Home About us Contact
|
Home About us Contact | ||
|
|
||
Heterogeneous Environments (heterogeneous + environment)
Selected AbstractsSELECTION IN HETEROGENEOUS ENVIRONMENTS MAINTAINS THE GENE ARRANGEMENT POLYMORPHISM OF DROSOPHILA PSEUDOOBSCURAEVOLUTION, Issue 12 2008Stephen W. Schaeffer Chromosomal rearrangements may play an important role in how populations adapt to a local environment. The gene arrangement polymorphism on the third chromosome of Drosophila pseudoobscura is a model system to help determine the role that inversions play in the evolution of this species. The gene arrangements are the likely target of strong selection because they form classical clines across diverse geographic habitats, they cycle in frequency over seasons, and they form stable equilibria in population cages. A numerical approach was developed to estimate the fitness sets for 15 gene arrangement karyotypes in six niches based on a model of selection,migration balance. Gene arrangement frequencies in the six different niches were able to reach a stable meta-population equilibrium that matched the observed gene arrangement frequencies when recursions used the estimated fitnesses with a variety of initial inversion frequencies. These analyses show that a complex pattern of selection is operating in the six niches to maintain the D. pseudoobscura gene arrangement polymorphism. Models of local adaptation predict that the new inversion mutations were able to invade populations because they held combinations of two to 13 local adaptation loci together. [source] splatche: a program to simulate genetic diversity taking into account environmental heterogeneityMOLECULAR ECOLOGY RESOURCES, Issue 1 2004M. Currat Abstract We present a program called splatche (SPatiaL And Temporal Coalescences in Heterogeneous Environments) to simulate the molecular diversity of samples of genes in an environmentally heterogeneous world. Simulations are performed by, first, simulating the colonization of the world using environmental information to constrain migrations and local densities. These simulated densities and migration rates recorded over time and space are then used to simulate genetic diversity under a coalescent framework. The program thus virtually allows the translation of ecological information into molecular diversity, a novel approach that can be used to study the effect of climatic change on genetic diversity. [source] Coordinated Development of Yeast Colonies: An Experimental Analysis of the Adaptation to Different Nutrient Concentrations , Part 1ENGINEERING IN LIFE SCIENCES (ELECTRONIC), Issue 2 2005T. Walther Abstract The development of yeast colonies on solid agar substrates served as a model system to investigate the growth of higher fungi in a heterogeneous environment. Applying a new analytical technique , which was based on the estimation of the intensity of transmitted light from microscopic images taken along the colony radius , cell-density distributions inside fungal mycelia were measured at an extremely high spatial resolution. Using this method, the adaptation of yeast colonies to the limitation of different nutrients was investigated. Under conditions of carbon or nitrogen limitation, populations of the dimorphic model yeasts Yarrowia lipolytica and Candida boidinii underwent a transition in their morphology from solid colony to mycelial colony patterns. When grown under conditions that induced the mycelial morphology, colonies extended linearly at a constant rate irrespective of the initial nutrient concentration. In general, the cell density within the population declined at higher degrees of limitation. Nitrogen-limited colonies of both model yeasts, as well as carbon-limited Y.,lipolytica colonies proceeded to extend until the growth field was finally covered by the population. Under these conditions, areas of fairly constant cell densities were formed during the growth process. Only carbon-limited C.,boidinii colonies stopped extending at a final diameter which was small when compared to the size of the growth field, and formed a cell-density profile which was monotonically declining. The observed differences in the final colony diameter, and in the cell-density profile morphology indicated the presence of different regulatory mechanisms that ruled the colony development of the model yeasts. The presented monitoring technique for the biomass distribution inside fungal populations provided the basis for a quantitative and non-invasive description of mycelial development. [source] Odor-mediated patch choice in the parasitoid Venturia canescens: temporal decision dynamicsENTOMOLOGIA EXPERIMENTALIS ET APPLICATA, Issue 2 2009Yin-Quan Liu Abstract Parasitoids foraging for hosts in a heterogeneous environment would greatly benefit if they could decide already from a distance in which areas search for resources would be most profitable and to avoid areas of low fitness returns. Interestingly, the temporal dynamics of the decision process in parasitoid patch choice have rarely been investigated. In a Y-tube olfactometer, we tested whether thelytokous and arrhenotokous females of the parasitoid wasp Venturia canescens (Gravenhorst) (Hymenoptera: Ichneumonidae) respond to differences in cues indicating the quality of a host-containing patch and choose more profitable patches. Special attention was given to the time it took females to make their choices (patch choice time) when differences in patch quality were either qualitative (absence vs. presence of hosts and kairomone) or quantitative (various concentrations of hosts and kairomone, and presence of competitors). We found that both thelytokous and arrhenotokous wasps only chose the higher-quality patch based on odor cues when the difference was qualitative. When patches differed only with respect to the number of hosts, or the presence or absence of competing female parasitoids, no significant preference could be found in females of either strain of the parasitoid. In contrast, both the time until females reached the junction of the Y-tube olfactometer (response time) and the time until females decided for either patch (decision time) varied with parasitoid strain and odor treatment. Thelytokous wasps were faster than arrhenotokous wasps in their response time and in their decision time. However, females of both strains responded faster with increasing number of total hosts releasing kairomone. Yet, decision time for patches did not significantly vary as a function of patch quality offered to Venturia wasps. [source] Storage dynamics and streamflow in a catchment with a variable contributing areaHYDROLOGICAL PROCESSES, Issue 16 2010C. Spence Abstract Storage heterogeneity effects on runoff generation have been well documented at the hillslope or plot scale. However, diversity across catchments can increase the range of storage conditions. Upscaling the influence of small-scale storage on streamflow across the usually more heterogeneous environment of the catchment has been difficult. The objective of this study was to observe the distribution of storage in a heterogeneous catchment and evaluate its significance and potential influence on streamflow. The study was conducted in the subarctic Canadian Shield: a region with extensive bedrock outcrops, shallow predominantly organic soils, discontinuous permafrost and numerous water bodies. Even when summer runoff was generated from bedrock hillslopes with small storage capacities, intermediary locations with large storage capacities, particularly headwater lakes, prevented water from transmitting to higher order streams. The topographic bounds of the basin thus constituted the maximum potential contributing area to streamflow and rarely the actual area. Topographic basin storage had little relation to basin streamflow, but hydrologically connected storage exhibited a strong hysteretic relationship with streamflow. This relationship defines the form of catchment function such that the basin can be defined by a series of storing and contributing curves comparable with the wetting and drying curves used in relating tension and hydraulic conductivity to water content in unsaturated soils. These curves may prove useful for catchment classification and elucidating predominant hydrological processes. Copyright © 2009 John Wiley & Sons, Ltd and Her Majesty the Queen in right of Canada. [source] Multimedia transmission with adaptive QoS based on real-time protocolsINTERNATIONAL JOURNAL OF COMMUNICATION SYSTEMS, Issue 3 2003Ch. Bouras Abstract In this paper, we describe a mechanism for adaptive transmission of multimedia data, which is based on real-time protocols. The proposed mechanism can be used for unicast or multicast transmission of multimedia data over heterogeneous networks, like the Internet, and has the capability to adapt the transmission of the multimedia data to network changes. In addition, the implemented mechanism uses an inter-receiver fairness function in order to treat the group of clients with fairness during the multicast transmission in a heterogeneous environment. The proposed mechanism uses a ,friendly' to the network users congestion control policy to control the transmission of the multimedia data. We implement a prototype application based on the proposed mechanism and we evaluate the proposed mechanism both in unicast and multicast transmission through a number of experiment and a number of simulations in order to examine its fairness to a group of clients and its behaviour against transport protocols (TCP) and UDP data streams. Copyright © 2003 John Wiley & Sons, Ltd. [source] Unfairness of measurement-based admission controls in a heterogeneous environmentINTERNATIONAL JOURNAL OF COMMUNICATION SYSTEMS, Issue 9 2001Yuan-Cheng Lai Abstract Admission controls are required to determine whether new connections should be admitted to networks. These controls ensure the quality of service (QoS) for data transmission. This paper introduces three measurement-based admission control algorithms called Measured Sum, Hoeffding Bound, and Adaptive Weight Factor. The unfairness of these algorithms in a heterogeneous environment is investigated. Simulation results indicate that the fairness of the Measured Sum exceeds that of the other methods. Admission of connections with large peak rates or travelling many hops can be difficult. Copyright © 2001 John Wiley & Sons, Ltd. [source] At-sea distribution and scale-dependent foraging behaviour of petrels and albatrosses: a comparative studyJOURNAL OF ANIMAL ECOLOGY, Issue 1 2007DAVID PINAUD Summary 1In order to study and predict population distribution, it is crucial to identify and understand factors affecting individual movement decisions at different scales. Movements of foraging animals should be adjusted to the hierarchical spatial distribution of resources in the environment and this scale-dependent response to environmental heterogeneity should differ according to the forager's characteristics and exploited habitats. 