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Glaciation
Kinds of Glaciation Terms modified by Glaciation Selected AbstractsGLACIATION OF MT ALLEN, STEWART ISLAND (RAKIURA): THE SOUTHERN MARGIN OF LGM GLACIATION IN NEW ZEALANDGEOGRAFISKA ANNALER SERIES A: PHYSICAL GEOGRAPHY, Issue 2 2009MARTIN S. BROOK ABSTRACT. The origin of two ridges on the eastern slopes of Mt Allen, southern Stewart Island, has remained equivocal, with differences of opinion over the exact process-mechanisms of formation. A variety of approaches was used to test a number of possible hypotheses about the origin of the ridges. These include topographic and spatial positioning, geomorphology, sedimentology and palaeoclimatological extrapolations to reconstruct two small former cirque glaciers with equilibrium line altitudes (ELAs) of c. 600 m. It would appear the two ridges reflect a glacial origin, the glaciers interpreted as forming during the Last Glacial Maximum (LGM) in New Zealand. Whilst glaciation during this time (18,19 ka) was extensive in the Southern Alps, the restricted nature of glaciation on Mt Allen suggests the low altitude restricted glaciation to niche sites on the lee side of upland areas. [source] THE LAST GLACIATION OF SHETLAND, NORTH ATLANTICGEOGRAFISKA ANNALER SERIES A: PHYSICAL GEOGRAPHY, Issue 1 2008N.R. GOLLEDGE ABSTRACT. Evidence relating to the extent, dynamics, and relative chronology of the last glaciation of the Shetland Islands, North Atlantic, is presented here, in an attempt to better illuminate some of the controversies that still surround the glacial history of the archipelago. We appraise previous interpretations and compare these earlier results with new evidence gleaned from the interpretation of a high resolution digital terrain model and from field reconnaissance. By employing a landsystems approach, we identify and describe three quite different assemblages of landscape features across the main islands of Mainland, Yell and Unst. Using the spatial interrelationship of these landsystems, an assessment of their constituent elements, and comparisons with similar features in other glaciated environments, we propose a simple model for the last glaciation of Shetland. During an early glacial phase, a coalescent British and Scandinavian ice sheet flowed approximately east to west across Shetland. The terrestrial land-forms created by this ice sheet in the north of Shetland suggest that it had corridors of relatively fast-flowing ice that were partially directed by bed topography, and that subsequent deglaciation was interrupted by at least one major stillstand. Evidence in the south of Shetland indicates the growth of a local ice cap of restricted extent that fed numerous radial outlet glaciers during, or after, ice-sheet deglaciation. Whilst the absolute age of these three landsystems remains uncertain, these new geo-morphological and palaeoglaciological insights reconcile many of the ideas of earlier workers, and allow wider speculation regarding the dynamics of the former British ice sheet. [source] THE EASTERN MARGIN OF GLACIATION IN THE BRITISH ISLES DURING THE YOUNGER DRYAS: THE BIZZLE CIRQUE, SOUTHERN SCOTLANDGEOGRAFISKA ANNALER SERIES A: PHYSICAL GEOGRAPHY, Issue 3 2006STEPHAN HARRISON ABSTRACT. Geomorphological and sedimentological evidence of former glaciation in the Bizzle valley in the Cheviot Hills of northern England and southern Scotland was used to reconstruct the dimensions of a small topographically constrained glacier with an equilibrium line altitude (ELA) of 535 m. This was interpreted as having formed during Younger Dryas cooling; this is the only glacier to have been described from the area and is the most easterly site of Younger Dryas glaciation in the British Isles. Whilst glaciation at this time was extensive in the Lake District to the southwest, the restricted nature of Cheviot ice cover suggests that a steep west,east precipitation gradient existed in this region during the Younger Dryas. [source] CHRONOLOGY OF THE LAST GLACIATION IN CENTRAL STRAIT OF MAGELLAN AND BAHÍA INÚTIL, SOUTHERNMOST SOUTH AMERICAGEOGRAFISKA ANNALER SERIES A: PHYSICAL GEOGRAPHY, Issue 2 2005R.D. McCULLOCH ABSTRACT. Glacier fluctuations in the Strait of Magellan tell of the climatic changes that affected southern latitudes at c. 53,55°S during the Last Glacial Maximum (LGM) and Late-glacial/Holocene transition. Here we present a revised chronology based on cosmogenic isotope analysis, 14C assays, amino acid racemisation and tephrochronology. We unpick the effect of bedrock-derived lignite which has affected many 14C dates in the past and synthesise new and revised dates that constrain five glacier advances (A to E). Advance A is prior to the LGM. LGM is represented by Advance B that reached and largely formed the arcuate peninsula Juan Mazia. Carbon-14and 10Be dating show it occurred after 31 250 cal yrs BP and culminated at 25 200,23 100 cal yrs BP and was then followed by the slightly less extensive advance C sometime before 22 400,20 300 cal yrs BP. This pattern of an early maximum is found elsewhere in South America and more widely. Stage D, considerably less extensive, culminated sometime before 17 700,17 600 cal yrs BP and was followed by rapid and widespread glacier retreat. Advance E, which dammed a lake, spanned 15 500,11770 cal yrs BP. This latter advance overlaps the Bølling-Allerød interstadials and the glacier retreat occurs during the peak of the Younger Dryas stadial in the northern hemisphere. However, the stage E advance coincides with the Antarctic Cold Reversal (c. 14800,12700 cal yrs BP) and may indicate that some millennial-scale climatic fluctuations in the Late-glacial period are out of phase between the northern and southern hemispheres. [source] A multi-dating approach applied to proglacial sediments attributed to the Most Extensive Glaciation of the Swiss AlpsBOREAS, Issue 3 2010ANDREAS DEHNERT Dehnert, A., Preusser, F., Kramers, J. D., Akçar, N., Kubik, P. W., Reber, R. & Schlüchter, C. 2010: A multi-dating approach applied to proglacial sediments attributed to the Most Extensive Glaciation of the Swiss Alps. Boreas, Vol. 39, pp. 620,632. 