presence or absence of flanges, dentary tooth number and palatal tooth row orientation subsume much less. Skull length was also found to be a poor descriptor of overall skull shape. Compared to basal rhynchocephalians members of more derived terrestrial radiations possess an enlarged postorbital area, a high parietal, and a jaw joint positioned ventral to the tooth row. Modification of these features is closely associated with increased biting performance and thus access to novel food items. Some of these same trends are apparent during Sphenodon ontogeny where skull growth is allometric and there is evidence for ontogenetic variation in diet. J. Morphol., 2008. © 2008 Wiley-Liss, Inc. [source]


    The structural evolution of the English Channel area

    JOURNAL OF QUATERNARY SCIENCE, Issue 3-4 2003
    J. L. Lagarde
    Abstract The structural evolution of the English Channel area is controlled by structure and particularly by the pre-existing Cadomian and Variscan crustal discontinuities, which have been reactivated repeatedly in post-Variscan times. They controlled the crustal subsidence that produced basin development in the Mesozoic, prior to the sea-floor spreading in the North Atlantic region. They were then reactivated during the Cenozoic compression and basin inversion. The English Channel development is ascribed to mid-Tertiary differential uplift (Oligocene to Miocene). During late Tertiary to Quaternary times the Channel displays characteristics of a tectonically controlled fluvial basin periodically invaded by the sea. At the lithospheric scale, the Channel can be considered as an active intraplate area influenced by the NW,SE ,Alpine push', the NW,SE ,Atlantic ridge push' and glacial rebound stresses. Copyright © 2003 John Wiley & Sons, Ltd. [source]


    Phylogenetic signal and the utility of 12S and 16S mtDNA in frog phylogeny

    JOURNAL OF ZOOLOGICAL SYSTEMATICS AND EVOLUTIONARY RESEARCH, Issue 1 2004
    S. Hertwig
    Abstract Genes selected for a phylogenetic study need to contain conserved information that reflects the phylogenetic history at the specific taxonomic level of interest. Mitochondrial ribosomal genes have been used for a wide range of phylogenetic questions in general and in anuran systematics in particular. We checked the plausibility of phylogenetic reconstructions in anurans that were built from commonly used 12S and 16S rRNA gene sequences. For up to 27 species arranged in taxon sets of graded inclusiveness, we inferred phylogenetic hypotheses based on different a priori decisions, i.e. choice of alignment method and alignment parameters, including/excluding variable sites, choice of reconstruction algorithm and models of evolution. Alignment methods and parameters, as well as taxon sampling all had notable effects on the results leading to a large number of conflicting topologies. Very few nodes were supported in all of the analyses. Data sets in which fast evolving and ambiguously aligned sites had been excluded performed worse than the complete data sets. There was moderate support for the monophyly of the Discoglossidae, Pelobatoidea, Pelobatidae and Pipidae. The clade Neobatrachia was robustly supported and the intrageneric relationships within Bombina and Discoglossus were well resolved indicating the usefulness of the genes for relatively recent phylogenetic events. Although 12S and 16S rRNA genes seem to carry some phylogenetic signal of deep (Mesozoic) splitting events the signal was not strong enough to resolve consistently the inter-relationships of major clades within the Anura under varied methods and parameter settings. Zusammenfassung Zur Anwendung in einer phylogenetische Analyse müssen die ausgewählten Gene konservierte und detektierbare Information zum untersuchten phylogenetischen Niveau enthalten. Ribosomale Gene des Mitochondriums wurden für ein breites Spektrum phylogenetischer Fragestellungen bei verschiedenen Gruppen und insbesondere bei Froschlurchen eingesetzt. Wir untersuchten die Frage, ob Rekonstruktionen der Anuren-Phylogenie, basierend auf 12S und 16S rRNA Gensequenzen, plausibel sind. An einer Auswahl von 27 Arten, arrangiert in Taxa-Gruppen abgestufter Hierarchie, rekonstruierten wir phylogenetische Hypothesen unter verschiedenen, a priori festgelegten Bedingungen. Dazu gehörten die Auswahl verschiedener Alinierungsmethoden und,parameter, der Umgang mit variabel alinierten Positionen, die Auswahl der Algorithmen zur Baumkonstruktion sowie die Auswahl alternativer Modelle der Sequenzevolution. Die Methoden und Parameter der Alinierung und der Rekonstruktion, sowie die Auswahl der Taxa, hatten bedeutenden Einfluss auf die Resultate. Daraus resultierte eine große Anzahl alternativer Topologien, in denen nur sehr wenige Knoten in allen Analysen Unterstützung fanden. Ausschluss variabel alinierter Positionen ergaben Topologien mit niedrigem Grad der Auflösung. Die Sequenzen enthielten ein gewisses Signal für die Monophylie von Discoglossidae, Pelobatoidea, Pelobatidae und Pipidae. Der Knoten Neobatrachia wurde deutlich unterstützt. Die robuste Auflösung intragenerischer Phylogenien von Bombina und Discoglossus weisen auf eine besondere Eignung der Gene für die Untersuchung junger Aufspaltungsereignisse hin. Obwohl 12S und 16S rRNA-Gene eine heterogene Unterstützung für wenige frühe (mesozoische) phylogenetische Ereignisse zeigten, war das Signal nicht geeignet, um die Beziehungen der Taxa höherer Ordnung der Anura unter variierten Parametern und Analysemethoden konsistent aufzulösen. [source]


