Habitat Choice (habitat + choice)

Distribution by Scientific Domains


Selected Abstracts


ENVIRONMENTAL FACTORS INFLUENCING THE DISTRIBUTION OF SOUTHERN RIGHT WHALES (EUBALAENA AUSTRALIS) ON THE SOUTH COAST OF SOUTH AFRICA I: BROAD SCALE PATTERNS

MARINE MAMMAL SCIENCE, Issue 3 2004
Simon H. Elwen
Abstract Aerial surveys over the last 32 yr have shown that the distribution of southern right whales Eubalaena australis along the south coast of South Africa is markedly discontinuous, but highly predictable. A GIS was used at a variety of scales to investigate whether this pattern was related to environmental characteristics. Whale distribution was analyzed as density per 20-min bin of longitude over two temporal and spatial scales, namely 15 bins for 32 yr, and a wider scale but shorter time period, 23 bins for 19 yr, as well as using three years of GPS accuracy data (15 bins) for finer scale analysis. Environmental factors tested were depth, distance from shore, sea floor slope, protection from swell, protection from wind, and shore type. The majority of whales were concentrated in areas that provided reasonable protection from open ocean swell and seasonal winds, and had sedimentary floors with gentle slopes. They generally avoided exposed rocky shorelines. Cow-calf pairs were found significantly closer to shore and in shallower water than unaccompanied whales, particularly off sandy beaches. Habitat choice at this time of year may be related both to energy conservation for calves and lactating females (calm sea conditions) and to protection of the new-born. [source]


Spatio-temporal shifts in gradients of habitat quality for an opportunistic avian predator

ECOGRAPHY, Issue 2 2003
Fabrizio Sergio
We used the conceptual framework of the theory of natural selection to study breeding habitat preferences by an opportunistic avian predator, the black kite Milvus migrans. In Europe, black kite populations are mostly found near large networks of aquatic habitats, usually considered optimal for foraging and breeding. We hypothesized that proximity to wetlands could vary among individuals and affect their fitness, and thus be subject to natural selection. We tested the hypothesis first on a population on Lake Lugano (Italian pre-Alps) which has been monitored for nine years, and then on seven other populations, each studied for four,five years, located along a continuum of habitat from large water bodies to scarce aquatic habitat of any kind. In the Lake Lugano population, black kite abundance was negatively related to distance to the lake in all the nine years of study, consistent with long-term natural selection. There was evidence of ongoing directional selection on strategic nest location in three of the years, and evidence of stabilizing selection in two years. In eight of the nine years the trend was for a linear increase in fitness with increasing proximity to the lake. At the population level, results were consistent with adaptive habitat choice in relation to the previous year's spatial variation in fitness: higher associations between fitness and distance to the lake (i.e. higher selection gradients) resulted in higher density variations in the following year, in turn related to the availability of fish, the main local prey. The progressive decline of inland pairs and increase in the density of lakeshore pairs caused a directional long-term trend of declining mean distance to the lake. Breeding near aquatic habitats was associated with higher foraging success, and higher frequency and biomass of prey deliveries to offspring. There was weak evidence of selection in other populations. The inland-wetland gradient of habitat quality may have been affected by predation risk, as estimated by density of a major predator of adults and nestlings, the eagle owl Bubo bubo. Behavioral decisions at the level of the individual probably translated into population effects on density and distribution at various spatial scales. Populations in optimal habitats showed higher density and produced six times as many young per unit space as those in sub-optimal habitats. [source]


Informed dispersal, heterogeneity in animal dispersal syndromes and the dynamics of spatially structured populations

ECOLOGY LETTERS, Issue 3 2009
Jean Clobert
Abstract There is accumulating evidence that individuals leave their natal area and select a breeding habitat non-randomly by relying upon information about their natal and future breeding environments. This variation in dispersal is not only based on external information (condition dependence) but also depends upon the internal state of individuals (phenotype dependence). As a consequence, not all dispersers are of the same quality or search for the same habitats. In addition, the individual's state is characterized by morphological, physiological or behavioural attributes that might themselves serve as a cue altering the habitat choice of conspecifics. These combined effects of internal and external information have the potential to generate complex movement patterns and could influence population dynamics and colonization processes. Here, we highlight three particular processes that link condition-dependent dispersal, phenotype-dependent dispersal and habitat choice strategies: (1) the relationship between the cause of departure and the dispersers' phenotype; (2) the relationship between the cause of departure and the settlement behaviour and (3) the concept of informed dispersal, where individuals gather and transfer information before and during their movements through the landscape. We review the empirical evidence for these processes with a special emphasis on vertebrate and arthropod model systems, and present case studies that have quantified the impacts of these processes on spatially structured population dynamics. We also discuss recent literature providing strong evidence that individual variation in dispersal has an important impact on both reinforcement and colonization success and therefore must be taken into account when predicting ecological responses to global warming and habitat fragmentation. [source]