2Using First-Passage Time analysis, we studied scales of search effort and habitat used by individuals of seven sympatric Indian Ocean Procellariiform species fitted with satellite transmitters. We characterized their search effort distribution and examined whether species differ in scale-dependent adjustments of their movements according to the marine environment exploited. 3All species and almost all individuals (91% of 122 individuals) exhibited an Area-Restricted Search (ARS) during foraging. At a regional scale (1000s km), foraging ranges showed a large spatial overlap between species. At a smaller scale (100s km, at which an increase in search effort occurred), a segregation in environmental characteristics of ARS zones (where search effort is high) was found between species. 4Spatial scales at which individuals increased their search effort differed between species and also between exploited habitats, indicating a similar movement adjustment for predators foraging in the same habitat. ARS zones of the two populations of wandering albatross Diomedea exulans (Crozet and Kerguelen) were similar in their adjustments (i.e. same ARS scale) as well as in their environmental characteristics. These two populations showed a weak spatial overlap in their foraging distribution, with males foraging in more southerly waters than females in both populations. 5This study demonstrates that predators of several species adjust their foraging behaviour to the heterogeneous environment and these scale-dependent movement adjustments depend on both forager and environment characteristics. [source] Modelling the establishment and spread of autotetraploid plants in a spatially heterogeneous environmentJOURNAL OF EVOLUTIONARY BIOLOGY, Issue 3 2004B.-H. Li Abstract The establishment and spread of autotetraploids from an original diploid population in a heterogeneous environment were studied using a stochastic simulation model. Specifically, we investigated the effects of heterogeneous habitats and nonrandom pollen/seed dispersal on the critical value (,) of unreduced 2n gamete production necessary for the establishment of autotetraploids as predicted by deterministic models. Introduction of a heterogeneous environment with random pollen/seed dispersal had little effect on the , value. In contrast, incorporating nonrandom pollen/seed dispersal into a homogeneous environment considerably reduced the , value. Incorporating both heterogeneous habitats and nonrandom pollen/seed dispersal may lead either to an increase or to a decrease in the , value compared to that with random dispersal, indicating that the two factors interact in a complex way. [source] Modelling the spatial configuration of refuges for a sustainable control of pests: a case study of Bt cottonJOURNAL OF EVOLUTIONARY BIOLOGY, Issue 3 2003C. Vacher Abstract The ,high-dose-refuge' (HDR) strategy is widely recommended by the biotechnology industry and regulatory authorities to delay pest adaptation to transgenic crops that produce Bacillus thuringiensis (Bt) toxins. This involves cultivating nontoxic plants (refuges) in close proximity to crops producing a high dose of Bt toxin. The principal cost associated with this strategy is due to yield losses suffered by farmers growing unprotected, refuge plants. Using a population genetic model of selection in a spatially heterogeneous environment, we show the existence of an optimal spatial configuration of refuges that could prevent the evolution of resistance whilst reducing the use of costly refuges. In particular, the sustainable control of pests is achievable with the use of more aggregated distributions of nontransgenic plants and transgenic plants producing lower doses of toxin. The HDR strategy is thus suboptimal within the context of sustainable agricultural development. [source] DYNAMICS OF SPATIAL EXPLOITATION: A METAPOPULATION APPROACHNATURAL RESOURCE MODELING, Issue 3 2001JAMES N. SANCHIRICO ABSTRACT. In this paper we present a bioeconomic model of a harvesting industry operating over a heterogeneous environment comprised of discrete biological populations interconnected by dispersal processes. The model generalizes the Gordon [1954]/Smith [1968] model of open-access rent dissipation by accounting for intertemporal and spatial "Ricardian" patterns of exploitation. This model yields a simple, but insightful, framework from which one can investigate factors that contribute to the evolution of resource exploitation patterns over space and time. For example, we find that exploitation patterns are driven by biological and fleet dispersal and biological and economic heterogeneity. We conclude that one cannot really understand the biological processes operating in an exploited system without knowing as much about the harvesting system as about the biological system. [source] Retrodicting patch use by foraging swans in a heterogeneous environment using a set of functional responsesOIKOS, Issue 3 2009Bart A. Nolet Effective conservation of important bird areas requires insight in the number of birds an area can support, and how this carrying capacity changes with habitat modifications. When food depletion is the dominant mechanism of competition, it should in principle be possible to calculate the total time foragers can spend per patch from their functional response (intake rate as a function of food density). However, in the field there are likely to be factors modulating the functional