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Island Size (island + size)
Selected AbstractsColonization success of carabid beetles on Baltic islandsJOURNAL OF BIOGEOGRAPHY, Issue 4 2000D. Johan Kotze Abstract Aims (1) To test whether there is a significant increase in carabid species richness with an increase in island size and, if so, if it is due to island area per se or habitat diversity. (2) To investigate whether scattered islands accumulate species quicker than islands close to each other, per island size. (3) To investigate changes in the proportions of carabid wing morphs between the Finnish mainland and islands in the Baltic Sea. Location Islands in the south-western archipelago of Finland, in the Baltic Sea. Methods Carabid beetles were collected using pitfall traps (diameter, 65 mm; volume, 170 mL), half-filled with an ethylene-glycol,water mixture, from 22 May to 20 September 1993 on 24 islands. Island size varied between 0.5 and c. 7000 ha, and each island had between one and four habitat types sampled. Results A total of 61 carabid species were captured on these islands. Pterostichus niger was numerically dominant on 15 of the 24 islands and made up 34.5% of the total catch. The islands had a significantly higher proportion of brachypterous species compared to the Finnish mainland. The islands also accumulated species at a much slower rate (z = 0.06) than that generally observed in the literature, and, for carabids, a mainly predacious group, habitat diversity had little predictive power in explaining species richness. Islands close to each other (a few hundred metres apart) accumulated species at a slower rate than did scattered islands, as island size increased. Main conclusions Although carabids disperse relatively easily to remote islands (perhaps a result of low Baltic Sea salinity and short interisland distances), colonization success appears to depend on a multitude of factors, including availability of suitable habitat on these islands, competitive superiority, survival ability during dispersal and island arrival sequence. [source] Patterns of bird species richness and composition on islands off Arnhem Land, Northern Territory, AustraliaAUSTRAL ECOLOGY, Issue 1 2001J. C. Z. Woinarski Abstract The bird faunas of the adjacent Wessel and English Company island chains were sampled at two scales (0.25 ha quadrats and entire islands). Ninety-six species were recorded from 226 quadrats, with the most frequently recorded species being mistletoebird Dicaeum hirundinaceum, brown honeyeater Lichmera indistincta, silver-crowned friarbird Philemon argenticeps, bar-shouldered dove Geopelia humeralis, northern fantail Rhipidura rufiventris and yellow white-eye Zosterops lutea. At the quadrat scale, vegetation type was a major determinant of the abundance of individual species (and hence species composition), species richness and total bird abundance. Bird species composition and richness at the quadrat scale was also significantly affected by island isolation (particularly the amount of land within 20 km of the island perimeter). Island size had no effect on quadrat-scale richness or total abundance. However, the abundance of 10 of the 38 most frequently recorded individual species was significantly related to island size, in most cases even when the comparison was restricted to similar habitats. The most striking cases were rufous fantail Rhipidura rufifrons, mangrove golden whistler Pachycephala melanura, brown honeyeater and yellow white-eye, which were all significantly more abundant on smaller islands. One hundred and seventy-one species were recorded from the 62 islands sampled. There was a very tight relationship between island size and the number of terrestrial species (73% of deviance explained) and of all species (84% of deviance explained). This relationship was improved (marginally) when isolation was included in the model. Ordination of islands by their terrestrial bird species composition was related to island size and isolation, and suggested an erratic species composition on small islands. [source] Minimizing energy expenditure facilitates vertebrate persistence on oceanic islandsECOLOGY LETTERS, Issue 5 2002Brian K. McNab Abstract The characteristics of terrestrial vertebrates on oceanic islands are examined. They often include a reduced body size, a tolerance of conspecifics, flightlessness, a reduced basal rate of metabolism, and a propensity to enter torpor. On oceanic islands ectotherms frequently replace endotherms. These changes reduce the energy expenditure and resource requirements of vertebrates. Such reductions are permitted by the absence of mammalian predators and facilitate the survival of island endemics in the face of a restricted resource base and a variable environment through an increase in population size. Some insular species increase body size, but this occurs only when the resource base is large, due either to a fortuitously abundant resource, or to the absence of other species that exploit normally abundant resources. Some questions are posed to guide future work. They examine of the characteristics that permit species to disperse over water barriers, the conditions that require a reduction in resource use, the rapidity of response by immigrants to island conditions, whether supertramps show physiological differentiation with respect to island distance or size, and whether island size is absolute or relative to the size of the immigrants. [source] Breaking taboos in the tropics: incest promotes colonization by wood-boring beetlesGLOBAL ECOLOGY, Issue 4 2001Bjarte H. Jordal Abstract 1,Inbreeding and parthenogenesis are especially frequent in colonizing species of plants and animals, and inbreeding in wood-boring species in the weevil families Scolytinae and Platypodidae is especially common on small islands. In order to study the relationship between colonization success, island attributes and mating system in these beetles, we analysed the relative proportions of inbreeders and outbreeders for 45 Pacific and Old World tropical islands plus two adjacent mainland sites, and scored islands for size, distance from nearest source population, and maximum altitude. 