Fish assemblages as influenced by environmental factors in streams in protected areas of the Czech Republic

ECOLOGY OF FRESHWATER FISH, Issue 1 2006
M. Humpl
Abstract,,, Three streams of comparable size located in different landscape-protected areas were selected for studying the effect of environmental factors on fish assemblages using indirect (detrended correspondence analysis, DCA) and direct (canonical correspondence analysis, CCA) gradient analysis. DCA of species showed well a gradient of assemblage changes in the longitudinal profile. DCA of sites stressed the variability between the fish assemblages of the three streams. This pattern was then confirmed by the highly significant between-stream CCA. In the within-site CCA, environmental factors explained 50.7% variability for presence,absence data and 58.3% for the relative abundance data. The analysis revealed that number of ponds and land use are the most influential factors of the strongest environmental gradient. However, in the partial CCAs, factor substratum type explained the largest proportion of the variability affecting fish in their habitat choice. Generally, presence,absence and relative abundance data of fish gave similar results in both DCA and CCA analyses; the same environmental factors proved to be important in both data type analyses. The environmental factors explain more variability than the regional (between-stream) one. The total proportion of variability explained by the presence,absence data analysis was 71.9% and in the relative abundance analysis even 80.8%. The environmental factors measured during the field survey explain 2.1- and 3.4-times more assemblages' variability than factors measured from a hydrological map. Resumen 1. Tres ríos de tamaño comparable localizados en diferentes áreas de paisaje protegido de la República Checa fueron seleccionados para estudiar el efecto de factores ambientales sobre los ensamblajes de peces. Para ello, utilizando análisis de gradientes indirectos (DCA) y directos (CCA). 2. El DCA para las especies enfatizó la variabilidad entre los ensamblajes de peces de los tres ríos. Este patrón fue confirmado por un CCA altamente significativo. Para la variabilidad dentro de la localidad, un CCA reveló que los factores ambientales explicaron un 50.7% para datos de presencia-ausencia y un 58.3% para las abundancias relativas. 3. Los análisis revelaron que el número de pozas y el uso del suelo fueron los factores de mayor influencia en el gradiente ambiental. Sin embargo, en el CCA parcial, el tipo de sustrato explicó la mayor proporción de la variabilidad que afecta a los peces en la elección de hábitat. 4. Generalmente los datos de presencia-ausencia y abundancia relativa produjeron resultados similares tanto en los análisis DCA como en los CCA; los mismos factores ambientales probaron ser importantes en los análisis de ambos tipos de datos. Los factores ambientales explicaron mas variabilidad que los regionales (entre ríos). La proporción total de variabilidad explicada por el análisis de los datos de presencia-ausencia fue 71.9% mientras que para las abundancias relativas fue de 80.8%. Los factores ambientales medidos durante los muestreos de campo explicaron 2.1 y 3.4 veces mas variabilidad que los factores medidos sobre mapas hidrológicos. [source]


Migration of rheophilic fish in the large lowland rivers Meuse and Rhine, the Netherlands

FISHERIES MANAGEMENT & ECOLOGY, Issue 5-6 2008
J. J. DE LEEUW
Abstract, Large-scale migratory patterns of adult rheophilic fish [barbel, Barbus barbus (L.), chub, Leuciscus cephalus (L.), ide, Leuciscus idus (L.), nase, Chondrostoma nasus (L.)] were studied in relation to habitat quality and possible migration barriers in the lower rivers Meuse and Rhine, the Netherlands, using a telemetry system with transponders and detection stations based on inductive coupling. Most fish moved over short distances (<10 km), especially those residing in river stretches with high habitat diversity year-round. About 16% of fish used river stretches over 50 km and two ide moved more than 200 km along free-flowing river stretches. One-third of barbel, chub and nase from a Weir-regulated river stretch moved upstream during the spawning season to spawning habitats. Some fish resided in the areas immediately downstream of weirs and fishways during the spawning season, although it was unclear to what extent these observations reflected habitat choice or barriers to migration. [source]