    Structure and diversity of the Mesozoic wood genus Xenoxylon in Far East Asia: implications for terrestrial palaeoclimates

    LETHAIA, Issue 4 2009
    MARC PHILIPPE
    Although the faunal elements of Far East Asian Mesozoic terrestrial biota have attracted much attention in recent years, their palaeoecology remains poorly known. In particular, features of the palaeoclimate are highly controversial. To address this point we used the Mesozoic fossil wood Xenoxylon, a genus recognized as an indicator of wet temperate biotopes and which is common in the area during the Carnian,Maastrichtian interval. We re-appraised bibliographic data and gathered new data for Xenoxylon in the Mesozoic of Far East Asia. This demonstrated that previous taxonomic approaches to the genus have been so far idiosyncratic. We examined the anatomical diversity of morphogenus Xenoxylon in Far East Asia and compared it to that of samples from Europe. This indicates that in an area centred on north-eastern China, Xenoxylon reached a level of anatomical diversity unmatched elsewhere in the world. We hypothesize that this diversity witnesses the persistence of palaeoecological conditions particularly suitable for Xenoxylon and that a wet temperate climate prevailed over most of the area throughout the Carnian,Maastrichtian interval. It is in this setting that the famous Jehol Biota probably evolved. [source]


    Soft-tissue imprints in fossil and Recent cephalopod septa and septum formation

    LETHAIA, Issue 4 2008
    CHRISTIAN KLUG
    Several soft-tissue imprints and attachment sites have been discovered on the inside of the shell wall and on the apertural side of the septum of various fossil and Recent ectocochleate cephalopods. In addition to the scars of the cephalic retractors, steinkerns of the body chambers of bactritoids and some ammonoids from the Moroccan and the German Emsian (Early Devonian) display various kinds of striations; some of these striations are restricted to the mural part of the septum, some start at the suture and terminate at the anterior limit of the annular elevation. Several of these features were also discovered in specimens of Mesozoic and Recent nautilids. These structures are here interpreted as imprints of muscle fibre bundles of the posterior and especially the septal mantle, blood vessels as well as the septal furrow. Most of these structures were not found in ammonoids younger than Middle Devonian. We suggest that newly formed, not yet mineralized (or only slightly), septa were more tightly stayed between the more numerous lobes and saddles in more strongly folded septa of more derived ammonoids and that the higher tension in these septa did not permit soft-parts to leave imprints on the organic preseptum. It is conceivable that this permitted more derived ammonoids to replace the chamber liquid faster by gas and consequently, new chambers could be used earlier than in other ectocochleate cephalopods, perhaps this process began even prior to mineralization. This would have allowed faster growth rates in derived ammonoids. [source]


