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Lower Devonian (lower + devonian)
Selected AbstractsTHE EXTRAORDINARY TRILOBITE FENESTRASPIS (DALMANITIDAE, SYNPHORIINAE) FROM THE LOWER DEVONIAN OF BOLIVIAPALAEONTOLOGY, Issue 4 2009DAVID J. HOLLOWAY Abstract: The hitherto poorly known, monotypic trilobite genus Fenestraspis from the Lower Devonian of Bolivia is revised and its original assignment to the Synphoriinae supported. The thoracic morphology of the genus remains very poorly known. Fenestraspis is morphologically unusual because of the development of extensive fenestrae in the pleural region of the pygidium and apparently of the thorax; the presence of upwardly directed spines on the cephalon, thorax and pygidium; and the exceptionally large and highly elevated eyes with the palpebral rim projecting outwards above the visual surface. The function of the fenestrae remains uncertain. If they formed openings in the body of the trilobite in life they may have allowed circulation of oxygenated water to the limb exites so that respiration could have been maintained while the trilobite was enrolled. If they were covered with a flexible membrane, they may have been secondary respiratory structures or had a sensory function. The Synphoriinae is regarded as a subfamily of the Dalmanitidae rather than as an independent family of the Dalmanitoidea as proposed by some authors. The type species of the poorly known monotypic genus Dalmanitoides from the Lower Devonian of Argentina is illustrated photographically for the first time and compared with Fenestraspis. [source] THE GENUS GIGANTASPIS HEINTZ, 1962 (VERTEBRATA, HETEROSTRACI) FROM THE LOWER DEVONIAN OF SPITSBERGENPALAEONTOLOGY, Issue 2 2007VINCENT PERNEGRE Abstract:, Material collected in 1969 by the CNRS-MNHN expedition to Spitsbergen includes a new species of Gigantaspis, G. minima, which is the smallest known species of this genus. The revision of Gigantaspis leads to the inclusion of Zascinaspis laticephala Blieck and Goujet in this genus. The phylogenetic analysis presented herein suggests that Gigantaspis is close to Zascinaspis, as suggested by Blieck. Moreover, it is also close to generalized representatives of the Protaspididae. Their shared character states allow the definition of a possible ancestral morphotype for the Protaspididae. [source] A NEW SPECIES OF BRITTLESTAR (OPHIUROIDEA, ECHINODERMATA) FROM THE HUNSRÜCK SLATE (LOWER EMSIAN, LOWER DEVONIAN) OF GERMANYPALAEONTOLOGY, Issue 5 2006ALEXANDER GLASS Abstract:,Lapworthura lehmanni, a new species of ophiuroid, is described from four specimens from the Lower Devonian Hunsrück Slate of Germany. It is the only known ophiuroid in the Hunsrück Slate with paired but unfused ambulacrals, and it exhibits unique rows of spine-bearing dorsal arm ossicles. The genus Lapworthura Gregory was previously known only from the Ordovician of Scotland and the Silurian of England and Australia. [source] Enameloid microstructure in the oldest known chondrichthyan teethACTA ZOOLOGICA, Issue 2009H. Botella Abstract Previous studies on tooth enameloid microstructure in several chondrichthyan taxa spanning the phylogeny of the group provided support for the homology of chondrichthyan tooth enameloid. This hypothesis requires that a single crystallite enameloid (SCE) monolayer must be present in the teeth of the most primitive chondrichthyan. However, the dental microstructure of the earliest sharks has yet to be investigated. We have studied the tooth enameloid microstructure of the two oldest tooth-bearing shark species currently known, Leonodus carlsi Mader (1986) and Celtiberina maderi Wang (1993), from the lowermost Lochkovian (Lower Devonian) of Spain. Our study demonstrates the presence of a SCE monolayer in the teeth of both species. These results show that a superficial cap of SCE is present in the oldest shark teeth known, which suggest its presence in the most basal chondrichthyans. [source] The braincase of the chondrichthyan Doliodus from the Lower Devonian Campbellton Formation of New Brunswick, CanadaACTA ZOOLOGICA, Issue 2009John Maisey Abstract The braincase of the late Lower Devonian (Emsian) chondrichthyan Doliodus is described for the first time. Its postorbital process is extended ventrally and