Effect of light and predator abundance on the habitat choice of plant-attached zooplankton

FRESHWATER BIOLOGY, Issue 3 2007
LEENA NURMINEN
Summary 1. The diurnal variations in the habitat choice of the periodically plant-attached cladoceran Sida crystallina together with light environment and predator abundance were studied. 2. The density of S. crystallina attached to floating leaves of Nuphar lutea increased between 18:00 and 20:00 hours, when light intensity underneath the leaves was temporarily increased, and decreased again when light intensity declined. A proportion of S. crystallina remained in the swimming mode underneath the leaves even during daylight, indicating that the water column sheltered by the leaves is safer than the open water. 3. In the water adjacent to the leaves, the density of S. crystallina increased steeply in the dark. The increase was not accompanied by a decrease in S. crystallina attached to plant leaves, indicating that the nocturnal increment in the open water density of S. crystallina was due to migration from daytime refuges other than floating leaves. 4. Sida crystallina was most intensively consumed by perch (Perca fluviatilis). Predation threat by fish had weaker effects on the density of S. crystallina attached to plant leaves than on cladocerans in the adjacent water. Cladocerans underneath floating plant leaves, whether attached or not, are probably less vulnerable to fish predation than those outside the leaf cover. 5. The results suggested that light intensity is the proximate factor regulating the attachment of zooplankton to the lower surfaces of floating macrophyte leaves. Light intensity has a positive effect on the density of S. crystallina attached to the floating leaves and a negative effect on density in the water. Predation threat by fish has a strong effect on the migration of zooplankters into the open water habitat. [source]


The energetic equivalence of changing salinity and temperature to juvenile salmon

FUNCTIONAL ECOLOGY, Issue 4 2006
S. J. WEBSTER
Summary 1The influence of salinity, temperature and physiological development on habitat choice by juvenile salmon is poorly understood. We measured differences in the net energetic cost of habitats that differed in salinity or temperature using behavioural titration of juvenile salmon and correlated these costs with body size and osmoregulatory enzyme activity to quantify how costs change with physiological development. 2Juvenile salmon showed a strong preference for saline water (27 or 15 vs 0) and for cold water (9 °C vs 14 °C). It was estimated to be 0·15 W and 0·11 W more costly for salmon to forage in fresh water than in 15 and 27 salt water, respectively, and 0·09 W more costly to forage in warm than in cold water. 3We expected fish to prefer 15 salt water to fresh water regardless of enzyme activity because they are iso-osmotic with this salt concentration. In contrast, preference for higher salt concentrations should increase with enzyme activity. Consistent with our expectations, enzyme activity was not correlated with preference for 15 salt water, but was positively correlated with preference for 27 salt water. 4The cost of changing salinity vs changing temperature were very similar, indicating that knowledge of both of these costs are necessary to understand juvenile salmon habitat choice. [source]


Vigilance and fitness in grey partridges Perdix perdix: the effects of group size and foraging-vigilance trade-offs on predation mortality

JOURNAL OF ANIMAL ECOLOGY, Issue 2 2007
MARK WATSON
Summary 1Vigilance increases fitness by improving predator detection but at the expense of increasing starvation risk. We related variation in vigilance among 122 radio-tagged overwintering grey partridges Perdix perdix (L.) across 20 independent farmland sites in England to predation risk (sparrowhawk Accipiter nisus L., kill rate), use of alternative antipredation behaviours (grouping and use of cover) and survival. 2Vigilance was significantly higher when individuals fed in smaller groups and in taller vegetation. In the covey period (in early winter when partridges are in flocks), vigilance and use of taller vegetation was significantly higher at sites with higher sparrowhawk predation risk, but tall vegetation was used less by larger groups. Individuals were constrained in reducing individual vigilance by group size and habitat choice because maximum group size was determined by overall density in the area during the covey period and by the formation of pairs at the end of the winter (pair period), when there was also a significant twofold increase in the use of tall cover. 3Over the whole winter individual survival was higher in larger groups and was lower in the pair period. However, when controlling for group size, mean survival decreased as vigilance increased in the covey period. This result, along with vigilance being higher at sites with increasing with raptor risk, suggests individual vigilance increases arose to reduce short-term predation risk from raptors but led to long-term fitness decreases probably because high individual vigilance increased starvation risk or indicated longer exposure to predation. The effect of raptors on survival was less when there were large groups in open habitats, where individual partridges can probably both detect predators and feed efficiently. 4Our study suggests that increasing partridge density and modifying habitat to remove the need for high individual vigilance may decrease partridge mortality. It demonstrates the general principle that antipredation behaviours may reduce fitness long-term via their effects on the starvation,predation risk trade-off, even though they decrease predation risk short-term, and that it may be ecological constraints, such as poor habitat (that lead to an antipredation behaviour compromising foraging), that cause mortality, rather than the proximate effect of an antipredation behaviour such as vigilance. [source]


