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Glacial Period (glacial + period)
Kinds of Glacial Period Selected AbstractsPatterns of isozyme variation as indicators of biogeographic history in Pilgerodendron uviferum (D. Don) FlorínDIVERSITY AND DISTRIBUTIONS, Issue 2 2002A. C. Premoli Abstract. The effects of Pleistocene glaciations on the genetic characteristics of the most austral conifer in the world, Pilgerodendron uviferum, were analysed with specific reference to the hypothesis that the species persisted locally in ice-free areas in temperate South America. It was expected that genetic variation would decrease with latitude, given that ice fields were larger in southern Patagonia and thus refugia were probably located towards the northern distributional limit of the species as suggested by the fossil record. In addition, an increase in among-population genetic divergence was expected with increasing distance to putative glacial refugia. We examined the relationship between location and within-population variability indices of 20 Pilgerodendron populations derived from isozyme analyses. We analysed possible refugia hypotheses by the distribution of allele frequencies using multivariate discriminant analysis. The degree of genetic differentiation with geographical distance between all population pairs was investigated by Mantel tests. Results indicated that Pilgerodendron populations are highly monomorphic, probably reflecting past population bottlenecks and reduced gene flow. Southernmost populations tend to be the least genetically variable and were therefore probably more affected by glacial activity than northern ones. Populations located outside ice limits seem to have been isolated during the glacial period. The presence of centres of genetic diversity, together with the lack of a significant correlation between genetic and geographical distances and the absence of geographical patterns of allelic frequencies at most analysed alleles, may indicate that Pilgerodendron did not advance southward after the last glaciation from a unique northern refugium, but spread from several surviving populations in ice-free areas in Patagonia instead. [source] Investigations regarding Alpine talus slopes using ground-penetrating radar (GPR) in the Bavarian Alps, GermanyEARTH SURFACE PROCESSES AND LANDFORMS, Issue 10 2001Oliver Sass Abstract The applicability of ground-penetrating radar (GPR) for the investigation of loose debris was tested at two sites (Viererkar and Zugspitzplatt). A pulseEKKO 100 GPR system equipped with 25 MHz antennae was utilized. The aim of the investigation was to record the base of the debris layer, and thereby acquire an estimation of the backweathering rates of the adjacent rockwalls. The study areas are situated in the Northern Alps near the German,Austrian border. The sites are characterized by steep limestone rockwalls and extensive talus accumulations. A total of six profiles was surveyed. The method is suitable and effective for a quick survey in this dry, high-ohmic substrate. The GPR system was able to deliver information about the subsurface stratigraphy to c. 70 m depth. The boundary line to the bedrock was discovered , depending upon the profile surveyed ,5 to 25 m below the surface. The base of the debris material sometimes shows no distinct reflection. Buried features (V-shaped furrows, zones overdeepened by ice action, geological structures) could be detected. Arched structures well below the talus,bedrock interface can be interpreted as drainage systems in the karstic bedrock. A thick scree layer of Late Glacial age was separated from a thinner layer on the talus surface, which was related to the Holocene. The backweathering rates were fixed by a calculation of talus volume to c. 100 mm/103 a during the Holocene (Viererkar) and 150,300 mm/103 a (Zugspitzplatt). The detrital formation in north-exposed sites is twice as intense as in south-exposed sites. These results match the rates of recent rockfall in the same area of investigation. The calculated backweathering for the late glacial period is 150,730 mm/103 a. The magnitude of the calculated rockwall retreat lies well within the range of previous measurements. The discrepancy between some weathering rates highlights the fact that recent and past relief formation must be differentiated. Otherwise recent removal rates may be overestimated. Copyright © 2001 John Wiley & Sons, Ltd. [source] A lock-in model for the complex Matuyama-Brunhes boundary record of the loess/palaeosol sequence at Lingtai (Central Chinese Loess Plateau)GEOPHYSICAL JOURNAL INTERNATIONAL, Issue 2 2003S. Spassov SUMMARY In most marine sedimentary records, the Matuyama-Brunhes boundary (MBB) has been found in interglacial oxygen isotope stage 19. In the magnetostratigraphic records of most Chinese loess/palaeosol profiles the MBB is located in loess layer L8, which was deposited during a glacial period. The MBB at Lingtai (central Chinese Loess Plateau) also occurs in L8 and is characterized by multiple polarity flips. The natural remanent magnetization is mainly carried by two coexisting components. The higher coercivity (harder) component dominates in loess layers and is thought to be of detrital origin. The lower coercivity (softer) component prevails in palaeosols and was most probably formed in situ by (bio-)chemical processes. A lock-in