10.1111/j.1502-3885.2010.00146.x. ISSN 0300-9483. The number and the timing of Quaternary glaciations of the Alps are poorly constrained and, in particular, the age of the Most Extensive Glaciation (MEG) in Switzerland remains controversial. This ice advance has previously been tentatively correlated with the Riss Glaciation of the classical alpine stratigraphy and with Marine Isotope Stage (MIS) 6 (186,127 ka). An alternative interpretation, based on pollen analysis and stratigraphic correlations, places the MEG further back in the Quaternary, with an age equivalent to MIS 12 (474,427 ka), or even older. To re-evaluate this issue in the Swiss glaciation history, a multi-dating approach was applied to proglacial deltaic ,Höhenschotter' deposits in locations outside the ice extent of the Last Glacial Maximum. Results of U/Th and luminescence dating suggest a correlation of the investigated deposits with MIS 6 and hence with the Riss Glaciation. Cosmogenic burial dating suffered from large measurement uncertainties and unusually high 26Al/10Be ratios and did not provide robust age estimates. [source] Regional assessment of the impact of climatic change on the distribution of a tropical conifer in the lowlands of South AmericaDIVERSITY AND DISTRIBUTIONS, Issue 6 2007Marie-Pierre Ledru Abstract For decades, palynologists working in tropical South America are using the genus Podocarpus as a climate indicator although without referring to any modern data concerning its distribution and limiting factors. With the aim to characterize the modern and past distribution of the southern conifer Podocarpus in Brazil and to obtain new information on the distribution of the Atlantic rainforest during the Quaternary, we examined herbarium data to locate the populations of three Brazilian endemic Podocarpus species: P. sellowii, P. lambertii, and P. brasiliensis, and extracted DNA from fresh leaves from 26 populations. Our conclusions are drawn in the light of the combination of these three disciplines: botany, palynology, and genetics. We find that the modern distribution of endemic Podocarpus populations shows that they are widely dispersed in eastern Brazil, from north to south and reveals that the expansion of Podocarpus recorded in single Amazonian pollen records may have come from either western or eastern populations. Genetic analysis enabled us to delimit regional expansion: between 5° and 15° S grouping northern and central populations of P. sellowii expanded c. 16,000 years ago; between 15° and 23° S populations of either P. lambertii or sellowii expanded at different times since at least the last glaciation; and between 23° and 30° S, P. lambertii appeared during the recent expansion of the Araucaria forest. The combination of botany, pollen, and molecular analysis proved to be a rapid tool for inferring distribution borders for sparse populations and their regional evolution within tropical ecosystems. Today the refugia of rainforest communities we identified are crucial hotspots to allow the Atlantic forest to survive under unfavourable climatic conditions and, as such, offer the only possible opportunity for this type of forest to expand in the event of a future climate change. [source] Patterns of isozyme variation as indicators of biogeographic history in Pilgerodendron uviferum (D. Don) FlorínDIVERSITY AND DISTRIBUTIONS, Issue 2 2002A. C. Premoli Abstract. The effects of Pleistocene glaciations on the genetic characteristics of the most austral conifer in the world, Pilgerodendron uviferum, were analysed with specific reference to the hypothesis that the species persisted locally in ice-free areas in temperate South America. It was expected that genetic variation would decrease with latitude, given that ice fields were larger in southern Patagonia and thus refugia were probably located towards the northern distributional limit of the species as suggested by the fossil record. In addition, an increase in among-population genetic divergence was expected with increasing distance to putative glacial refugia. We examined the relationship between location and within-population variability indices of 20 Pilgerodendron populations derived from isozyme analyses. We analysed possible refugia hypotheses by the distribution of allele frequencies using multivariate discriminant analysis. The degree of genetic differentiation with geographical distance between all population pairs was investigated by Mantel tests. Results indicated that Pilgerodendron populations are highly monomorphic, probably reflecting past population bottlenecks and reduced gene flow. Southernmost populations tend to be the least genetically variable and were therefore probably more affected by glacial activity than northern ones. Populations located outside ice limits seem to have been isolated during the glacial period. The presence of centres of genetic diversity, together with the lack of a significant correlation between genetic and geographical distances and the absence of geographical patterns of allelic frequencies at most analysed alleles, may indicate that Pilgerodendron did not advance southward after the last glaciation from a unique northern refugium, but spread from several surviving populations in ice-free areas in Patagonia instead. [source] Slope development reconstruction at two sites in the Bohemian Forest MountainsEARTH SURFACE PROCESSES AND LANDFORMS, Issue 4 2010Filip Hartvich Abstract Although the Bohemian Forest is generally considered to be geomorphologically a rather stable region, there are localities which bear proof of a surprisingly high intensity of recent and even present day processes. A multidisciplinary methodology based on the analysis of slope systems was used for researching the present and past dynamics of two hillslopes. Using the results of direct dilatometric monitoring (the slow creep of rock blocks reached a maximum of 1,mm year,1), GPS field mapping, sedimentology, geodetic measurements and DEM analyses, several common rules have been observed for the behaviour of two model localities and a likely polycyclic development established, with recurrence of mass-wasting cycles related to material supply thresholds. Radiometric dating of debris flow activity showed that the cycles span approximately 4000 years. In this area of rather stable bedrock conditions, at least two factors that decrease slope stability need to act together to activate slope processes: i.e. a favourable structural predisposition (dip of major joint or bedding planes) and an increase in relief through either glaciation or river incision. Copyright © 2010 John Wiley & Sons, Ltd. [source] Meltwater discharge through the subglacial bed and its land-forming consequences from numerical experiments in the Polish lowland during the last glaciationEARTH SURFACE PROCESSES AND LANDFORMS, Issue 4 2009Jan A. Piotrowski Abstract Numerical experiments suggest that the last glaciation severely affected the upper lithosphere groundwater system in NW Poland: primarily its flow pattern, velocities and fluxes. We have simulated subglacial groundwater flow in two and three spatial dimensions using finite difference codes for steady-state and transient conditions. The results show how profoundly the ice sheet modifies groundwater pressure heads beneath and some distance beyond the ice margin. All model runs show water discharge at the ice forefield driven by ice-sheet-thickness-modulated, down-ice-decreasing hydraulic heads. In relation to non-glacial times, the transient 3D model shows significant changes in the groundwater flow directions in a regionally extensive aquifer ca. 90 m below the ice,bed interface and up to 40 km in front of the glacier. Comparison with empirical data suggests that, depending on the model run, only between 5 and 24% of the meltwater formed at the ice sole drained through the bed as groundwater. This is consistent with field observations