Effect of wave exposure dynamics on gut content mass and growth of young-of-the-year fishes in the littoral zone of lakes

JOURNAL OF FISH BIOLOGY, Issue 7 2010
S. Stoll
Total length, body mass and gut content mass of young-of-the-year (YOY) perch Perca fluviatilis, dace Leuciscus leuciscus and bleak Alburnus alburnus were recorded over the summer of 2006 at three littoral sites at Upper Lake Constance. In P. fluviatilis and L. leuciscus, gut content mass correlated positively with wave-induced energy flux (EF) of the respective site and sampling day, while no correlation of gut content mass with EF was found in A. alburnus. It was assumed that benthivorous P. fluviatilis and L. leuciscus profited from suspended or uncovered benthic food items generated by wave action at sites and periods with high EF. Alburnus alburnus, in contrast, feeding mainly on zooplankton in upper parts of the water column, could not profit from increased EF. In P. fluviatilis, increased gut content mass during periods of high EF resulted in higher growth rates. For L. leuciscus, no real growth rates in local fish populations could be determined, as individuals were less sedentary, and when increased growth occurred at sites during the periods of high EF, migration of fish levelled out the resulting size differences within few days. The results of this study show that dynamic habitat variables affect site profitability in the littoral zone of lakes, especially in benthivorous fishes. Therefore, dynamic habitat variables should be considered in addition to fixed habitat properties in analyses of habitat choice of fishes in the littoral zone of lakes. [source]


The trade-off between food and temperature in the habitat choice of bluegill sunfish

JOURNAL OF FISH BIOLOGY, Issue 5 2001
M. L. Wildhaber
A model is presented to describe the trade-off between food and temperature in bluegills Lepomis macrochirus, where temperature was the primary factor used in determining the patch in which to reside. [source]


ENVIRONMENTAL FACTORS INFLUENCING THE DISTRIBUTION OF SOUTHERN RIGHT WHALES (EUBALAENA AUSTRALIS) ON THE SOUTH COAST OF SOUTH AFRICA II: WITHIN BAY DISTRIBUTION

MARINE MAMMAL SCIENCE, Issue 3 2004
Simon H. Elwen
Abstract Environmental factors are thought to strongly influence the distribution and predictability of the coastal distribution of southern right whales (Eubalaena australis) off South Africa. Preferred habitat had generally shallow sloping sedimentary floors and was characteristically protected from open ocean swell and prevalent seasonal winds. This study investigated whether habitat choices at smaller scales (within bays) were similar. Fine scale distribution patterns (GPS) from three years' surveys (1997, 1999, 2000) were analyzed separately within the three main concentration areas St Sebastian Bay, De Hoop, and Walker Bay (containing ,73% of cow-calf pairs and ,49% of unaccompanied adults in the whole survey region). Whale density at this scale of within particular bays did not correlate well with predicted variables, but Chi-squared analysis strongly supported results at broader scales, in all bays. Post-hoc"choice" tests between similar areas differing in only one variable revealed that cow-calves preferred (presumed) sandy substrates and especially protection from swell. The strength and predictability of preferences shown at fine scale (where individual movement and weather variability could have great influence) provide strong support for findings at larger scales and emphasize the importance of environmental factors in the habitat choice of wintering right whales. [source]


Intraspecific competition drives multiple species resource polymorphism in fish communities