model for the Lingtai MBB record has been developed by extending the lithologically controlled PDRM model of Bleil & von Dobeneck (1999). It assumes two lock-in zones. The NRM of the magnetically harder component is physically locked by consolidation shortly after loess deposition, whereas the softer component is formed at greater depth by pedogenesis and acquires a chemical remanent magnetization of younger age. At polarity boundaries, grains carrying reversed and normal directions may therefore occur together within a single horizon. The model uses ARM coercivity spectra to estimate the relative contributions of the two components. It is able to explain the observed rapid multiple polarity flips and low magnetization intensities as well as the stratigraphic shift of the Lingtai MBB with respect to the marine records. [source] The importance of low atmospheric CO2 and fire in promoting the spread of grasslands and savannasGLOBAL CHANGE BIOLOGY, Issue 7 2003W. J. BOND Abstract The distribution and abundance of trees can be strongly affected by disturbance such as fire. In mixed tree/grass ecosystems, recurrent grass-fuelled fires can strongly suppress tree saplings and therefore control tree dominance. We propose that changes in atmospheric [CO2] could influence tree cover in such metastable ecosystems by altering their postburn recovery rates relative to flammable herbaceous growth forms such as grasses. Slow sapling recovery rates at low [CO2] would favour the spread of grasses and a reduction of tree cover. To test the possible importance of [CO2]/fire interactions, we first used a Dynamic Global Vegetation Model (DGVM) to simulate biomass in grassy ecosystems in South Africa with and without fire. The results indicate that fire has a major effect under higher rainfall conditions suggesting an important role for fire/[CO2] interactions. We then used a demographic model of the effects of fire on mesic savanna trees to test the importance of grass/tree differences in postburn recovery rates. We adjusted grass and tree growth in the model according to the DGVM output of net primary production at different [CO2] relative to current conditions. The simulations predicted elimination of trees at [CO2] typical of the last glacial period (180 ppm) because tree growth rate is too slow (15 years) to grow to a fire-proof size of ca. 3 m. Simulated grass growth would produce an adequate fuel load for a burn in only 2 years. Simulations of preindustrial [CO2] (270 ppm) predict occurrence of trees but at low densities. The greatest increase in trees occurs from preindustrial to current [CO2] (360 ppm). The simulations are consistent with palaeo-records which indicate that trees disappeared from sites that are currently savannas in South Africa in the last glacial. Savanna trees reappeared in the Holocene. There has also been a large increase in trees over the last 50,100 years. We suggest that slow tree recovery after fire, rather than differential photosynthetic efficiencies in C3 and C4 plants, might have been the significant factor in the Late Tertiary spread of flammable grasslands under low [CO2] because open, high light environments would have been a prerequisite for the spread of C4 grasses. Our simulations suggest further that low [CO2] could have been a significant factor in the reduction of trees during glacial times, because of their slower regrowth after disturbance, with fire favouring the spread of grasses. [source] Putative glacial refugia of Cedrus atlantica deduced from Quaternary pollen records and modern genetic diversityJOURNAL OF BIOGEOGRAPHY, Issue 7 2009R. Cheddadi Abstract Aim, To investigate the impact of past environmental changes on Cedrus atlantica and its current genetic diversity, and to predict its future distribution. Location, Morocco, Algeria and Tunisia. Methods, Eleven fossil pollen records from these three countries were used to locate putative glacial refugia and to reconstruct past climate changes. A mechanistic vegetation distribution model was used to simulate the distribution of C. atlantica in the year 2100. In addition, a genetic survey was carried out on modern Moroccan C. atlantica. Results, Pollen records indicate that Cedrus was present during the last glacial period, probably in scattered refugia, in Tunisia, Algeria and Morocco. In the Tunisian and Algerian sites, cedar expanded during the late glacial and the early Holocene, then disappeared after c. 8000 yr bp. Reconstructed mean annual precipitation and January temperature show that the last glacial period in Morocco was cooler by 10,15°C and drier by c. 300,400 mm year,1 than the climate today. Modern chloroplast microsatellites of 15 C. atlantica populations in Morocco confirm the presence of multiple refugia and indicate that cedar recolonized the Moroccan mountains fairly recently. Model simulation indicates that by the year 2100 the potential distribution of C. atlantica will be much restricted with a potential survival area located in the High Atlas. Main conclusions, Environmental changes in northern Africa since the last glacial period have had an impact on the geographical distribution of C. atlantica and on its modern genetic diversity. It is possible that by the end of this century C. atlantica may be unable to survive in its present-day locations. To preserve the species, we suggest that seedlings