documenting abundant occurrence of tunnel valleys, indicating that the remaining portion of basal meltwater was evacuated through a channelized subglacial drainage system. Groundwater flow simulation suggests that in areas of very low hydraulic conductivity and adverse subglacial slopes water ponding at the ice sole was likely. In these areas the relief shows distinct palaeo-ice lobes, indicating fast ice flow, possibly triggered by the undrained water at the ice,bed interface. Owing to the abundance of low-permeability strata in the bed, the simulated groundwater flow depth is less than ca. 200 m. Copyright © 2009 John Wiley & Sons, Ltd. [source] Stream geomorphology in a mountain lake district: hydraulic geometry, sediment sources and sinks, and downstream lake effectsEARTH SURFACE PROCESSES AND LANDFORMS, Issue 4 2007C. D. Arp Abstract Lakes are common in glaciated mountain regions and geomorphic principles suggest that lake modifications to water and sediment fluxes should affect downstream channels. Lakes in the Sawtooth Mountains, Idaho, USA, were created during glaciation and we sought to understand how and to what extent glacial morphology and lake disruption of fluxes control stream physical form and functions. First, we described downstream patterns in channel form including analyses of sediment entrainment and hydraulic geometry in one catchment with a lake. To expand on these observations and understand the role of glacial legacy, we collected data from 33 stream reaches throughout the region to compare channel form and functions among catchments with lakes, meadows (filled lakes), and no past or present lakes. Downstream hydraulic geometry relationships were weak for both the single catchment and regionally. Our data show that downstream patterns in sediment size, channel shape, sediment entrainment and channel hydraulic adjustment are explained by locations of sediment sources (hillslopes and tributaries) and sediment sinks (lakes). Stream reaches throughout the region are best differentiated by landscape position relative to lakes and meadows according to channel shape and sediment size, where outlets are wide and shallow with coarse sediment, and inlets are narrow and deep with finer sediment. Meadow outlets and lake outlets show similarities in the coarse-sediment fraction and channel capacity, but meadow outlets have a smaller fine-sediment fraction and nearly mobile sediment. Estimates of downstream recovery from lake effects on streams suggest 50 per cent recovery within 2,4 km downstream, but full recovery may not be reached within 20 km downstream. These results suggest that sediment sinks, such as lakes, in addition to sources, such as tributaries, are important local controls on mountain drainage networks. Copyright © 2006 John Wiley & Sons, Ltd. [source] Reach-scale channel geometry of a mountain riverEARTH SURFACE PROCESSES AND LANDFORMS, Issue 8 2004Ellen Wohl Abstract Mountain rivers can be subject to strong constraints imposed by changes in gradient and grain size supplied by processes such as glaciation and rockfall. Nonetheless, adjustments in the channel geometry and hydraulics of mountain rivers at the reach scale can produce discernible patterns analogous to those in fully alluvial rivers. Mountain rivers can differ in that imposed reach-scale gradient is an especially important control on reach-scale channel characteristics, as indicated by examination of North St Vrain Creek in Colorado. North St Vrain Creek drains 250 km2 of the Rocky Mountains. We used 25 study reaches within the basin to examine controls on reach-scale channel geometry. Variables measured included channel geometry, large woody debris, grain size, and mean velocity. Drainage area at the study reaches ranged from 2·2 to 245 km2, and gradient from 0·013 to 0·147 m m,1. We examined correlations among (1) potential reach-scale response variables describing channel bankfull dimension and shape, hydraulics, bedform wavelength and amplitude, grain size, ,ow resistance, standard deviation of hydraulic radius, and volume of large woody debris, and (2) potential control variables that change progressively downstream (drainage area, discharge) or that are likely to re,ect a reach-speci,c control (bed gradient). We tested the hypothesis that response variables correlate most strongly with local bed gradient because of the segmented nature of mountain channels. Results from simple linear regression analyses indicate that most response variables correlate best with gradient, although channel width and width/depth ratio correlate best with discharge. Multiple regression analyses using Mallow's Cp selection criterion and log-transformation of all variables produced similar results in that most response variables correlate strongly with gradient. These results suggest that the hypothesis is partially supported: channel bed gradient is likely to be a good predictor for many reach-scale response variables along mountain rivers, but discharge is also an important predictor for some response variables. Copyright © 2004 John Wiley & Sons, Ltd. [source] A rare landform: Yerköprü travertine bridges in the Taurids Karst Range, TurkeyEARTH SURFACE PROCESSES AND LANDFORMS, Issue 6 2002C. Serdar Bayari Abstract Two examples of travertine bridges are observed at 8 to 15 m above stream level in the Lower Zamanti Basin, Eastern Taurids, Turkey. Yerköprü-1 and Yerköprü-2 bridges are currently being deposited from cool karstic groundwaters with log PCO2 > 10,2 atm. The surface area and the total volume of travertine in Yerköprü-1 bridge are 4350 m2 and 40 000 m3, whereas the values for Yerköprü-2 are 2250 m2 and 20 000 m3, respectively. The interplay of hydrogeological structure, local topography, calcite-saturated hanging springs, algal activity and rapid downcutting in the streambed appear to have led to the formation of travertine bridges. Aeration through cascades and algal uptake causes efficient carbon dioxide evasion that enhances travertine formation. Algal curtains aid lateral development of travertine rims across the stream. Model calculations based on a hypothetical deposit in the form of a half-pyramid implied that lateral development should have occurred from both banks of the stream in the Yerköprü-1 bridge, whereas one-sided growth has been sufficient for Yerköprü-2. The height difference between travertine springs and the main stream appears to be a result of Pleistocene glaciation during which karstic base-level lowering was either stopped or slowed down while downcutting in the main stream continued. Copyright © 2002 John Wiley & Sons, Ltd. [source] Chalk micropalaeontology and the provenancing of middle pleistocene lowestoft formation till in eastern EnglandEARTH SURFACE PROCESSES AND LANDFORMS, Issue 9 2001P. R. Fish Abstract The distribution and deposits of British and Scandinavian Middle Pleistocene ice sheets in eastern England remain problematic. A new till provenancing technique based on Chalk micropalaeontology is described, with the object of refining