probably enclosed part of the infraorbital sensory canal, as in some placoderms. Doliodus has a shark-like dentition, but its upper anterior tooth files were supported by the internasal cartilage of the braincase, not by the palatoquadrates. Modern selachian jaws and dentitions are not representative of primitive crown-group gnathostomes because they display a mixture of conserved and derived character states. Separation of the palatoquadrates by the internasal cartilage is probably a primitive condition for crown-group gnathostomes. Continuity of the upper dental arcade across the ethmoid region may represent a synapomorphy of chondrichthyans and some acanthodians (the condition is not found in placoderms or osteichthyans). Exclusion of the arcade from the ethmoid region is probably apomorphic within elasmobranchs. Doliodus has curious bar-like, paired subcranial ridges ending posteriorly at the hyomandibular articulation. These superficially resemble visceral arch infrapharyngohyals fused to the floor of the braincase, adding circumstantial palaeontological support to the old proposal that parts of visceral arches may be incorporated into the gnathostome braincase, although it seems more plausible that they formed in the lateral margins of the embryonic parachordal or hypotic lamina. [source] Exceptionally preserved conulariids and an edrioasteroid from the Hunsrück Slate (Lower Devonian, SW Germany)PALAEONTOLOGY, Issue 2 2010HEYO VAN ITEN Abstract:, Nineteen partial specimens of Conularia sp., together with an articulated agelacrinitid edrioasteroid and several discinid brachiopods, occur in close association with a probable biological substrate on a small slab of silty Hunsrück Slate (Lower Devonian, Emsian) from Bundenbach, Germany. Most of the conulariids occur in V-like pairs or in a single cluster of 12 specimens arranged in a fan-like radial pattern. Together with the edrioasteroid and (possibly) brachiopods, the conulariids probably were attached to the substrate in life and then were buried and possibly killed by a single influx of silty mud. The apertural end of many of the conulariids is partially covered by inwardly folded short lappets, which may have closed in response to rapid (but gentle) burial. Rock matrix in the apertural region of the peridermal cavity of nearly all of the conulariids exhibits irregular, variably dense concentrations of pyrite. The concentrations occur almost exclusively within the conulariids, where they probably formed as a result of the decay of retracted conulariid soft parts. Although the concentrations lack clearly defined anatomical features that can be unambiguously homologized with particular anatomical structures of any extant taxon, their form and distribution within the conulariids are consistent with the hypothesis that conulariids were polypoid scyphozoans. [source] THE EXTRAORDINARY TRILOBITE FENESTRASPIS (DALMANITIDAE, SYNPHORIINAE) FROM THE LOWER DEVONIAN OF BOLIVIAPALAEONTOLOGY, Issue 4 2009DAVID J. HOLLOWAY Abstract: The hitherto poorly known, monotypic trilobite genus Fenestraspis from the Lower Devonian of Bolivia is revised and its original assignment to the Synphoriinae supported. The thoracic morphology of the genus remains very poorly known. Fenestraspis is morphologically unusual because of the development of extensive fenestrae in the pleural region of the pygidium and apparently of the thorax; the presence of upwardly directed spines on the cephalon, thorax and pygidium; and the exceptionally large and highly elevated eyes with the palpebral rim projecting outwards above the visual surface. The function of the fenestrae remains uncertain. If they formed openings in the body of the trilobite in life they may have allowed circulation of oxygenated water to the limb exites so that respiration could have been maintained while the trilobite was enrolled. If they were covered with a flexible membrane, they may have been secondary respiratory structures or had a sensory function. The Synphoriinae is regarded as a subfamily of the Dalmanitidae rather than as an independent family of the Dalmanitoidea as proposed by some authors. The type species of the poorly known monotypic genus Dalmanitoides from the Lower Devonian of Argentina is illustrated