from modern C. atlantica populations located in the High Atlas mountains, where a high genetic diversity is found, be transplanted into the western High Atlas. [source] Glacial survival or late glacial colonization?JOURNAL OF BIOGEOGRAPHY, Issue 12 2006Phylogeography of the root vole (Microtus oeconomus) in north-west Norway Abstract Aim, It has been proposed that the root vole subspecies, Microtus oeconomus finmarchicus, survived the last glacial period on islands on the north-west coast of Norway. The Norwegian island of Andøya may have constituted the only site with permanent ice-free conditions. Geological surveys and fossil finds from Andøya demonstrate that survival throughout the last glacial maximum was probably possible for some plants and animals. In this study we aim to infer the recent evolutionary history of Norwegian root vole populations and to evaluate the glacial survival hypothesis. Methods, DNA sequence variation in the mitochondrial cytochrome b gene was studied in 46 root voles from 19 localities. Location, Northern Fennoscandia and north-west Russia with a focus on islands on the north-west coast of Norway. Results The phylogeographical analyses revealed two North European phylogroups labelled ,Andøya' and ,Fennoscandia'. The Andøya phylogroup contained root voles from the Norwegian islands of Andøya, Ringvassøya and Reinøya and two localities in north-west Russia. The Fennoscandian phylogroup encompassed root voles from the three Norwegian islands of Kvaløya, Håkøya and Arnøya and the remaining specimens from Norway, northern Sweden and Finland. Nucleotide diversity within the Andøya and Fennoscandian phylogroups was similar, ranging from 0.5% to 0.7%. Main conclusions Both our genetic data and previously published morphological data are consistent with in situ glacial survival of root voles on Andøya during the last glacial maximum. However, the level of genetic diversity observed in the extant island populations, the past periods of severe climatic conditions on Andøya and the ecology of the root vole are somewhat difficult to reconcile with this model. A biogeographical scenario involving late glacial recolonization along the northern coasts of Russia and Norway therefore represents a viable alternative. Our results demonstrate that complex recolonization and extinction histories can generate intricate phylogeographical patterns and relatively high levels of genetic variation in northern populations. [source] Evidence for a lacustrine faunal refuge in the Larsemann Hills, East Antarctica, during the Last Glacial MaximumJOURNAL OF BIOGEOGRAPHY, Issue 7 2006Louise Cromer Abstract Aim, There is no previous direct evidence for the occurrence of lacustrine refuges for invertebrate fauna in Antarctica spanning the Last Glacial Maximum (LGM). In the absence of verified LGM lacustrine refuges many species are believed to result from Holocene dispersal from sub-Antarctic islands and continents further north. If freshwater lake environments were present throughout the LGM, extant freshwater species may have been associated with Antarctica prior to this glacial period. This study looked at faunal microfossils in a sediment core from an Antarctic freshwater lake. This lake is unusual in that, unlike most Antarctic lakes, the sediment record extends to c. 130,000 yr bp, i.e. prior to the LGM. Location, Lake Reid, Larsemann Hills, East Antarctica (76°23, E; 69°23, S). Methods, Palaeofaunal communities in Lake Reid were identified through examination of faunal microfossils in a sediment core that extended to c. 130,000 yr bp. Results, Ephippia and mandibles from the cladoceran Daphniopsis studeri and loricae of the rotifer Notholca sp. were found at all depths in the sediment, indicating that these two species have been present in the lake for up to 130,000 years. Copepod mandibles were also present in the older section of the core, yet were absent from the most recent sediments, indicating extinction of this species from Lake Reid during the LGM. Main conclusion, The presence of D. studeri and Notholca sp. microfossils throughout the entire Lake Reid core is the first direct evidence of a glacial lacustrine refugium for invertebrate animals in Antarctica, and indicates the presence of a relict fauna on the Antarctic continent. [source] Patterns and determinants of shorebird species richness in the circumpolar ArcticJOURNAL OF BIOGEOGRAPHY, Issue 3 2005Sara S. Henningsson Abstract Aim, The intention with this study was first to investigate and describe the broad-scale geographical patterns of species richness of breeding shorebirds (Charadriiformes; families: Charadriidae, Scolopacidae and Haematopodidae) throughout the arctic tundra biome. Secondly, after compensating for the positive relationship between net primary productivity (NPP) and species richness, the relative importance of additional ecological and historical variables for species richness was investigated. The main variables considered are NPP, length of snow- and ice-free season, accessibility of regions depending on migratory flyway systems, tundra area at Pleistocene (120 and 20,18 ka bp) and Holocene (8 ka bp) times, and tundra area at present. Methods, Information on shorebird species breeding distributions was compiled from distribution atlases and species accounts. The breeding distributions of shorebirds with ranges