OIKOS, Issue 1 2008
Richard Svanbäck
It has been hypothesized that inter-specific competition will reduce species niche utilization and drive morphological evolution in character displacement. In the absence of a competitor, intra-specific competition may favor an expansion of the species niche and drive morphological evolution in character release. Despite of this theoretical framework, we sometimes find potential competitor species using the same niche range without any partitioning in niche. We used a database on test fishing in Sweden to evaluate the factors (inter- and intraspecific competition, predation, and abiotic factors) that could influence habitat choice of two competitor species. The pattern from the database shows that the occurrence of perch and roach occupying both littoral and pelagic habitats of lakes in Sweden is a general phenomenon. Furthermore, the results from the database suggest that this pattern is due to intra-specific competition rather than inter-specific competition or predation. In a field study, we estimated the morphological variation in perch and roach and found that, individuals of both species caught in the littoral zone were more deeper bodied compared to individuals caught in the pelagic zone. Pelagic perch fed more on zooplankton compared to littoral perch, independent of size, whereas the littoral perch had more macroinvertebrates and fish in their diet. Pelagic roach fed more on zooplankton compared to littoral roach, whereas littoral individuals fed more on plant material. Furthermore, we sampled littoral and pelagic fish from another lake to evaluate the generality of our first results and found the same habitat associated morphology in both perch and roach. The results show a consistent multi-species morphological separation in the littoral and pelagic habitats. This study suggests that intra-specific competition is possibly more important than inter-specific competition for the morphological pattern in the perch-roach system. [source]


Experimental manipulation reveals the importance of refuge habitat temperature selected by lizards

AUSTRAL ECOLOGY, Issue 3 2010
MICHA ANDERSSON
Abstract Refuges provide shelter from predators, and protection from exposure to the elements, as well as other fitness benefits to animals that use them. In ectotherms, thermal benefits may be a critical aspect of refuges. We investigated microhabitat characteristics of refuges selected by a heliothermic scincid lizard, Carlia rubrigularis, which uses rainforest edges as habitat. We approached lizards in the field, simulating a predator attack, and quantified the refuge type used, and effect of environmental temperatures (air temperature, substrate temperature and refuge substrate temperature) on the amount of time skinks remained in refuges after hiding (emergence time). In respone to our approach, lizards were most likely to flee into leaf litter, rather than into rocks or woody debris, and emergence time was dependent on refuge substrate temperature, and on refuge substrate temperature relative to substrate temperature outside the refuge. Lizards remained for longer periods in warmer refuges, and in refuges that were similar in temperature to outside. We examined lizard refuge choice in response to temperature and substrate type in large, semi-natural outdoor enclosures. We experimentally manipulated refuge habitat temperature available to lizards, and offered them equal areas of leaf litter, woody debris and rocks. When refuge habitat temperature was unmanipulated, lizards (85%) preferred leaf litter, as they did in the field. However, when we experimentally manipulated the temperature of the leaf litter by shading, most skinks (75%) changed their preferred refuge habitat from leaf litter to woody debris or rocks. These results suggest that temperature is a critical determinant of refuge habitat choice for these diurnal ectotherms, both when fleeing from predators and when selecting daytime retreats. [source]


Habitat disturbance, not predation, is all that is required to influence habitat choice in juvenile snakes: A rejoinder to Lill

AUSTRAL ECOLOGY, Issue 7 2006
JONATHAN K. WEBB
No abstract is available for this article. [source]


Flying Foxes Prefer to Forage in Farmland in a Tropical Dry Forest Landscape Mosaic in Fiji

BIOTROPICA, Issue 2 2010
Article first published online: 29 SEP 200, Matthew Scott Luskin
ABSTRACT To test flying fox adaptations to a habitat mosaic with extreme deforestation, the abundance, habitat choice and feeding behavior of the Pacific flying fox, Pteropus tonganus, were investigated across 16 islands of the Yasawa archipelago, Fiji. The habitat mosaic is formed by 4.3 percent tropical dry forest and 3.3 percent farmland, leaving exotic grasslands and stands of Leucaena leucocephala to overrun the vast majority of land. Pteropus tonganus abundance was high (5757 bats) despite deforestation and hunting. Roosting sites were restricted to native forest fragments. Grasslands and stands of L. leucocephala were completely void of bats at all times. The mean foraging density in farmland was four times higher than in forests and foraging competition was routinely observed in farmland but was extremely rare in forests. The author suggests that during the study, extensive foraging in farmland was supporting the high P. tonganus population. Additionally, the preferential foraging in farmland was responsible for the low foraging densities within forests and dramatically less intraspecies competition for forest resources. Further research is needed on seed dispersal within forests and to test for seasonal variations in bat abundance and feeding. [source]


Assessments of habitat preferences and quality depend on spatial scale and metrics of fitness