2,The numbers of wood-borer species decreased with decreasing island size, as expected; the degree of isolation and maximum island altitude had negligible effects on total species numbers. 3,Numbers of outbreeding species decreased more rapidly with island size than did those of inbreeders. Comparing species with similar ecology (e.g. ambrosia beetles) showed that this difference was best explained by differential success in colonization, rather than by differences in resource utilization or sampling biases. This conclusion was further supported by analyses of data from small islands, which suggested that outbreeding species have a higher degree of endemism and that inbreeding species are generally more widespread. 4,Recently established small populations necessarily go through a period of severe inbreeding, which should affect inbreeding species much less than outbreeding ones. In addition, non-genetic ecological and behavioural (,Allee') effects are also expected to reduce the success of outbreeding colonists much more than that of inbreeders: compared with inbreeders, outbreeders are expected to have slower growth rates, have greater difficulties with mate-location and be vulnerable to random extinction over a longer period. [source] Colonization success of carabid beetles on Baltic islandsJOURNAL OF BIOGEOGRAPHY, Issue 4 2000D. Johan Kotze Abstract Aims (1) To test whether there is a significant increase in carabid species richness with an increase in island size and, if so, if it is due to island area per se or habitat diversity. (2) To investigate whether scattered islands accumulate species quicker than islands close to each other, per island size. (3) To investigate changes in the proportions of carabid wing morphs between the Finnish mainland and islands in the Baltic Sea. Location Islands in the south-western archipelago of Finland, in the Baltic Sea. Methods Carabid beetles were collected using pitfall traps (diameter, 65 mm; volume, 170 mL), half-filled with an ethylene-glycol,water mixture, from 22 May to 20 September 1993 on 24 islands. Island size varied between 0.5 and c. 7000 ha, and each island had between one and four habitat types sampled. Results A total of 61 carabid species were captured on these islands. Pterostichus niger was numerically dominant on 15 of the 24 islands and made up 34.5% of the total catch. The islands had a significantly higher proportion of brachypterous species compared to the Finnish mainland. The islands also accumulated species at a much slower rate (z = 0.06) than that generally observed in the literature, and, for carabids, a mainly predacious group, habitat diversity had little predictive power in explaining species richness. Islands close to each other (a few hundred metres apart) accumulated species at a slower rate than did scattered islands, as island size increased. Main conclusions Although carabids disperse relatively easily to remote islands (perhaps a result of low Baltic Sea salinity and short interisland distances), colonization success appears to depend on a multitude of factors, including availability of suitable habitat on these islands, competitive superiority, survival ability during dispersal and island arrival sequence. [source] Analysis of plant species diversity with respect to island characteristics on the Channel Islands, CaliforniaJOURNAL OF BIOGEOGRAPHY, Issue 3 2000Aaron Moody Abstract Aim Species richness of native, endemic, and exotic plant groups is examined relative to island area, disturbance history, geological history, and other physical characteristics. Of particular interest are the biogeographic factors that underlie (a) differences in species-area and species-isolation relationships between plant groups; and (b) adherence or departure of individual islands and/or plant groups from expected patterns. Location The eight Channel Islands lie along the continental margin between the U.S./Mexico border and Point Conception, CA. They range in size from 2.6 to 249 km2, and are located from 20 to 100 km off the coast. The islands are known for their high degree of plant endemism, and they have undergone a long history of human occupation by indigenous peoples, followed by over a century of intensive grazing and other biotic disturbances. Methods The study is based on linear regression and residual analysis. Cases where individual islands and/or specific plant groups do not adhere to patterns expected under species-area and species-isolation paradigms, are evaluated with respect to other island characteristics that are not captured by considering only island size and isolation. Results All three plant groups exhibit strong, positive relationships between species richness and island size. For native species, the variance that remains after consideration of island size is largely explained by island isolation. For exotic species, residuals from the species-area relationship are unrelated to isolation. For endemic species, residuals from the species-area relationship are negatively related to isolation. Several islands are outliers for endemic and exotic species, for which richness values are not explained by either island area or isolation. Main,conclusions Species-area and species-isolation relationships for native, endemic, and exotic plant groups differ in accordance with hypothesized differences in the biogeographic factors that govern species diversity for these three groups. Most notably, endemic richness increases with isolation, suggesting the influence of this variable on processes of speciation and relictualism. These general relationships persist despite a long and varied history of human activity on the islands. Analysis of residuals suggests that deviations from expected patterns correspond to island-specific biogeographic factors. It is hypothesized that primary among these factors are land-use history, island environmental characteristics, and