understanding of the ice sheet which deposited the Lowestoft Formation till (Anglian/Elsterian) and its relationship to Scandinavian ice sheets. The technique involves extracting foraminifera from Chalk erratics and till matrix obtained from stratigraphically controlled till sections and comparing their micropalaeontology with that of Upper Cretaceous Chalk bedrock. Application to the Lowestoft Formation till of eastern England suggests that current models of ice-flow in this region require revision involving reinstatement of some earlier ideas. Chalk provenance data indicate an initial phase of glaciation, with ice streaming southwards across eastern England before fanning across East Anglia from the position of the Fen basin. This was followed by a later phase in which the main southward trajectory of ice-flow was located further east in the North Sea Basin, but again with ice fanning out across East Anglia. These ice-flow trajectories imply constraint of the British ice sheet by Scandinavian ice. Copyright © 2001 John Wiley & Sons, Ltd. [source] Atlantic salmon Salmo salar L., brown trout Salmo trutta L. and Arctic charr Salvelinus alpinus (L.): a review of aspects of their life historiesECOLOGY OF FRESHWATER FISH, Issue 1 2003A. Klemetsen Abstract ,,,Among the species in the family Salmonidae, those represented by the genera Salmo, Salvelinus, and Oncorhynchus (subfamily Salmoninae) are the most studied. Here, various aspects of phenotypic and life-history variation of Atlantic salmon Salmo salar L., brown trout Salmo trutta L., and Arctic charr Salvelinus alpinus (L.) are reviewed. While many strategies and tactics are commonly used by these species, there are also differences in their ecology and population dynamics that result in a variety of interesting and diverse topics that are challenging for future research. Atlantic salmon display considerable phenotypic plasticity and variability in life-history characters ranging from fully freshwater resident forms, where females can mature at approximately 10 cm in length, to anadromous populations characterised by 3,5 sea-winter (5SW) salmon. Even within simple 1SW populations, 20 or more spawning life-history types can be identified. Juveniles in freshwater can use both fluvial and lacustrine habitats for rearing, and while most smolts migrate to sea during the spring, fall migrations occur in some populations. At sea, some salmon undertake extensive oceanic migrations while other populations stay within the geographical confines of areas such as the Baltic Sea. At the other extreme are those that reside in estuaries and return to freshwater to spawn after spending only a few months at sea. The review of information on the diversity of life-history forms is related to conservation aspects associated with Atlantic salmon populations and current trends in abundance and survival. Brown trout is indigenous to Europe, North Africa and western Asia, but was introduced into at least 24 countries outside Europe and now has a world-wide distribution. It exploits both fresh and salt waters for feeding and spawning (brackish), and populations are often partially migratory. One part of the population leaves and feeds elsewhere, while another part stays as residents. In large, complex systems, the species is polymorphic with different size morphs in the various parts of the habitat. Brown trout feed close to the surface and near shore, but large individuals may move far offshore. The species exhibits ontogenetic niche shifts partly related to size and partly to developmental rate. They switch when the amount of surplus energy available for growth becomes small with fast growers being younger and smaller fish than slow growers. Brown trout is an opportunistic carnivore, but individuals specialise at least temporarily on particular food items; insect larvae are important for the young in streams, while littoral epibenthos in lakes and fish are most important for large trout. The sexes differ in resource use and size. Females are more inclined than males to become migratory and feed in pelagic waters. Males exploit running water, near-shore and surface waters more than females. Therefore, females feed more on zooplankton and exhibit a more uniform phenotype than males. The Arctic charr is the northernmost freshwater fish on earth, with a circumpolar distribution in the Holarctic that matches the last glaciation. Recent mtDNA studies indicate that there are five phylogeographic lineages (Atlantic, Arctic, Bering, Siberian and Acadian) that may be of Pleistocene origin. Phenotypic expression and ecology are more variable in charr than in most fish. Weights at maturation range from 3 g to 12 kg. Population differences in morphology and coloration are large and can have some genetic basis. Charr live in streams, at sea and in all habitats of oligotrophic lakes, including very deep areas. Ontogenetic habitat shifts between lacustrine habitats are common. The charr feed on all major prey types of streams, lakes and near-shore marine habitats, but has high niche flexibility in competition. Cannibalism is expressed in several cases, and can be important for developing and maintaining bimodal size distributions. Anadromy is found in the northern part of its range and involves about 40, but sometimes more days in the sea. All charr overwinter in freshwater. Partial migration is common, but the degree of anadromy varies greatly among populations. The food at sea includes zooplankton and pelagic fish, but also epibenthos. Polymorphism and sympatric morphs are much studied. As a prominent fish of glaciated lakes, charr is an important species for studying ecological speciation by the combination of field studies and experiments, particularly in the fields of morphometric heterochrony and comparative behaviour. [source] ESTIMATING A GEOGRAPHICALLY EXPLICIT MODEL OF POPULATION DIVERGENCEEVOLUTION, Issue 3 2007L. Lacey Knowles Patterns of genetic variation can provide valuable insights for deciphering the relative roles of different evolutionary processes in species differentiation. However, population-genetic models for studying divergence in geographically structured species are generally lacking. Since these are the biogeographic settings where genetic drift is expected to predominate, not only are population-genetic tests of hypotheses in geographically structured species constrained, but generalizations about the evolutionary processes that promote species divergence may also be potentially biased. Here we estimate a population-divergence model in montane grasshoppers from the sky islands of the Rocky Mountains. Because this region was directly impacted by Pleistocene glaciation, both the displacement into glacial refugia and recolonization of montane habitats may contribute to differentiation. Building on the tradition of using information from the genealogical relationships of alleles to infer the geography of divergence, here the additional consideration of the process of gene-lineage sorting is used to obtain a