    The Ordovician Biodiversification: revolution in the oceanic trophic chain

    LETHAIA, Issue 2 2008
    THOMAS SERVAIS
    The Early Palaeozoic phytoplankton (acritarch) radiation paralleled a long-term increase in sea level between the Early Cambrian and the Late Ordovician. In the Late Cambrian, after the SPICE ,13Ccarb excursion, acritarchs underwent a major change in morphological disparity and their taxonomical diversity increased to reach highest values during the Middle Ordovician (Darriwilian). This highest phytoplankton diversity of the Palaeozoic was possibly the result of palaeogeography (greatest continental dispersal) and major orogenic and volcanic activity, which provided maximum ecospace and large amounts of nutrients. With its warm climate and high atmospheric CO2 levels, the Ordovician was similar to the Cretaceous: a period when phytoplankton diversity was at its maximum during the Mesozoic. With increased phytoplankton availability in the Late Cambrian and Ordovician a radiation of zooplanktonic organisms took place at the same time as a major diversification of suspension feeders. In addition, planktotrophy originated in invertebrate larvae during the Late Cambrian,Early Ordovician. These important changes in the trophic chain can be considered as a major palaeoecological revolution (part of the rise of the Palaeozoic Evolutionary Fauna of Sepkoski). There is now sufficient evidence that this trophic chain revolution was related to the diversification of the phytoplankton, of which the organic-walled fraction is partly preserved. [source]


    Intense drilling in the Carboniferous brachiopod Cardiarina cordata Cooper, 1956

    LETHAIA, Issue 2 2003
    ALAN P. HOFFMEISTER
    The brachiopod Cardiarina cordata, collected from a Late Pennsylvanian (Virgilian) limestone unit in Grapevine Canyon (Sacramento Mts., New Mexico), reveals frequent drillings: 32.7% (n = 400) of these small, invariably articulated specimens (<2 mm size) display small (<0.2 mm), round often beveled holes that are typically single and penetrate one valve of an articulated shell. The observed drilling frequency is comparable with frequencies observed in the Late Mesozoic and Cenozoic. The drilling organism displayed high valve and site selectivity, although the exact nature of the biotic interaction recorded by drill holes (parasitism vs. predation) cannot be established. In addition, prey/host size may have been an important factor in the selection of prey/host taxa by the predator/parasite. These results suggest that drilling interactions occasionally occurred at high (Cenozoic-like) frequencies in the Paleozoic. However, such anomalously high frequencies may have been restricted to small prey/host with small drill holes. Small drillings in C. cordata, and other Paleozoic brachiopods, may record a different guild of predators/parasites than the larger, but less common, drill holes previously documented for Paleozoic brachiopods, echinoderms, and mollusks. [source]


    TAXONOMY AND ZOOGEOGRAPHY OF THE MESOZOIC CYTHERURID OSTRACODA FROM WEST-CENTRAL ARGENTINA

    PALAEONTOLOGY, Issue 1 2009
    SARA C. BALLENT
    Abstract:, The status of thirty four species of the cytheroidean ostracod family Cytheruridae from the Neuquén Basin, west-central Argentina is reviewed. These species belong to the following nine genera: Eucytherura Müller, ranging in this study from Pliensbachian to Valanginian and represented by fourteen species; Kangarina Coryell and Fields, Aalenian,Bajocian, with a single species; Acrocythere Neale, Hauterivian, one species; Paranotacythere Bassiouni, Berriasian, two species; Procytherura Whatley, Pliensbachian to Hauterivian, twelve species; Cytheropteron Sars, Aalenian,Bajocian, one species: Eocytheropteron Alexander, Hauterivian, one species; Paradoxorhyncha Chapman, Aalenian,Bajocian, one species; and Paracytheridea Müller, Berriasian, one species. Of the three subfamilies of the Cytheruridae, the Cytherurinae, with thirty species are by far the most numerous and abundant throughout the study. The other two subfamilies, the Cytheropterinae are represented by three species and the Paracytherideinae by a single species of the nominative genus. Several of the species are very widely distributed geographically and have, for example, also been recorded from Europe; others indicate close links with South Africa and Australia. The stratigraphical ranges of certain genera have been extended as a result of this study. For example, Kangarina has not previously been recorded below the Cretaceous. The Cytheruridae are clearly the most diverse cytheroidean ostracodes in the Mesozoic of the Neuquén Basin and are more diverse than all other groups of ostracodes combined. Four new species, Eucytherura tessae, Eucytherura yunga, Procytherura amygdala and Eocytheropteron immodicus are described. Eucytherura guillaumeae nom. nov. for Eucytherura tuberculata Brenner and Oertli and Eucytherura paranuda nom. nov. for Eucytherura nuda (Brand) are proposed. [source]