community-type richness. [source] Genetic diversity and population size: island populations of the common shrew, Sorex araneusMOLECULAR ECOLOGY, Issue 10 2007THOMAS A. WHITE Abstract Populations of many species are currently being fragmented and reduced by human interactions. These processes will tend to reduce genetic diversity within populations and reduce individual heterozygosities because of genetic drift, inbreeding and reduced migration. Conservation biologists need to know the effect of population size on genetic diversity, as this is likely to influence a population's ability to persist. Island populations represent an ideal natural experiment with which to study this problem. In a study of common shrews (Sorex araneus) on offshore Scottish islands, 497 individuals from 13 islands of different sizes and 6 regions on the mainland were trapped and genotyped at eight microsatellite loci. Previous genetic work had revealed that most of the islands in this study were highly genetically divergent from one another and the mainland. We found that most of the islands exhibited lower genetic diversity than the mainland populations. In the island populations, mean expected heterozygosity, mean observed heterozygosity and mean allelic richness were significantly positively correlated with log island size and log population size, which were estimated using habitat population density data and application of a Geographic Information System. [source] Scale as a lurking factor: incorporating scale-dependence in experimental ecologyOIKOS, Issue 9 2009Brody Sandel Ecologists have recognized for decades the importance of spatial scale in ecological processes and patterns, as well as the complications scale poses for understanding ecological mechanisms. Here we highlight the opportunity attention to scale offers experimental ecology. Despite many advantages to considering scale, a review of the literature indicates that multi-scale experimental studies are rare. Although much work has focused on scale as a primary factor (e.g. island size), we draw attention to scale as a ,lurking' variable: one which influences the relationship between two or more variables that are not usually understood to be scale-dependent. We highlight three basic observations from which scale-dependence arises: abundance increases with area, environmental conditions vary across space, and the effect of an organism on its environment is spatially limited. From these arise first-order scale-dependence, which relates an ecological variable of interest to a measure of scale. Combining first-order relationships together, we can produce second-order scale-dependencies, which occur when the relationship between two or more variables is mediated by scale. It is these relationships that are of particular interest, as they have the potential to confound experimental results. Most ecological experiments have incorporated scale either implicitly or not at all. We suggest that an explicit consideration of scale could help resolve some long-standing debates when scale is turned from a lurking variable into a working variable. Finally, we review and evaluate four different experimental sampling designs and corresponding statistical analyses that can be used to address the effects of scale in ecological experiments. [source] Kinetically controlled engineering of quantum dot arraysPHYSICA STATUS SOLIDI (B) BASIC SOLID STATE PHYSICS, Issue 2 2003V. G. Dubrovskii Abstract A kinetic model for the formation of arrays of coherent strained islands is developed. The derived expressions provide a detailed characterization of quantum dot arrays in terms of island size and density as functions of substrate temperature, growth rate, exposition time and the amount of deposit. The obtained results give a guideline to the kinetically controlled engineering of quantum dot arrays with required structural properties. [source] Patterns of bird species richness and composition on islands off Arnhem Land, Northern Territory, AustraliaAUSTRAL ECOLOGY, Issue 1 2001J. C. Z. Woinarski Abstract The bird faunas of the adjacent Wessel and English Company island chains were sampled at two scales (0.25 ha quadrats and entire islands). Ninety-six species were recorded from 226 quadrats, with the most frequently recorded species being mistletoebird Dicaeum hirundinaceum, brown honeyeater Lichmera indistincta, silver-crowned friarbird Philemon argenticeps, bar-shouldered dove Geopelia humeralis, northern fantail Rhipidura rufiventris and yellow white-eye Zosterops lutea. At the quadrat scale, vegetation type was a major determinant of the abundance of individual species (and hence species composition), species richness and total bird abundance. Bird species composition and richness at the quadrat scale was also significantly affected by island isolation (particularly the amount of land within 20 km of the island perimeter). Island size had no effect on quadrat-scale richness or total abundance. However, the abundance of 10 of the 38 most frequently recorded individual species was significantly related to island size, in most cases even when the comparison was restricted to similar habitats. The most striking cases were rufous fantail Rhipidura rufifrons, mangrove golden whistler Pachycephala melanura, brown honeyeater and yellow white-eye, which were all significantly more abundant on smaller islands. One hundred and seventy-one species were recorded from the 62 islands sampled. There was a very tight relationship between island size and the number of terrestrial species (73% of deviance explained) and of all species (84% of deviance explained). This relationship was improved (marginally) when isolation was included in the model. Ordination of islands by their terrestrial bird species composition was related to island size and isolation, and suggested an erratic species composition on small islands. [source] Gene flow and melanism in garter snakes revisited: a comparison of molecular markers and island vs. coalescent modelsBIOLOGICAL JOURNAL OF THE