quantitative estimate of population relationships and historical associations (i.e., a population tree) from the gene trees of five anonymous nuclear loci and one mitochondrial locus in the broadly distributed species Melanoplus oregonensis. Three different approaches are used to estimate a model of population divergence; this comparison allows us to evaluate specific methodological assumptions that influence the estimated history of divergence. A model of population divergence was identified that significantly fits the data better compared to the other approaches, based on per-site likelihood scores of the multiple loci, and that provides clues about how divergence proceeded in M. oregonensis during the dynamic Pleistocene. Unlike the approaches that either considered only the most recent coalescence (i.e., information from a single individual per population) or did not consider the pattern of coalescence in the gene genealogies, the population-divergence model that best fits the data was estimated by considering the pattern of gene lineage coalescence across multiple individuals, as well as loci. These results indicate that sampling of multiple individuals per population is critical to obtaining an accurate estimate of the history of divergence so that the signal of common ancestry can be separated from the confounding influence of gene flow,even though estimates suggest that gene flow is not a predominant factor structuring patterns of genetic variation across these sky island populations. They also suggest that the gene genealogies contain information about population relationships, despite the lack of complete sorting of gene lineages. What emerges from the analyses is a model of population divergence that incorporates both contemporary distributions and historical associations, and shows a latitudinal and regional structuring of populations reminiscent of population displacements into multiple glacial refugia. Because the population-divergence model itself is built upon the specific events shaping the history of M. oregonensis, it provides a framework for estimating additional population-genetic parameters relevant to understanding the processes governing differentiation in geographically structured species and avoids the problems of relying on overly simplified and inaccurate divergence models. The utility of these approaches, as well as the caveats and future improvements, for estimating population relationships and historical associations relevant to genetic analyses of geographically structured species are discussed. [source] EVIDENCE FOR HISTORICAL INTROGRESSION ALONG A CONTACT ZONE BETWEEN TWO SPECIES OF CHAR (PISCES: SALMONIDAE) IN NORTHWESTERN NORTH AMERICAEVOLUTION, Issue 5 2002Z. Redenbach Abstract Phylogeographic analyses can yield valuable insights into the geographic and historical contexts of contact and hybridization between taxa. Two species of char (Salmonidae), Dolly Varden (Salvelinus malma) and bull trout (S. confluentus) have largely parapatric distributions in watersheds of northwestern North America. They are, however, sympatric in several localities and hybridization and some introgression occurs across a broad area of contact. We conducted a comparative phylogenetic analysis of Dolly Varden and bull trout to gain a historical perspective of hybridization between these species and to test for footprints of historical introgression. We resolved two major Dolly Varden mitochondrial DNA (mtDNA) clades (with 1.4,2.2% sequence divergence between haplotypes) that had different geographical distributions. Clade N is distributed across most of the range of Dolly Varden, from southern British Columbia through to the Kuril Islands in Asia. Clade S had a much more limited distribution, from Washington state, at the southern limit of the Dolly Varden range, to the middle of Vancouver Island. The distribution and inferred ages of the mtDNA clades suggested that Dolly Varden survived the Wisconsinan glaciation in a previously unsuspected refuge south of the ice sheet, and that Dolly Varden and bull trout were probably in continuous contact over most of the last 100,000 years. When bull trout were included in the phylogenetic analysis, however, the mtDNA of neither species was monophyletic: Clade S Dolly Varden clustered within the bull trout mtDNA clade. This pattern was discordant with two nuclear phylogenies produced (growth hormone 2 and rRNA internal transcribed sequence 1), in which Dolly Varden and bull trout were reciprocally monophyletic. This discordance between mtDNA- and nDNA-based phylogenies indicates that historical introgression of bull trout mtDNA into Dolly Varden occurred. Percent sequence divergence within these introgressed Dolly Varden (clade S) was 0.2,0.6%, implying that the introgression occurred prior to the most recent glaciation. Our analysis and other evidence of contact between divergent lineages in northwestern North America strongly suggests that the area may be the site of previously unsuspected suture zones of aquatic biotas. [source] SYNTHESIS: Evolutionary history of Pacific salmon in dynamic environmentsEVOLUTIONARY APPLICATIONS (ELECTRONIC), Issue 2 2008Robin S. Waples Abstract Contemporary evolution of Pacific salmon (Oncorhynchus spp.) is best viewed in the context of the evolutionary history of the species and the dynamic ecosystems they inhabit. Speciation was complete by the late Miocene, leaving c. six million years for intraspecific diversification. Following the most recent glacial maximum, large areas became available for recolonization. Current intraspecific diversity is thus the product of recent evolution overlaid onto divergent historical lineages forged during recurrent episodes of Pleistocene glaciation. In northwestern North America, dominant habitat features have been relatively stable for the past 5000 years, but salmon ecosystems remain dynamic because of disturbance regimes (volcanic eruptions, landslides, wildfires, floods, variations in marine and freshwater productivity) that occur on a variety of temporal and spatial scales. These disturbances both create selective pressures for adaptive responses by salmon and inhibit long-term divergence by periodically extirpating local populations and creating episodic dispersal events that erode emerging differences. Recent anthropogenic changes are replicated pervasively across the landscape and interrupt processes that allow natural habitat recovery. If anthropogenic changes can be shaped to produce disturbance regimes that more closely mimic (in both space and time) those under which the species evolved, Pacific salmon should be well-equipped to deal with future challenges, just as they have throughout their evolutionary history. [source] Groundwater biodiversity patterns in the Lessinian Massif of northern ItalyFRESHWATER BIOLOGY, Issue 4 2009DIANA M. P. GALASSI Summary 1. The distribution patterns of stygobiotic invertebrates were examined with a stratified sampling design at 197 sites selected among four hydrogeographic basins in the Lessinian Massif (northern Italy). The sites were approximately evenly distributed among four hydrogeological zones: unsaturated and saturated zone of karstic aquifers, and hyporheic and saturated zone of porous aquifers. 