photographically for the first time and compared with Fenestraspis. [source] DEVONIAN TRILOBITES FROM THE FALKLAND ISLANDSPALAEONTOLOGY, Issue 1 2006MARIA DA GLORIA PIRES DE CARVALHO Abstract:, New trilobite material is described from the Fox Bay Formation (Lower Devonian; Pragian) of the Falkland Islands (South Atlantic). The fossils were collected from many localities in both East and West Falkland and represent the first significant collection of trilobites from East Falkland. The trilobites belong to three higher taxa: Proetida (the first proetid record in these islands); Homalonotidae, Burmeisteria Salter, 1865; and Calmoniidae, including Bainella Rennie, 1930 and Metacryphaeus Reed, 1907. All of the forms except proetids have now been found in both East and West Falkland. [source] Hsüa deflexa sp. nov. from the Xujiachong Formation (Lower Devonian) of eastern Yunnan, ChinaBOTANICAL JOURNAL OF THE LINNEAN SOCIETY, Issue 3 2003DE-MING WANG A new vascular plant, Hsüa deflexa sp. nov., is documented from the Lower Devonian ((upper) Pragian-lower Emsian) Xujiachong Formation, near Xujiachong village, Qujing District, eastern Yunnan, South China. In three dimensions, the branching system comprises a robust creeping main axis and comparatively slender erect lateral axes inserted oppositely or alternately. The lateral axes depart at right angles from the main axis. Towards the apex, the main axis is perhaps erect with the lateral axes attached at smaller angles. The lateral axes dichotomize equally one to three times in different planes and decrease in diameter and length acropetally. They bear sparse and irregular tiny spines. Apices of a pair of distal lateral axes curve in opposite directions and each terminates in a rounded to reniform sporangium. The sporangium dehisces into two equal valves along an indistinct convex marginal thickening. The xylem is possibly centrarch. This plant fits Hsüa in branching pattern, sporangial characters and xylem maturation. Hsüa deflexa sp. nov. differs mainly from the type species H. robusta in the presence of axial spines, perpendicular extension of lateral axes from the main axis and curving of distal lateral axes. Based on the possibly centrarch xylem and terminal sporangium, this plant relates to the rhyniophytes (sensu Banks, 1968). In view of the equal sporangial valves with marginal thickenings it resembles the zosterophyllophytes (sensu Banks, 1968). Hsüa is now treated as incertae sedis. © 2003 The Linnean Society of London, Botanical Journal of the Linnean Society, 2003, 142, 255,271. [source] Diversity in conducting cells in early land plants and comparisons with extant bryophytesBOTANICAL JOURNAL OF THE LINNEAN SOCIETY, Issue 3 2003D. EDWARDS Anatomical screening using scanning electron microscopy (SEM) of short lengths of smooth coalified axes (mesofossils) from a Lochkovian (Lower Devonian) locality in the Welsh Borderland, Shropshire has revealed extensive diversity in the architecture of centrally aggregated, elongate cells. At least 14 types have been discovered, each distinguished by variation in wall architecture and combination of the cells in the central strand. End walls have not been seen. These elongate cells may have smooth, uniformly thick or thin walls, walls with smooth projections either traversing or lining the lumen, or bilayered walls, the innermost perforated by pores of plasmodesmata dimensions. The latter type may be further divided on presence or absence of projections which may line the lumen, but usually cross it and are highly disorganized. Indeed, none of the cells shows the regularity associated with the secondary thickenings of tracheids, but the imperforate/pitted forms with projections superficially resemble the S-type tracheids of the Rhyniopsida in basic construction. Simply pitted types show greater similarity with the water-conducting cells (WCCs) of liverworts and Takakia. To facilitate direct comparison with bryophyte conducting elements, SEM studies were undertaken on the WCCs of a number of mosses and liverworts and on the leptoids of mosses, in conjunction with a range of degradation experiments designed to assess the fossilization potential of these cells. With the exception of polytrichaceous hydroids, the latter demonstrated