response. In this study previously published results of experiments on captive Bewick's swans were used to obtain functional responses of swans digging for tubers of Fennel pondweed on different foraging substrates: sandy and clayey sediment, and in shallow and deep water. In a field study, four 250×250 m sections belonging to different types (sandy,shallow, clayey,shallow, sandy,deep and clayey,deep) were delineated. Here tubers were sampled with sediment corers in three years, both before and after swan exploitation in autumn, and swans were observed and mapped from a hide in two of these years. Giving-up tuber biomass densities varied among sections. Substitution of these giving-up densities in the derived patch-type-specific functional responses yielded the quitting net energy intake rates in the four sections. As expected from the marginal value theorem, the quitting net energy intake rates did not vary among sections. Moreover, the observed foraging pressure (total foraging time per area) per patch type was in quantitative agreement with the integrated functional responses. These results suggest that in spatially heterogeneous environments, patch exploitation by foragers can be predicted from their functional responses after accounting for foraging substrate. [source] The influence of bite size on foraging at larger spatial and temporal scales by mammalian herbivoresOIKOS, Issue 12 2007Lisa A. Shipley Organisms respond to their heterogeneous environment in complex ways at many temporal and spatial scales. Here, I examine how the smallest scale process in foraging by mammalian herbivores, taking a bite, influences plants and herbivores over larger scales. First, because cropping bites competes with chewing them, bite size influences short-term intake rate of herbivores within plant patches. On the other hand, herbivores can chew bites while searching for new ones, thus influencing the time spent vigilant and intake rate as animals move among food patches. Therefore, bite size affects how much time herbivores must spend foraging each day. Because acquiring energy is necessary for fitness, herbivores recognize the importance of bite size and select bites, patches and diets based on tradeoffs between harvesting rates, digestion, and sheering forces. In turn, induced structural defenses of plants, such as thorns, allow plants to respond immediately to herbivory by reducing bite size and thus tissue loss. Over evolutionary time, herbivores have adapted mouth morphology that allows them to maximize bite size on their primary forage plant, whereas plants faced with large mammalian herbivores have adapted structures such as divarication that minimize bite size and protect themselves from herbivory. Finally, bite size available among plant communities can drive habitat segregation and migration of larger herbivores across landscapes. [source] Backbone structure of a small helical integral membrane protein: A unique structural characterizationPROTEIN SCIENCE, Issue 1 2009Richard C. Page Abstract The structural characterization of small integral membrane proteins pose a significant challenge for structural biology because of the multitude of molecular interactions between the protein and its heterogeneous environment. Here, the three-dimensional backbone structure of Rv1761c from Mycobacterium tuberculosis has been characterized using solution NMR spectroscopy and dodecylphosphocholine (DPC) micelles as a membrane mimetic environment. This 127 residue single transmembrane helix protein has a significant (10 kDa) C-terminal extramembranous domain. Five hundred and ninety distance, backbone dihedral, and orientational restraints were employed resulting in a 1.16 Å rmsd backbone structure with a transmembrane domain defined at 0.40 Å. The structure determination approach utilized residual dipolar coupling orientation data from partially aligned samples, long-range paramagnetic relaxation enhancement derived distances, and dihedral restraints from chemical shift indices to determine the global fold. This structural model of Rv1761c displays some influences by the membrane mimetic illustrating that the structure of these membrane proteins is dictated by a combination of the amino acid sequence and the protein's environment. These results demonstrate both the efficacy of the structural approach and the necessity to consider the biophysical properties of membrane mimetics when interpreting structural data of integral membrane proteins and, in particular, small integral membrane proteins. [source] Asymmetric amino acid compositions of transmembrane ,-strandsPROTEIN SCIENCE, Issue 8 2004Aaron K. Chamberlain Abstract In contrast to water-soluble proteins, membrane proteins reside in a heterogeneous environment, and their surfaces must interact with both polar and apolar membrane regions. As a consequence, the composition of membrane proteins' residues varies substantially between the membrane core and the interfacial regions. The amino acid compositions of helical membrane proteins are also known to be different on the cytoplasmic and extracellular sides of the membrane. Here we report that in the 16 transmembrane ,-barrel structures, the amino acid compositions of lipid-facing residues are different near the N and C termini of the individual strands. Polar amino acids are more prevalent near the C termini than near the N termini, and hydrophobic amino acids show the opposite trend. We suggest that this difference arises because