2. Outlying Mean Index (OMI) analysis which assesses deviation of habitat conditions from reference conditions, was used to evaluate the importance of 14 selected environmental variables in shaping groundwater biodiversity patterns in the region (total of 89 stygobiotic species). The measured variables explained 80% of the variability in the data set. 3. Sampling sites were distributed along the environmental gradients defined by OMI analysis. Significant differences were detected between karstic and porous site, as well as among sites located in the four hydrogeological zones. Differences among the four hydrogeographic basins were not observed. 4. Ordination of stygobiotic species along the environmental gradients was best explained by historical variables (mainly Würmian glaciation and age of the underlying geological formation), while variables related to hydrogeology (mainly pH, calcium concentration and habitat fragmentation) influenced species distributions in the hydrogeological zones. An Environmental Integrity Index and nitrate concentration were significantly correlated with altitude, but appeared not to play a significant role in determining stygobiotic biodiversity patterns at the regional scale. 5. Results of the OMI analysis were highly significant for all taxa, suggesting that stygobiotic species are sensitive to the environmental factors studied. Thirty-five species showed high habitat specialisation (OMI index > 10). These species were usually rare and endemic to the Lessinian Massif. Most of them were found in a single hydrogeological zone. 6. Quaternary glaciations appear not to have lowered stygobiotic species richness in the Lessinian Massif. This may be because of the marginal location of the region with respect to the Würmian glacier limit and because of extensive networks of fractures in the vadose zone of the karst, which may have allowed stygobionts to move deep down in the aquifers to seek refuge during surface freezing and to recolonise ancestral habitats after the glaciers retreated. [source] Geoarchaeology and late glacial landscapes in the western lake superior region, Central North AmericaGEOARCHAEOLOGY: AN INTERNATIONAL JOURNAL, Issue 1 2007Christopher L. Hill The transition from full glacial to interglacial conditions along the southern margin of the Laurentide ice sheet resulted in dramatic changes in landscapes and biotic habitats. Strata and landforms resulting from the Wisconsin Episode of glaciation in the area directly west of Lake Superior indicate a context for late Pleistocene biota (including human populations) connected to ice margins, proglacial lakes, and postglacial drainage systems. Late Glacial landscape features that have the potential for revealing the presence of Paleoindian artifacts include abandoned shorelines of proglacial lakes in the Superior and Agassiz basins and interior drainages on deglaciated terrains. The linkage between Late Pleistocene human populations and Rancholabrean fauna has yet to be demonstrated in the western Lake Superior region, although isolated remains of mammoth ( Mammuthus) have been documented, as well as fluted points assigned to Clovis, Folsom, and Holcombe-like artifact forms. Agate Basin and Hell Gap (Plano-type) artifacts also imply the presence of human groups in Late Glacial landscapes associated with the Agassiz and Superior basins. © 2007 Wiley Periodicals, Inc. [source] Late Wisconsinan Port Eliza Cave deposits and their implications for human coastal migration, Vancouver Island, CanadaGEOARCHAEOLOGY: AN INTERNATIONAL JOURNAL, Issue 4 2006M. Al-Suwaidi Sediments of Port Eliza Cave provide a record of the Last Glacial Maximum (LGM) on Vancouver Island that has important implications for human migration along the debated coastal migration route. Lithofacies changes from nonglacial diamict to glacial laminated silt and clay and till, then a return to nonglacial conditions with oxidized clay, colluvial block beds, and speleothems, along with radiocarbon and U/Th dates, define glacial,nonglacial transitions. Scanning electron microscope studies and clay mineralogy confirm that the laminated fines represent glaciation. Preglacial faunal evidence shows a diverse range from small species, including birds, fish, vole, and marmot, to larger species, such as mountain goat. Pollen data from the same unit show a cold, dry tundra environment with sparse trees. Deglaciation occurred prior to an age of 12.3 ka B.P. based on dated mountain goat bone. These data support the viability of the coastal migration route for humans prior to ,16 ka B.P. and then as early as ,13 ka B.P. © 2006 Wiley Periodicals, Inc. [source] The Ohalo II prehistoric camp (19.5 Ky): New evidence for environmental and tectonic changes at the Sea of GalileeGEOARCHAEOLOGY: AN INTERNATIONAL JOURNAL, Issue 5 2002Shmuel Belitzky Combined archaeological data, shore surveys, and aerial photos of submerged sediments in the Sea of Galilee provide new insights into environmental and tectonic events, their dating, and their impact on the Ohalo II prehistoric camp (ca. 19,500 yr B.P.) and its surroundings. The Ohalo II waterlogged campsite contains excellently preserved brush hut remains and other in situ features, all embedded in late Pleistocene lacustrine strata. The findings indicate relatively short occupation of the site, not more than months or several years at a time. The high quality in situ preservation of delicate organic materials, as well as the short occupation period, suggests a quick and gentle burial by fine sediments. The evident fast submergence (water level rise of the Sea of Galilee) could have been the result of climatic fluctuations towards the end of the last glaciation and/or small-scale tectonic subsidence. The site is located on a tectonic block formed in the western fault belt of the Dead Sea Rift. We present new evidence of post-occupational folding of the late Pleistocene strata and recent tilting and faulting. A westward tectonic tilt may have caused the blockage of the old Jordan River outlet after A. D. 1106. Excellent preservation of the fault traces to the east of the site is attributed to the young age of the displacement on the fault. The last displacement apparently post-dates the blockage of the old Jordan River. © 2002 Wiley Periodicals, Inc. [source] The age of Rubisco: the evolution of oxygenic photosynthesisGEOBIOLOGY, Issue 4 2007E. G. NISBET ABSTRACT The evolutionary history of oxygenesis is controversial. Form I of ribulose 1,5-bisphosphate carboxylase/oxygenase (Rubisco) in oxygen-tolerant organisms both enables them to carry out oxygenic extraction of carbon from air and enables the competitive process of photorespiration. Carbon isotopic evidence is presented from ~2.9 Ga stromatolites from Steep Rock, Ontario, Canada, ~2.9 Ga stromatolites