LINNEAN SOCIETY, Issue 3 2003TONYA D. BITTNER Within populations, the stochastic effect of genetic drift and deterministic effect of natural selection are potentially weakened or altered by gene flow among populations. The influence of gene flow on Lake Erie populations of the common garter snake has been of particular interest because of a discontinuous colour pattern polymorphism (striped vs. melanistic) that is a target of natural selection. We reassessed the relative contributions of gene flow and genetic drift using genetic data and population size estimates. We compared all combinations of two marker systems and two analytical approaches to the estimation of gene flow rates: allozymes (data previously published), microsatellite DNA (new data), the island model (FST -based approach), and a coalescence-based approach. For the coalescence approach, mutation rates and sampling effects were also investigated. While the two markers produced similar results, gene flow based on FST was considerably higher (Nm > 4) than that from the coalescence-based method (Nm < 1). Estimates of gene flow are likely to be inflated by lack of migration-drift equilibrium and changing population size. Potentially low rates of gene flow (Nm < 1), small population size at some sites, and positive correlations of number of microsatellite DNA alleles and island size and between M, mean ratio of number of alleles to range in allele size, and island size suggest that in addition to selection, random genetic drift may influence colour pattern frequencies. © 2003 The Linnean Society of London, Biological Journal of the Linnean Society, 2003, 79, 389,399. [source] Floristic biogeography of the Hawaiian Islands: influences of area, environment and paleogeographyJOURNAL OF BIOGEOGRAPHY, Issue 3 2004Jonathan P. Price Abstract Aim, A detailed database of distributions and phylogenetic relationships of native Hawaiian flowering plant species is used to weigh the relative influences of environmental and historical factors on species numbers and endemism. Location, The Hawaiian Islands are isolated in the North Pacific Ocean nearly 4000 km from the nearest continent and nearly as distant from the closest high islands, the Marquesas. The range of island sizes, environments, and geological histories within an extremely isolated archipelago make the Hawaiian Islands an ideal system in which to study spatial variation in species distributions and diversity. Because the biota is derived from colonization followed by extensive speciation, the role of evolution in shaping the regional species assemblage can be readily examined. Methods, For whole islands and regions of each major habitat, species,area relationships were assessed. Residuals of species,area relationships were subjected to correlation analysis with measures of endemism, isolation, elevation and island age. Putative groups of descendents of each colonist from outside the Hawaiian Islands were considered phylogenetic lineages whose distributions were included in analyses. Results, The species,area relationship is a prominent pattern among islands and among regions of each given habitat. Species number in each case correlates positively with number of endemics, number of lineages and number of species per lineage. For mesic and wet habitat regions, island age is more influential than area on species numbers, with older islands having more species, more single-island endemics, and higher species : lineage ratios than their areas alone would predict. Main conclusions, Because species numbers and endemism are closely tied to speciation in the Hawaiian flora, particularly in the most species-rich phylogenetic lineages, individual islands' histories are central in shaping their biota. The Maui Nui complex of islands (Maui, Moloka,i, L,na,i and Kaho,olawe), which formed a single large landmass during most of its history, is best viewed in terms of either the age or area of the complex as a whole, rather than the individual islands existing today. [source] Biological images of geological history: through a glass darkly or brightly face to face?JOURNAL OF BIOGEOGRAPHY, Issue 2 2003Jeremy D. Holloway Abstract Aim, To explore the implications for historical biogeography of a recent review of island biogeographical theory in three main thematic areas and to suggest ways in which a synthesis between the two approaches might be achieved to the benefit of both. Location, The Indo-Australian tropics. Theme 1, discusses the relationship of species number to area, and how the nestedness of faunas may influence the methodology used for some types of analysis and also the quality of data expected from an archipelago embracing an extreme range of island sizes. Theme 2, examines the way in which the processes of speciation may lead to development of biogeographical patterns through a complex archipelago, illustrated in particular with reference to Sulawesi where biotic enrichment from different lepidopteran groups follows predictions from island biogeographical theory. This also has implications for patterns of endemism in the archipelago, another constraint on the quality of data available for historical biogeography. Theme 3, addresses ecological determinism as an influence in development of biogeographical pattern, focusing on the theme of specificity in insect,plant relationships and the potential for parallel development of pattern in an insect group and its particular plant host group. This theme is developed with particular reference to moth and plant groups that may represent Gondwanan elements in the Oriental fauna, with an analysis of Sarcinodes, a geometrid moth genus associated with Proteaceae. Main conclusions, Prospects are assessed for the synthesis of the two approaches of island biogeography and historical biogeography. Modelling pattern development with the former may complement the methods of analysis of the latter, particularly if some satisfactory method for dating events of pattern development can also be incorporated. [source] | ||||||||||||||||||||||