    A NEW SPINICAUDATAN GENUS (CRUSTACEA: ,CONCHOSTRACA') FROM THE LATE CRETACEOUS OF MADAGASCAR

    PALAEONTOLOGY, Issue 5 2008
    ALYCIA L. STIGALL
    Abstract:, A new spinicaudatan genus and species, Ethmosestheria mahajangaensis gen. et sp. nov., is described from the Anembalemba Member (Upper Cretaceous, Maastrichtian) of the Maevarano Formation, Mahajanga Basin, Madagascar. This is the first spinicaudatan reported from the post-Triassic Mesozoic of Madagascar. The new species is assigned to the family Antronestheriidae based on the cavernous or sievelike ornamentation on the carapace. Of well-documented Mesozoic spinicaudatan genera, Ethmosestheria mahajangaensis is most closely related to Antronestheria Chen and Hudson from the Great Estuarine Group (Jurassic) of Scotland. However, relatively poor documentation of the ornamentation of most Gondwanan Mesozoic spinicaudatan species precludes detailed comparison among taxa. Ethmosestheria mahajangaensis exhibits ontogenetic trends in carapace growth: a change in carapace outline from subcircular/subelliptical to elliptical, and from very wide juvenile growth bands to narrow adult growth bands. Ornamentation style, however, does not vary with ontogeny. Ethmosestheria mahajangaensis individuals lived in temporary pools in a broad channel-belt system within a semiarid environment; preserved desiccation structures on carapaces indicate seasonal drying out of pools within the river system. Specimens of Ethmosestheria mahajangaensis are preserved with exquisite detail in debris flow deposits; these are the first spinicaudatans reported from debris flow deposits. These deposits also contain a varied vertebrate fauna, including dinosaurs, crocodyliforms, turtles, and frogs. Rapid entombment of the spinicaudatan carapaces likely promoted early fossil diagenesis leading to highly detailed preservation. [source]


    A Special Orogenic-type Rare Earth Element Deposit in Maoniuping, Sichuan, China: Geology and Geochemistry

    RESOURCE GEOLOGY, Issue 3 2001
    Denghong WANG
    Abstract: The Maoniuping REE deposit is the second largest light rare earth elements deposit in China, explored recently in the northern Jinpingshan Mountains, a Cenozoic intracontinental orogenic belt in southwestern China. It is a vein-type deposit hosted within, and genetically related to, carbonatite-alkalic complex. Field investigation and new geochemical data of the carbonatites from the carbonatite-alkalic complex support an igneous origin for the Maoniuping carbonatites and related REE mineralization. Carbonatite itself carries rare earth elements which were enriched by hydrothermal solution. It is known that most of the REE deposits related to carbonatite-alkalic complexes were formed in relatively stable tectonic setting such as cratonic or rifting environment. The Maoniuping deposit, however, was formed during the processes of Cenozoic orogeny. Although the Maoniuping deposit is located in the north sector of the Panxi paleo-rift zone, the rift had been closed before early Cenozoic and evolved into an intracontinental orogenic belt, i.e., the Jinpingshan Orogen, which was formed since later Mesozoic to early Cenozoic. Geochronological and geochemical data also prove that the Maoniuping REE deposit was formed in an intracontinental orogenic belt instead of rift system or stationary block. The Maoniuping REE deposit is similar to the Mountain Pass REE deposit in many respects such as the high contents of bastnaesite and barite, the low content of niobium, and the common occurrence of sulfides. The discovery of the Maoniuping deposit and other REE deposits during the past two decades suggest a good potential for prospecting REE deposits along the alkalic complex belt located on the eastern side of the Qinghai,Xizang,West Sichuan Plateau. [source]


    Mudstone compaction curves in basin modelling: a study of Mesozoic and Cenozoic Sediments in the northern North Sea