it is easier for polar atoms to escape from the apolar regions of the bilayer at the C terminus of a ,-strand. This new characteristic of ,-barrel membrane proteins enhances our understanding of how a sequence encodes a membrane protein structure and should prove useful in identifying and predicting the structures of trans-membrane ,-barrels. [source] Grassland invertebrate assemblages in managed landscapes: Effect of host plant and microhabitat architectureAUSTRAL ECOLOGY, Issue 6 2007ADELE M. REID Abstract Grasslands are often considered as two-dimensional habitats rather than complex, multilayered habitats. However, native grasslands are complex habitats, with multiple layers of annual and perennial grasses, sedges, shrubs and mosses. Vegetation complexity, including plant type, quality and three-dimensional structure is important for providing a variety of food and habitat resources for insects. Grazing by domestic livestock can affect these processes through the loss or fragmentation of habitats, as well as altering the vertical and horizontal vegetation structure. This study aimed to investigate the role of host plants and microhabitat architecture for determining foliage invertebrate assemblages. Different plant species supported distinct invertebrate assemblages and less complex host plants supported fewer invertebrate individuals and species. Manipulations of plant architecture changed the species composition of invertebrates, with most species found in more complex vegetation. This study illustrates the importance of host diversity and pasture complexity for invertebrate communities. Management practices that encourage a heterogeneous environment with diverse and structurally complex pastures should also sustain a more diverse and functional invertebrate assemblage. [source] Local adaptation in four Iris species tested in a common-garden experimentBIOLOGICAL JOURNAL OF THE LINNEAN SOCIETY, Issue 2 2009MICHAEL DORMAN Local adaptation is a commonly observed result of natural selection acting in heterogeneous environment. Common-garden experiments are a method of detecting local adaptation, as well as studying phenotypic plasticity and gradients of traits. The present study aimed to analyse reaction norms of four closely-related Iris species of section Oncocyclus and to identify a role of environmentally-specific natural selection in their plastic responses. The plant vegetative and phenological, as well as performance traits were measured in a full factorial common-garden experiment with three levels of water amount and three soil types. We found a significant effect of species identity on all traits measured. Water amount and soil type affected many of the traits, but soil type did not affect the performance. There was no significant difference in the effect of water amount and soil type on performance as reflected by rhizome growth; in other words, there was no significant genotype × environment interaction for performance. Plasticity levels and directions of response were also similar among the species. We conclude that phenotypic differences among species are of genetic origin, although no adaptive value was demonstrated for them at the time and life-stages ,frame' of this experiment. © 2009 The Linnean Society of London, Biological Journal of the Linnean Society 2009, 98, 267,277. [source] Quantifying spatial classification uncertainties of the historical Wisconsin landscape (USA)ECOGRAPHY, Issue 2 2005Janine Bolliger Landscape feature can be classified by creating categories based on aggregation of spatially explicit information. However, many landscape features appear continuous rather than discrete. The aggregation process likely involves loss of information and introduces a variety of uncertainties whose degree and extent may differ spatially. Since landscape classifications have found wide application in e.g. natural resource policies or ecological research, assessments of spatial classification uncertainties are required. We present a quantitative framework to identify the degree of landscape continuity (fuzziness) and structure (categorization) based on fuzzy classification and offer measures to quantify uncertainties originating from aggregating features into categories. Fuzzy classification is a non-hierarchical, quantitative method of assessing class definitions using degrees of association between features and class. This results in classes which are well defined and compositionally distinct, as well as classes which are less clearly defined but which, to various degrees, share characteristics with some or all classes. The spatial variation in the degree of class definition on the landscape is used to assess classification uncertainties. The two aspects of uncertainty investigated are the degree of association of a feature with the overall class definitions (membership diffusion), and the class-specific degree of association of each pixel on the landscape with each class (membership saturation). Three classification scenarios, one fuzzy and one discrete, of the historical landscape of Wisconsin (USA) were compared for spatial classification uncertainties. Membership diffusion is highest in topographically heterogeneous environments, or areas characterized by many species occupying similar ecological niches. Classification uncertainties for individual classes show that differentiated species distributions can be identified, not only distribution centers. [source] A comprehensive framework