JOURNAL OF APPLIED ECOLOGY, Issue 5 2007
ANNA D. CHALFOUN
Summary 1Identifying the habitat features that influence habitat selection and enhance fitness is critical for effective management. Ecological theory predicts that habitat choices should be adaptive, such that fitness is enhanced in preferred habitats. However, studies often report mismatches between habitat preferences and fitness consequences across a wide variety of taxa based on a single spatial scale and/or a single fitness component. 2We examined whether habitat preferences of a declining shrub steppe songbird, the Brewer's sparrow Spizella breweri, were adaptive when multiple reproductive fitness components and spatial scales (landscape, territory and nest patch) were considered. 3We found that birds settled earlier and in higher densities, together suggesting preference, in landscapes with greater shrub cover and height. Yet nest success was not higher in these landscapes; nest success was primarily determined by nest predation rates. Thus landscape preferences did not match nest predation risk. Instead, nestling mass and the number of nesting attempts per pair increased in preferred landscapes, raising the possibility that landscapes were chosen on the basis of food availability rather than safe nest sites. 4At smaller spatial scales (territory and nest patch), birds preferred different habitat features (i.e. density of potential nest shrubs) that reduced nest predation risk and allowed greater season-long reproductive success. 5Synthesis and applications. Habitat preferences reflect the integration of multiple environmental factors across multiple spatial scales, and individuals may have more than one option for optimizing fitness via habitat selection strategies. Assessments of habitat quality for management prescriptions should ideally include analysis of diverse fitness consequences across multiple ecologically relevant spatial scales. [source]


ENVIRONMENTAL FACTORS INFLUENCING THE DISTRIBUTION OF SOUTHERN RIGHT WHALES (EUBALAENA AUSTRALIS) ON THE SOUTH COAST OF SOUTH AFRICA II: WITHIN BAY DISTRIBUTION

MARINE MAMMAL SCIENCE, Issue 3 2004
Simon H. Elwen
Abstract Environmental factors are thought to strongly influence the distribution and predictability of the coastal distribution of southern right whales (Eubalaena australis) off South Africa. Preferred habitat had generally shallow sloping sedimentary floors and was characteristically protected from open ocean swell and prevalent seasonal winds. This study investigated whether habitat choices at smaller scales (within bays) were similar. Fine scale distribution patterns (GPS) from three years' surveys (1997, 1999, 2000) were analyzed separately within the three main concentration areas St Sebastian Bay, De Hoop, and Walker Bay (containing ,73% of cow-calf pairs and ,49% of unaccompanied adults in the whole survey region). Whale density at this scale of within particular bays did not correlate well with predicted variables, but Chi-squared analysis strongly supported results at broader scales, in all bays. Post-hoc"choice" tests between similar areas differing in only one variable revealed that cow-calves preferred (presumed) sandy substrates and especially protection from swell. The strength and predictability of preferences shown at fine scale (where individual movement and weather variability could have great influence) provide strong support for findings at larger scales and emphasize the importance of environmental factors in the habitat choice of wintering right whales. [source]


The influence of increasing population size and vegetation productivity on elephant distribution in the Kruger National Park

AUSTRAL ECOLOGY, Issue 3 2009
KIM D. YOUNG
Abstract Decisions to reduce the impacts of large herbivores on biodiversity in protected areas are often based on controlling their numbers. However, numbers per se may not be the foremost consideration when managing impacts. This is because density-related changes in distribution can also affect habitat utilization and hence, impact. In this study we tested whether changes in the distribution of African elephants are associated with increasing population size. We used spatially explicit count data collected during the dry seasons from 1998 to 2004 in South Africa's Kruger National Park. We did this at five spatial scales and in landscapes defined by vegetation, geology, climate and soils. We then investigated whether observed distributions and grid-cell-specific densities were associated with the remotely sensed Normalized Difference Vegetation Index (NDVI) as a measure of productivity and therefore food resource availability at the landscape scale. Consistent with density mediated changes, we found that elephant grid-cell occupancy increased with population size, while grid-cell-specific density became less variable. In addition, the combined distribution of bull groups and breeding herds became less clumped with increasing population size. We further found that within landscapes elephants were present on grid-cells with higher NDVI values, but that the influence of NDVI during the dry season on densities among landscapes may be weak. These results suggest that NDVI was more indicative of structural habitat choices such as woody vegetation than food availability per se. Our study highlights the need to consider factors other than population size alone when formulating management decisions to reduce large herbivore impacts on biodiversity in protected areas. [source]