    BASIN RESEARCH, Issue 3 2010
    Ø. Marcussen
    ABSTRACT Basin modelling studies are carried out in order to understand the basin evolution and palaeotemperature history of sedimentary basins. The results of basin modelling are sensitive to changes in the physical properties of the rocks in the sedimentary sequences. The rate of basin subsidence depends, to a large extent, on the density of the sedimentary column, which is largely dependent on the porosity and therefore on the rate of compaction. This study has tested the sensitivity of varying porosity/depth curves and related thermal conductivities for the Cenozoic succession along a cross-section in the northern North Sea basin, offshore Norway. End-member porosity/depth curves, assuming clay with smectite and kaolinite properties, are compared with a standard compaction curve for shale normally applied to the North Sea. Using these alternate relationships, basin geometries of the Cenozoic succession may vary up to 15% from those predicted using the standard compaction curve. Isostatic subsidence along the cross-section varies 2.3,4.6% between the two end-member cases. This leads to a 3,8% difference in tectonic subsidence, with maximum values in the basin centre. Owing to this, the estimated stretching factors vary up to 7.8%, which further gives rise to a maximum difference in heat flow of more than 8.5% in the basin centre. The modelled temperatures for an Upper Jurassic source rock show a deviation of more than 20 °C at present dependent on the thermal conductivity properties in the post-rift succession. This will influence the modelled hydrocarbon generation history of the basin, which is an essential output from basin modelling analysis. Results from the northern North Sea have shown that varying compaction trends in sediments with varying thermal properties are important parameters to constrain when analysing sedimentary basins. [source]


    Unravelling the multi-stage burial history of the Swiss Molasse Basin: integration of apatite fission track, vitrinite reflectance and biomarker isomerisation analysis

    BASIN RESEARCH, Issue 1 2006
    Martin Mazurek
    ABSTRACT A complex basin evolution was studied using various methods, including thermal constraints based on apatite fission-track (AFT) analysis, vitrinite reflectance (VR) and biomarker isomerisation, in addition to a detailed analysis of the regional stratigraphic record and of the lithological properties. The study indicates that (1) given the substantial amount of data, the distinction and characterisation of successive stages of heating and burial in the same area are feasible, and (2) the three thermal indicators (AFT, VR and biomarkers) yield internally consistent thermal histories, which supports the validity of the underlying kinetic algorithms and their applicability to natural basins. All data pertaining to burial and thermal evolution were integrated in a basin model, which provides constraints on the thickness of eroded sections and on heat flow over geologic time. Three stages of basin evolution occurred in northern Switzerland. The Permo-Carboniferous strike,slip basin was characterised by high geothermal gradients (80,100°C km,1) and maximum temperature up to 160°C. After the erosion of a few hundreds of metres in the Permian, the post-orogenic, epicontinental Mesozoic basin developed in Central Europe, with subsidence triggered by several stages of rifting. Geothermal gradients in northern Switzerland during Cretaceous burial were relatively high (35,40°C km,1), and maximum temperature typically reached 75°C (top middle Jurassic) to 100°C (base Mesozoic). At least in the early Cretaceous, a stage of increased heat flow is needed to explain the observed maturity level. After erosion of 600,700 m of Cretaceous and late Jurassic strata during the Paleocene, the wedge-shaped Molasse Foreland Basin developed. Geothermal gradients were low at this time (,20°C km,1). Maximum temperature of Miocene burial exceeded that of Cretaceous burial in proximal parts (<35 km from the Alpine front), but was lower in more distal parts (>45 km). Thus, maximum temperature as well as maximum burial depth ever reached in Mesozoic strata occurred at different times in different regions. Since the Miocene, 750,1050 m were eroded, a process that still continues in the proximal parts of the basin. Current average geothermal gradients in the uppermost 2500 m are elevated (32,47°C km,1). They are due to a Quaternary increase of heat flow, most probably triggered by limited advective heat transport along Paleozoic faults in the crystalline basement. [source]


    The stratigraphic and structural evolution of the Dzereg Basin, western Mongolia: clastic sedimentation, transpressional faulting and basin destruction in an intraplate, intracontinental setting