for global patterns in biodiversityECOLOGY LETTERS, Issue 1 2004Robert E. Ricklefs Abstract The present study proposes to reconcile the different spatial and temporal scales of regional species production and local constraint on species richness. Although interactions between populations rapidly achieve equilibrium and limit membership in ecological communities locally, these interactions occur over heterogeneous environments within large regions, where the populations of species are stably regulated through competition and habitat selection. Consequently, exclusion of species from a region depends on long-term regional-scale environmental change or evolutionary change among interacting populations, bringing species production and extinction onto the same scale and establishing a link between local and regional processes. [source] Functional redundancy in heterogeneous environments: implications for conservationECOLOGY LETTERS, Issue 3 2001Todd Wellnitz It has been argued that one of the best ways to conserve biological diversity is to maintain the integrity of functional processes within communities, and this can be accomplished by assessing how much ecological redundancy exists in communities. Evidence suggests, however, that the functional roles species play are subject to the influences of local environmental conditions. Species may appear to perform the same function (i.e. be redundant) under a restricted set of conditions, yet their functional roles may vary in naturally heterogeneous environments. Incorporating the environmental context into ecological experiments would provide a critical perspective for examining functional redundancy among species. [source] Allowing for redundancy and environmental effects in estimates of home range utilization distributionsENVIRONMETRICS, Issue 1 2005W. G. S. Hines Abstract Real location data for radio tagged animals can be challenging to analyze. They can be somewhat redundant, since successive observations of an animal slowly wandering through its environment may well show very similar locations. The data set can possess trends over time or be irregularly timed, and they can report locations in environments with features that should be incorporated to some degree. Also, the periods of observation may be too short to provide reliable estimates of characteristics such as inter-observation correlation levels that can be used in conventional time-series analyses. Moreover, stationarity (in the sense of the data being generated by a source that provides observations of constant mean, variance and correlation structure) may not be present. This article considers an adaptation of the kernel density estimator for estimating home ranges, an adaptation which allows for these various complications and which works well in the absence of exact (or precise) information about correlation structure and parameters. Modifications to allow for irregularly timed observations, non-stationarity and heterogeneous environments are discussed and illustrated. Copyright © 2004 John Wiley & Sons, Ltd. [source] Plasticity of clonal integration in the perennial herb Linaria vulgaris after damageFUNCTIONAL ECOLOGY, Issue 3 2006K. HELLSTRÖM Summary 1Clonal integration in plants can improve their ability to cope with habitat heterogeneity. Integration may increase in response to damage, such as herbivore attack, if undamaged ramets support damaged ones. To test this, we studied the effects of apex removal and substantial defoliation on the performance of the clonal perennial herb Linaria vulgaris Mill. in a common-garden growth experiment and a 13C-labelling study. 2In the growth experiment, contrary to expectations, the target ramet could compensate for damage better when the other ramets in the clone were also damaged, indicating within-clone competition for resources rather than support to damaged ramets. 3In the 13C-labelling experiment, 5·7% of the label moved to a neighbour ramet in controls. Apex removal resulted in a negative net translocation of 13C in the damaged ramet, but defoliation led to zero net translocation. 4The observed lack of support to damaged ramets in Linaria suggests that plasticity of clonal integration in this species includes competition between sibling ramets. Although young ramets may be supported, resources are not directed towards a single damaged ramet if there are more viable intact ramets in the clone. Our main results are consistent with the notion that resource allocation among ramets depends on their relative value in terms of expected fitness profits in heterogeneous environments. [source] A network-centric approach for access and interface selection in heterogeneous wireless environmentsINTERNATIONAL JOURNAL OF COMMUNICATION SYSTEMS, Issue 5 2008George Koundourakis Abstract In this paper, we introduce a network-based approach for access and interface selection (AIS) in the context of resource management in heterogeneous wireless environments (UMTS, WLAN and DVB-T). We focus on the optimization of resource utilization, while ensuring acceptable quality of service (QoS) provision to the end users. Our objective is to optimally manage the overall system resources and minimize the possibility of QoS handovers (non-mobility handovers). The adopted architecture applies to typical heterogeneous environments and network entities (Access Routers) are enhanced with extra functionalities. We propose an AIS algorithm that exploits the multihoming concept and globally