from Mushandike, Zimbabwe, and ~2.7 Ga stromatolites in the Belingwe belt, Zimbabwe. The data imply that in all three localities the reef-building autotrophs included organisms using Form I Rubisco. This inference, though not conclusive, is supported by other geochemical evidence that these stromatolites formed in oxic conditions. Collectively, the implication is that oxygenic photosynthesizers first appeared ~2.9 Ga ago, and were abundant 2.7,2.65 Ga ago. Rubisco specificity (its preference for CO2 over O2) and compensation constraints (the limits on carbon fixation) may explain the paradox that despite the inferred evolution of oxygenesis 2.9 Ga ago, the Late Archaean air was anoxic. The atmospheric CO2:O2 ratio, and hence greenhouse warming, may reflect Form I Rubisco's specificity for CO2 over O2. The system may be bistable under the warming Sun, with liquid oceans occurring in either anoxic (H2O with abundant CH4 plus CO2) or oxic (H2O with more abundant CO2, but little CH4) greenhouse states. Transition between the two states would involve catastrophic remaking of the biosphere. Build-up of a very high atmospheric inventory of CO2 in the 2.3 Ga glaciation may have allowed the atmosphere to move up the CO2 compensation line to reach stability in an oxygen-rich system. Since then, Form I Rubisco specificity and consequent compensation limits may have maintained the long-term atmospheric disproportion between O2 and CO2, which is now close to both CO2 and O2 compensation barriers. [source] THE OMISSION OF CONFLICTING EVIDENCE FROM THE PAPER BY GOLLEDGE ET AL. (2008)GEOGRAFISKA ANNALER SERIES A: PHYSICAL GEOGRAPHY, Issue 4 2009DEREK FLINN ABSTRACT. In their paper on the last glaciation of Shetland, Golledge et al. (2008) concluded on the basis of a remote sensing study that during that period Shetland had been overrun by ice from Scandinavia. Since the method of study they used does not reveal the sense of the direction in which the ice flowed and since they ignored earlier ground-based work involving striations which do reveal the sense of direction of ice flow their conclusions have no scientific basis. [source] GLACIATION OF MT ALLEN, STEWART ISLAND (RAKIURA): THE SOUTHERN MARGIN OF LGM GLACIATION IN NEW ZEALANDGEOGRAFISKA ANNALER SERIES A: PHYSICAL GEOGRAPHY, Issue 2 2009MARTIN S. BROOK ABSTRACT. The origin of two ridges on the eastern slopes of Mt Allen, southern Stewart Island, has remained equivocal, with differences of opinion over the exact process-mechanisms of formation. A variety of approaches was used to test a number of possible hypotheses about the origin of the ridges. These include topographic and spatial positioning, geomorphology, sedimentology and palaeoclimatological extrapolations to reconstruct two small former cirque glaciers with equilibrium line altitudes (ELAs) of c. 600 m. It would appear the two ridges reflect a glacial origin, the glaciers interpreted as forming during the Last Glacial Maximum (LGM) in New Zealand. Whilst glaciation during this time (18,19 ka) was extensive in the Southern Alps, the restricted nature of glaciation on Mt Allen suggests the low altitude restricted glaciation to niche sites on the lee side of upland areas. [source] THE LAST GLACIATION OF SHETLAND, NORTH ATLANTICGEOGRAFISKA ANNALER SERIES A: PHYSICAL GEOGRAPHY, Issue 1 2008N.R. GOLLEDGE ABSTRACT. Evidence relating to the extent, dynamics, and relative chronology of the last glaciation of the Shetland Islands, North Atlantic, is presented here, in an attempt to better illuminate some of the controversies that still surround the glacial history of the archipelago. We appraise previous interpretations and compare these earlier results with new evidence gleaned from the interpretation of a high resolution digital terrain model and from field reconnaissance. By employing a landsystems approach, we identify and describe three quite different assemblages of landscape features across the main islands of Mainland, Yell and Unst. Using the spatial interrelationship of these landsystems, an assessment of their constituent elements, and comparisons with similar features in other glaciated environments, we propose a simple model for the last glaciation of Shetland. During an early glacial phase, a coalescent British and Scandinavian ice sheet flowed approximately east to west across Shetland. The terrestrial land-forms created by this ice sheet in the north of Shetland suggest that it had corridors of relatively fast-flowing ice that were partially directed by bed topography, and that subsequent deglaciation was interrupted by at least one major stillstand. Evidence in the south of Shetland indicates the growth of a local ice cap of restricted extent that fed numerous radial outlet glaciers during, or after, ice-sheet deglaciation. Whilst the absolute age of these three landsystems remains uncertain, these new geo-morphological and palaeoglaciological insights reconcile many of the ideas of earlier workers, and allow wider speculation regarding the dynamics of the former British ice sheet. [source] THE EASTERN MARGIN OF GLACIATION IN THE BRITISH ISLES DURING THE YOUNGER DRYAS: THE BIZZLE CIRQUE, SOUTHERN SCOTLANDGEOGRAFISKA ANNALER SERIES A: PHYSICAL GEOGRAPHY, Issue 3 2006STEPHAN HARRISON ABSTRACT. Geomorphological and sedimentological evidence of former glaciation in the Bizzle valley in the Cheviot Hills of northern England and southern Scotland was used to reconstruct the dimensions of a small topographically constrained glacier with an equilibrium line altitude (ELA) of 535 m. This was interpreted as having formed during Younger Dryas cooling; this is the only glacier to have been described from the area and is the most easterly site of Younger Dryas glaciation in the British Isles. Whilst glaciation at this time was extensive in the Lake District to the southwest, the restricted nature of Cheviot ice cover suggests that a steep west,east precipitation gradient existed in this region during the Younger Dryas. [source] Quantifying the erosional impact of the Fennoscandian ice sheet in the Torneträsk,Narvik corridor, northern Sweden, based on cosmogenic radionuclide dataGEOGRAFISKA ANNALER SERIES A: PHYSICAL GEOGRAPHY, Issue 3-4 2002Arjen P. Stroeven Despite spectacular landform evidence of a dominant role for glacial action in shaping landscapes under former northern hemisphere ice sheets, there is little quantitative evidence for rates and patterns of erosion associated with specific glaciations. Here we use cosmogenic nuclide data to assess rates of subglacial erosion underneath the Fennoscandian ice sheet. By testing whether there are remnant nuclide concentrations in samples taken from sites that include both relict areas and features and landscapes typically associated with vigorous glacial erosion, we can constrain the level and pattern of modification that resulted from the last glaciation. Cosmogenic 10Be and 36Cl data from the Torneträsk region confirm the temporal and spatial variability of glacial erosion suggested by geomorphological