    BASIN RESEARCH, Issue 1 2003
    J. P. Howard
    ABSTRACT The Dzereg Basin is an actively evolving intracontinental basin in the Altai region of western Mongolia. The basin is sandwiched between two transpressional ranges, which occur at the termination zones of two regional-scale dextral strike-slip fault systems. The basin contains distinct Upper Mesozoic and Cenozoic stratigraphic sequences that are separated by an angular unconformity, which represents a regionally correlative peneplanation surface. Mesozoic strata are characterized by northwest and south,southeast-derived thick clast-supported conglomerates (Jurassic) overlain by fine-grained lacustrine and alluvial deposits containing few fluvial channels (Cretaceous). Cenozoic deposits consist of dominantly alluvial fan and fluvial sediments shed from adjacent mountain ranges during the Oligocene,Holocene. The basin is still receiving sediment today, but is actively deforming and closing. Outwardly propagating thrust faults bound the ranges, whereas within the basin, active folding and thrusting occurs within two marginal deforming belts. Consequently, active fan deposition has shifted towards the basin centre with time, and previously deposited sediment has been uplifted, eroded and redeposited, leading to complex facies architecture. The geometry of folds and faults within the basin and the distribution of Mesozoic sediments suggest that the basin formed as a series of extensional half-grabens in the Jurassic,Cretaceous which have been transpressionally reactivated by normal fault inversion in the Tertiary. Other clastic basins in the region may therefore also be inherited Mesozoic depocentres. The Dzereg Basin is a world class laboratory for studying competing processes of uplift, deformation, erosion, sedimentation and depocentre migration in an actively forming intracontinental transpressional basin. [source]


    Historical biogeography of Hexabathynella , a cosmopolitan genus of groundwater Syncarida (Crustacea, Bathynellacea, Parabathynellidae)

    BIOLOGICAL JOURNAL OF THE LINNEAN SOCIETY, Issue 4 2003
    ANA I. CAMACHO
    Hexabathynella is the only cosmopolitan genus of the order Bathynellacea (Crustacea). The known species number 18, found in Europe (9), Africa (1), South America (2), North America (3) and Australia and New Zealand (3). Phylogenetic analysis suggests that the least derived species are those from South America and the most derived those from the Iberian Peninsula, North America and Australia. The five species with the most plesiomorphic characters occur in salt or brackish water, which supports a marine origin for the genus. Phylogenetic and biogeographical analyses suggest that the distribution of the genus can be explained by dispersion and a double vicariant biogeographical model based on plate tectonics and the evolution of the Tethys during the Mesozoic and Cenozoic. © 2003 The Linnean Society of London . Biological Journal of the Linnean Society , 2003, 78 , 457,466. [source]


    Testing co-evolutionary hypotheses over geological timescales: interactions between Mesozoic non-avian dinosaurs and cycads

    BIOLOGICAL REVIEWS, Issue 1 2009
    Richard J. Butler
    Abstract The significance of co-evolution over ecological timescales is well established, yet it remains unclear to what extent co-evolutionary processes contribute to driving large-scale evolutionary and ecological changes over geological timescales. Some of the most intriguing and pervasive long-term co-evolutionary hypotheses relate to proposed interactions between herbivorous non-avian dinosaurs and Mesozoic plants, including cycads. Dinosaurs have been proposed as key dispersers of cycad seeds during the Mesozoic, and temporal variation in cycad diversity and abundance has been linked to dinosaur faunal changes. Here we assess the evidence for proposed hypotheses of trophic and evolutionary interactions between these two groups using diversity analyses, a new database of Cretaceous dinosaur and plant co-occurrence data, and a geographical information system (GIS) as a visualisation tool. Phylogenetic evidence suggests that the origins of several key biological properties of cycads (e.g. toxins, bright-coloured seeds) likely predated the origin of dinosaurs. Direct evidence of dinosaur,cycad interactions is lacking, but evidence from extant ecosystems suggests that dinosaurs may plausibly have acted as seed dispersers for cycads, although it is likely that other vertebrate groups (e.g. birds, early mammals) also played a role. Although the Late Triassic radiations of dinosaurs and cycads appear to have been approximately contemporaneous, few significant changes in dinosaur faunas coincide with the late Early Cretaceous cycad decline. No significant spatiotemporal associations between particular dinosaur groups and cycads can be identified , GIS visualisation reveals disparities between the spatiotemporal distributions of some dinosaur groups (e.g. sauropodomorphs) and cycads that are inconsistent with co-evolutionary hypotheses. The available data provide no unequivocal support for any of the proposed co-evolutionary interactions between cycads and herbivorous dinosaurs , diffuse co-evolutionary scenarios that are proposed to operate over geological timescales are plausible, but such hypotheses need to be firmly grounded on direct evidence of interaction and may be difficult to support given the patchiness of the fossil record. [source]