manages network resources at both radio access and IP backbone networks. The algorithm can estimate near-optimal solutions in real time and we also introduce a novel triggering policy. We present simulation results of typical scenarios that demonstrate the advantages of our approach. System performance metrics, derived from the simulations, show minimum degradations in high load and congestion situations. Copyright © 2007 John Wiley & Sons, Ltd. [source] Combined fair packet scheduling policy and multi-class adaptive CAC scheme for QoS provisioning in multimedia cellular networksINTERNATIONAL JOURNAL OF COMMUNICATION SYSTEMS, Issue 2 2006Tarek Bejaoui Abstract In this paper, we propose a combined multi-class adaptive connection admission control algorithm and a new fair packet scheduling policy, based on an enhanced weighted fair queuing algorithm. It takes into account a realistic behaviour of traffic while considering the spatial variation of the system characterizing both the user mobility and the signal propagation impairments due to the surrounding effects. The proposed allocation scheme provides enhanced traffic performance in heterogeneous environments and achieves a good level of capacity gain. The new adaptive quality of service (QoS) oriented connection admission control (CAC) function is proposed to meet the rapidly increasing demand for providing multimedia services with diversified quality requirements. It decides whether the connection request is to be admitted into the system, on the basis of the generated interference level and the waiting time before access. This CAC scheme, based on service class differentiation, aims at maximizing the use of available radio resource and meeting the QoS requirement of higher priority users as much as possible while maintaining the minimum requirements of lower priority users, especially when the system suffers from congestion. The results indicate that significantly increased traffic performance can be achieved in comparison with other conventional investigated policies. Copyright © 2006 John Wiley & Sons, Ltd. [source] Effect of genetic variance in plant quality on the population dynamics of a herbivorous insectJOURNAL OF ANIMAL ECOLOGY, Issue 4 2009Nora Underwood Summary 1Species diversity can affect many ecological processes; much less is known about the importance of population genetic diversity, particularly for the population dynamics of associated species. Genetic diversity within a host species can create habitat diversity; when associated species move among hosts, this variation could affect populations additively (an effect of average habitat) or non-additively (an effect of habitat variance). Mathematical theory suggests that non-additive effects of variance among patches should influence population size, but this theory has not been tested. 2This prediction was tested in the field by asking whether aphid population dynamics parameters on strawberry plant genotype mixtures were additive or non-additive functions of parameters on individual plant genotypes in monoculture using model fitting. 3Results show that variance in quality among plant genotypes can have non-additive effects on aphid populations, and that the form of this effect depends on the particular plant genotypes involved. 4Genetic variation among plants also influenced the spatial distribution of aphids within plant populations, but the number of plant genotypes per population did not affect aphid populations. 5These results suggest that predicting the behaviour of populations in heterogeneous environments can require knowledge of both average habitat quality and variance in quality. [source] Local adaptation at the range peripheries of Sitka spruceJOURNAL OF EVOLUTIONARY BIOLOGY, Issue 2 2010M. MIMURA Abstract High-dispersal rates in heterogeneous environments and historical rapid range expansion can hamper local adaptation; however, we often see clinal variation in high-dispersal tree species. To understand the mechanisms of the species' distribution, we investigated local adaptation and adaptive plasticity in a range-wide context in Sitka spruce, a wind-pollinated tree species that has recently expanded its range after glaciations. Phenotypic traits were observed using growth chamber experiments that mimicked temperature and photoperiodic regimes from the limits of the species realized niche. Bud phenology exhibited parallel reaction norms among populations; however, putatively adaptive plasticity and strong divergent selection were seen in bud burst and bud set timing respectively. Natural selection appears to have favoured genotypes that maximize growth rate during available frost-free periods in each environment. We conclude that Sitka spruce has developed local adaptation and adaptive plasticity throughout its range in response to current climatic conditions despite generally high pollen flow and recent range expansion. [source] Environmental stress and the costs of whole-organism phenotypic plasticity in tadpolesJOURNAL OF EVOLUTIONARY BIOLOGY, Issue 1 2008U. K. STEINER Abstract Costs of phenotypic plasticity are important for the evolution of plasticity because they prevent organisms from shaping themselves at will to match heterogeneous environments. These costs occur when plastic genotypes have relatively low fitness regardless of the trait value expressed. We report two experiments in which we measured selection on predator-induced plasticity in the behaviour and external morphology