mapping. At some sites, glacial erosion estimates in what appear to be heavily scoured areas indicate erosion of only c. 2 ± 0.4 m of bedrock, based on cosmogenic nuclide inheritance. This implies that the generation of severely scoured terrain in this study area required multiple glaciations. The overall modification produced by ice sheets along glacial corridors may be more restricted than previously thought, or may have occurred preferentially during earlier Quaternary glacial periods. [source] Age and Origin of Upland block Fields on Melville Peninsula, Eastern Canadian ArcticGEOGRAFISKA ANNALER SERIES A: PHYSICAL GEOGRAPHY, Issue 4 2000Lynda A. Dredge Block fields have developed on gently graded uplands of granite and gneiss on central and southern Melville Peninsula. The location of block fields is not controlled by elevation, but rather by areas covered by cold,based ice during the last glaciation. Block fields consist either of angular boulders, sorted circles 3,4 m across having blocky rims and central areas of weathered grus, concentrations of openwork boulders, or in the southeast, of immature bouldery till. The block fields are primarily relict features predating the last glaciation, produced by weakening of bedrock by weathering along joints, followed by frost heave, although some block fields have been modified by meltwater from regional glacier down,wasting. Frost processes are active in the modern environment but the extent of riven bedrock and the size of recent patterned ground forms on postglacial surfaces are insufficient to account for the forms in the block fields. [source] The diversification and extinction of Doushantuo-Pertatataka acritarchs in South China: causes and biostratigraphic significanceGEOLOGICAL JOURNAL, Issue 3-4 2007Zhou Chuanming Abstract The Ediacaran Period immediately follows the last Cryogenian glaciation,the ,635,Ma Marinoan or Nantuo glaciation, and it is also punctuated by another brief glaciation,the ,582,Ma Gaskiers glaciation. It is possible that these glaciations may have had significant impact on Ediacaran biological evolution (e.g. the appearance or disappearance of Doushantuo-Pertatataka acritarchs). Alternative hypotheses propose that the diversification of Doushantuo-Pertatataka acritarchs was caused by the Acraman bolide impact or by emerging eumetazoans. To test these hypotheses, high-resolution geochronological and biostratigraphic data are required. The Doushantuo Formation in South China, radiometrically constrained between ,635 and ,551,Ma, has the potential to clarify the global picture of early-middle Ediacaran evolution. Here we present preliminary biostratigraphic data from the Doushantuo Formation in the East Yangtze Gorges area and new ,13C chemostratigraphic data from the Doushantuo Formation at Weng'an. These and previously published palaeontological data, aided by the tool of ,13C chemostratigraphy, indicate that the biostratigraphic record of the Doushantuo Formation is locally sensitive to the availability of specific taphonomic windows. In the East Yangtze Gorges area, Doushantuo-Pertatataka acritarchs first appeared shortly after the termination of the Nantuo glaciation and gradually diversified throughout the Doushantuo Formation. At Weng'an, however, such acritarchs first appear,abruptly and in much greater diversity,in phosphorite of the upper Doushantuo Formation, immediately above a subaerial exposure surface. Thus, the biostratigraphic pattern in the East Yangtze Gorges area permits, whereas that at Weng'an apparently disallows, a causal relationship between the Nantuo glaciation and the diversification of Doushantuo-Pertatataka acritarchs. We conclude that the biostratigraphic record is incomplete at Weng'an, where the early Ediacaran evolutionary history is not preserved. The South China data indicate that special attention has to be paid to taphonomic and palaeoenvironmental analysis before extrapolating local and regional biostratigraphic ranges to make global interpretations. It is less clear when Doushantuo-Pertatataka acritarchs disappeared. Previous investigators have variously suggested that they disappeared before, at, or after, the Gaskiers glaciation. These hypotheses are difficult to test because of the lack of sedimentary evidence for the Gaskiers glaciation in South China and other regions (e.g. South Australia) where Doushantuo-Pertatataka acritarchs are abundant. In Australia, Doushantuo-Pertatataka acritarchs are thought to have experienced a sharp decline after the Egan glaciation, which may well be equivalent to the Gaskiers glaciation. If true, then Doushantuo-Pertatataka acritarchs are largely restricted to the interval between the Nantuo and Gaskiers glaciations. This conclusion allows us to place constraints on the possible causes of the diversification and extinction of Doushantuo-Pertatataka acritarchs and has important implications for the biostratigraphic significance of Doushantuo-Pertatataka acritarchs. Copyright © 2006 John Wiley & Sons, Ltd. [source] Facies development, depositional settings and sequence stratigraphy across the Ordovician,Silurian boundary: a new perspective from the Barrandian area of the Czech RepublicGEOLOGICAL JOURNAL, Issue 2 2006torch Abstract The Hirnantian and Llandovery sedimentary succession of the Barrandian area has been assigned to middle and outer clastic-shelf depositional settings, respectively. Deposition was influenced by the remote Gondwanan glaciation and subsequent, long-persisting, post-glacial anoxia triggered by a current-driven upwelling system. High-resolution graptolite stratigraphy, based upon 19 formally defined biozones,largely interval zones,and five subzones, enabled a detailed correlation between 42 surface sections and boreholes, and enabled linking of the sedimentary record, graptoloid fauna dynamics, organic-content fluctuations and spectral gamma-ray curves. The Hirnantian and Llandovery succession has been subdivided into four biostratigraphically dated third-order sequences (units 1,4). Time,spatial facies distribution recorded early and late Hirnantian glacio-eustatic sea-level lowstands separated by a remarkable mid-Hirnantian rise in sea-level. A major part of the post-glacial sea-level rise took place within the late Hirnantian. The highstand of Unit 2 is apparently at the base of the Silurian succession. Short-term relative sea-level drawdown and a third-order sequence boundary followed in the early Rhuddanian upper acuminatus Zone. Early Aeronian and late Telychian sea-level highstands and late Aeronian drawdown of likely eustatic origin belong to units 3 and 4. Sea-level rise culminated in the late Telychian, which may also be considered as a highstand episode of a second-order Hirnantian,early Silurian cycle. Facies and sequence-stratigraphic analysis supports recent interpretations on nappe structures in the core part of the Ordovician,Middle Devonian Prague Synform of the Barrandian. Copyright © 2006 John Wiley & Sons, Ltd. [source] | ||||||||||||||||||||||||||||