    Current Knowledge of Mesozoic Coleoptera from Daohugou and Liaoning (Northeast China)

    ACTA GEOLOGICA SINICA (ENGLISH EDITION), Issue 4 2010
    Alexander G. KIREJTSHUK
    Abstract: The present paper is devoted to an overview on fossil Coleoptera studied from Inner Mongolia, Daohugou (Middle Jurassic, Jiulongshan Formation) and Liaoning (Upper Jurassic-Lower Cretaceous, Yixian Formation) deposited in Chinese collections. As a result, species of the tribe Sperchopsini and Hydrophilini from Hydrophilidae, families and subfamilies Silphidae, Syndesinae from Lucanidae, Pleocomidae, Trogidae, Trogissitidae, Pyrochroidae, Diaperinae from Tenebrionidae, and Cerambycidae were first registered in the Mesozoic and some families were defined as new. It was shown that many superfamilies represented in the Recent Fauna were formed within the Middle Jurassic and Lower Cretaceous. The materials examined confirm the hypothesis that Cucujiformian beetles are a younger group than other infraordera of Polyphaga (Staphyliniformia and Elateriformia) and, therefore, they appeared in the fossil record only in the late Mesozoic. It was shown and confirmed that most superfamilies appeared in the fossil records before Cucujoidea. The synonymy of Notocupes Ponomarenko, 1964; Sinocupes Lin, 1976, syn. nov.; Amblomma Tan, Ren et Liu 2005, syn. nov.; Euryomma Tan, Ren et Shih, 2006, syn. nov., non Stein, 1899 and Ovatocupes Tan et Ren, 2006, syn. nov.; synonymy of Tetraphalerus Waterhouse, 1901 and Odontomma Tan, Ren et Ge 2006, syn. nov.; and synonymy of Priacmopsis Ponomarenko, 1966 and Latocupes Tan et Ren, 2006, syn. nov. are proposed. Sinorhombocoleus papposus Tan et Ren, 2009 is transferred from the family Rhombocoleidae to Schizophoridae. Cervicatinius complanus Tan, Ren et Shih, 2007 and Forticatinius elegans Tan, Ren et Shih, 2007 are transferred from the family Catiniidae (suborder Archostemata) to superfamily Cleroidea (suborder Polyphaga: first among the family Peltidae and second as a closely related group to the latter family). The family Parandrexidae is transferred from the superfamily Cucujoidea to Cleroidea. The ecological circumstances of the past ecosystems and hypotheses of historical development of the order Coleoptera are discussed. The age of faunas examined is considered. The list of the taxa described from Daohugou and Liaoning is compiled. [source]


    Cockroach as the Earliest Eusocial Animal

    ACTA GEOLOGICA SINICA (ENGLISH EDITION), Issue 4 2010
    ANSKÝ, Peter VR
    Abstract: A completely preserved cockroach Sociala perlucida gen. et sp. nov. (Blattida: Socialidae fam. nov.) is described from the Mesozoic (Albian Early Cretaceous) amber of Archingeay in France. It is categorized within the new family Socialidae, originating from the Mesozoic cockroach family Liberiblattinidae, and representing the sister group to the most basal known eutermite families. Numerous direct and indirect morphological evidence (such as a unique narrow body and pronotum with reduced coloration, legs without carination and with numerous sensillar pitts (forelegs) as adaptations to life in nests analogical to the living termitophillous species; enlarged head, long palps for communication, a general venation modified in a direction towards termites (subcosta and radial vein [R] approximated, R branches approximated, simplified, cross-veins reduced), and possibly also the development of the breaking forewing sutura for detaching wings after marriage flight), suggest its eusocial mode of life. Thus, the first eusociality originated within cockroaches, prior to giving birth to termites; termites and Cryptocercus are not directly related. The group of eusocial cockroaches makes termite morphotaxon polyphyletic, but also in this case, the erection of a new order for mastotermites (including Cratomastotermitidae) and/or the inclusion of eusocial, morphological cockroaches within termites appears counterproductive, thus an example of such a general exception to the taxonomical procedure is provided. [source]