the resilience of hydroids and leptoids to the chemical treatments. In addition, dehydration of the leptoids produced globular residues similar to those seen in some of the fossils. This combination of techniques raises the possibility that food-conducting cells might well be preserved in coalified fossils, and hence extends the interpretation of the functions of the elongate cells. Broadly speaking, imperforate bilayered examples may have been involved in water conduction, cells with globular residues with or without pitting involved in metabolite movement, and smooth walled examples with or without projections involved in support. The wider affinities of the plants which produced the axes remain equivocal and in the absence of sporangia it is impossible to assign them to a genus. However, this anatomical diversity in vegetative remains of extreme simplicity demonstrates far greater diversity in early land vegetation than is apparent from perusal of species lists. © 2003 The Linnean Society of London, Botanical Journal of the Linnean Society, 2003, 141, 297,347. [source] New perspectives on Cooksonia from the Lower Devonian of the Welsh BorderlandBOTANICAL JOURNAL OF THE LINNEAN SOCIETY, Issue 4 2002K. S. HABGOOD New collections of coalified mesofossils of Cooksonia have been made from a Lochkovian (Lower Devonian) locality in Shropshire, England. They allow detailed description of sporangial anatomy in Cooksonia pertoni ssp. apiculispora with inferences for spore dispersal involving disintegration of the distal sporangial wall and maintenance of a rigid rim. Trilete spores, Aneurospora sp. and Streelispora newportensis have been recovered from the same sporangium. A new subspecies Cooksonia pertoni subsp reticulispora contains miospores with laevigate proximal surface bearing a simple triradiate apertural fold, equatorial crassitude and a coarse reticulum of muri on a slightly thickened distal wall. Spores are tentatively assigned to Synorisporites sp.. Cooksonia banksiisp. nov. is based on terminal sporangia in which the spore-containing area is enclosed within the expanded apex of the subtending axis. The cavity is lined by a smooth resilient acellular layer. Well-defined cells are apparent only in the pronounced sporangial rim. Spores are assigned to Ambitisporites avitus, a taxon of particular significance, because it is recorded in the Upper Ordovician and Lower Silurian assemblages which contain the earliest trilete monads. However, it is concluded that spores of such simple morphology may have been produced by a variety of taxa and that ultrastructural studies are necessary to confirm this possibility. These new specimens thus allow further insights into the disparity (via anatomy and in situ spores) of very simple plants and, in providing information on cell construction and arrangement, allow for functional interpretations, particularly relating to spore dispersal. © 2002 The Linnean Society of London, Botanical Journal of the Linnean Society, 2002, 139, 339,359. [source] Two Zosterophyll Plants from the Lower Devonian (Lochkovian) Xitun Formation of Northeastern Yunnan, ChinaACTA GEOLOGICA SINICA (ENGLISH EDITION), Issue 3 2009Jinzhuang XUE Abstract: Two zosterophyll plants are described from the Lower Devonian (Lochkovian) Xitun Formation of Qujing, Yunnan, China. Xitunia spinitheca gen. et sp. nov. has stalked sporangia laterally attached on the axis in a helical arrangement. Sporangia are dorsoventrally flattened and composed of two unequal valves; the adaxial valve is round in face view, while the abaxial valve is larger than the former, triangular or wedge-shaped, and radially bears long spiny appendages along the distal margin. Xitunia shows new variation of sporangial morphology within the zosterophylls. Zosterophyllum minorstachyum sp. nov. has K-shaped branchings at the basal parts and small-sized terminal spikes, which consist of round to elliptical sporangia arranged helically. This paper provides new data on the diversity of plant types during Lochkovian when rare vascular plants were reported. As for various species of Zosterophyllum in South China, their apparent evolutionary trend of features from the Late Silurian to Early Devonian (Emsian) is discussed. [source] | |||||||||||||