of frog tadpoles (Rana temporaria). We assessed costs under stressful and benign conditions, measured fitness as larval growth rate or competitive ability and focused analysis on aggregate measures of whole-organism plasticity. There was little convincing evidence for a cost of phenotypic plasticity in our experiments, and costs of canalization were nearly as frequent as costs of plasticity. Neither the magnitude of the cost nor the variation around the estimate (detectability) was sensitive to environmental stress. [source] Stable genetic polymorphism in heterogeneous environments: balance between asymmetrical dispersal and selection in the acorn barnacleJOURNAL OF EVOLUTIONARY BIOLOGY, Issue 2 2006D. VÉLIZ Abstract Elucidating the processes responsible for maintaining polymorphism at ecologically relevant genes is intimately related to understanding the interplay between selection imposed by habitat heterogeneity and a species' capacity for dispersal in the face of environmental constraints. In this paper, we used a model-based approach to solve equilibria of balanced polymorphism, given values of fitness and larval dispersal among different habitats in the acorn barnacle Semibalanus balanoides from the Gulf of St Lawrence. Our results showed that allele frequencies observed at both MPI* and GPI* loci represented stable equilibria, given empirical estimates of fitness values, and that considerably more larvae dispersed from one region (north) to the other (south) than vice versa. Dispersal conditions were predicted to be similar for the maintenance of polymorphism at both loci. Moreover, the values of asymmetrical dispersal required by the model to reach stable equilibria were compatible with empirical estimates of larval dispersal and oceanic circulation documented in this system. Overall, this study illustrated the usefulness of a modified and computable version of Bulmer's model (1972) in order to test hypotheses of balanced polymorphism resulting from interactions between spatial selection and asymmetrical dispersal. [source] The experimental evolution of specialists, generalists, and the maintenance of diversityJOURNAL OF EVOLUTIONARY BIOLOGY, Issue 2 2002R. Kassen Environmental heterogeneity may be a general explanation for both the quantity of genetic variation in populations and the ecological niche width of individuals. To evaluate this hypothesis, I review the literature on selection experiments in heterogeneous environments. The niche width usually , but not invariably , evolves to match the amount of environmental variation, specialists evolving in homogeneous environments and generalists evolving in heterogeneous environments. The genetics of niche width are more complex than has previously been recognized, particularly with respect to the magnitude of costs of adaptation and the putative constraints on the evolution of generalists. Genetic variation in fitness is more readily maintained in heterogeneous environments than in homogeneous environments and this diversity is often stably maintained through negative frequency-dependent selection. Moreover environmental heterogeneity appears to be a plausible mechanism for at least two well-known patterns of species diversity at the landscape scale. I conclude that environmental heterogeneity is a plausible and possibly very general explanation for diversity across the range of scales from individuals to landscapes. [source] Numerical simulation of a permittivity probe for measuring the electric properties of planetary regolith and application to the near-surface region of asteroids and cometsMETEORITICS & PLANETARY SCIENCE, Issue 6 2008Klaus SPITZER Our simulation techniques aim at accompanying hardware development and conducting virtual experiments, e.g., to assess the response of arbitrary heterogeneous conductivity and permittivity distributions or to scrutinize possibilities for spatial reconstruction methods using inverse schemes. In a first step, we have developed a finite element simulation code on the basis of unstructured, adaptive triangular grids for arbitrary two-dimensional axisymmetric distributions of conductivity and permittivity. The code is able to take into account the spatial geometry of the probe and allows for possible inductive effects. In previous studies, the non-inductive approach has been used to convert potential and phase data into apparent material properties. By our simulations, we have shown that this approach is valid for the frequency range from 102 Hz to 107 Hz and electric conductivities of 10,8 S/m that are typical for the near-surface region of asteroids and comets composed of chondritic materials and/or frozen volatiles such as H2O and CO2 ice. We prove the accuracy of our code to be better than 10%, using mixed types of boundary conditions and present a simulated vertical log through a horizontally stratified subsurface layer as a representative example of a heterogeneous distribution of the electrical properties. Resolution studies for the given electrode separation reveal that the material parameters of layers having thicknesses of less than about half the electrode spread are not reconstructible if only apparent quantities are considered. Therefore, spatial distributions of the complex sensitivity are presented having in mind a future data inversion concept that will permit the multi-dimensional reconstruction of material parameters in heterogeneous environments. [source] | |||||||||||||||||||||||||||||||