partly or completely in the Arctic (a total of 50 species) were mapped in ArcView 3.2 to create a raster grid layer of shorebird species richness at a 1° latitude × longitude resolution. The total and mean species richness value was calculated per each 10° of longitude sector of the Arctic. The relationships between species richness and the different climatic and environmental variables were analysed on the basis of this sector-wise division of the arctic tundra. The influence of each variable on species richness was investigated using univariate and multivariate analyses (multivariate linear regression and general linear model). Results, We found that patterns of breeding shorebird species richness in the Arctic tundra biome are to a large degree determined by the NPP, the length of the snow- or ice-free season, the diversity of migratory flyways, as well as the historical extent of tundra habitat area during the maximum cooling of the last glacial period. Essentially, two main regions are distinguishable in the circumpolar Arctic regarding shorebird community richness. These are a species-rich Beringia-centred region and a species-poor Atlantic-centred region. Main conclusions, The underlying explanations to these major trends may primarily be attributed to factors that operate at present through accessibility of areas from contemporary migration flyways, as well as processes that operated in the past during and after the last glacial cycle. The most prominent influence on the shorebird diversity was found for NPP in combination with the diversity of migratory flyways. These flyways provide the links between breeding and wintering resources, often separated by huge distances, and the geographical and ecological conditions associated with the shorebirds' migration seem to be of particular importance for their breeding diversity in different sectors of circumpolar tundra. [source] Genetic divergence between sympatric Arctic charr Salvelinus alpinus morphs in Gander Lake, Newfoundland: roles of migration, mutation and unequal effective population sizesJOURNAL OF FISH BIOLOGY, Issue 8 2008D. Gomez-Uchida A suite of 10 microsatellite loci was used to examine genetic divergence between two sympatric morphs of Arctic charr Salvelinus alpinus (,dark' and ,pale') inhabiting Gander Lake, Newfoundland. Results can be summarized as follows: (1) the morphs are strongly reproductively isolated , gene flow,migration estimates were consistently low in long and short-term evolutionary timescales of analysis; (2) intermorph divergence based on allele size (RST) was significantly larger than those based on allele state (,) implying a cumulative effect of stepwise-like mutations; (3) historical (coalescent) and current (linkage disequilibrium) point estimates of effective population size (Ne) were consistently higher for dark than for pale S. alpinus. The first and second findings lend support to the hypothesis that divergence between forms may have preceded the last glacial period (ending c. 12 000 years bp). The third finding argues for significant differences in habitat quantity and quality between morphs, which were emphasized in a previous study. Overall, these analyses underscore the importance of genetic assessment and monitoring in the conservation of fish diversity, with emphasis on ,rare' or under-represented forms among temperate species pairs. [source] Chernozem,Soil of the Year 2005JOURNAL OF PLANT NUTRITION AND SOIL SCIENCE, Issue 6 2005Manfred Altermann Abstract The proclamation of the "Soil of the Year" was made for the first time in Germany in 2005 on occasion of the World Soil Day. Chernozems were selected for this purpose. In this paper an overview of these groups of soils is given. Chernozems are concentrated in the drought region of Central Germany. A standard profile from the core area of Chernozems developed from loess is presented with comprehensive laboratory analysis. Chernozems developed primarily upon carbonatic loess substrates under summer-dry climatic conditions in an open park-like landscape with isolated forest stands. The development of Chernozems began as early as the late glacial period, and they were fully developed by the Atlantikum age. The far-reaching, uniformly thick humus horizons indicate substrate differences in the loess cover, which are partly the result of bioturbation. Within Germany, Chernozems and Chernozem-like soils make up approx. 3% of the surface area and 5% (approx. 11,000 km2) of the arable land. The results of the Static Fertilization Experiment in Bad Lauchstädt, founded in 1902, clarify the high value of Chernozem for biomass production and the environment. Each loss due to erosion or decrease in surface area reduces the fulfillment of soil ecological functions of the soils and is comparable to a loss of animal and plant species. Therefore, soil scientists and the results of soil research must be more comprehensively implemented for soil preservation, protection, and politics. For acceptance of these goals among the general public and the political-decision makers, the campaign "Soil of the Year" should give some thought-provoking impulses. Schwarzerde , Boden des Jahres 2005 Anlässlich des Weltbodentages wurde in Deutschland für 2005 mit der Schwarzerde erstmalig ein ,Boden des Jahres" proklamiert. Damit soll in der Bevölkerung und bei politischen Entscheidungsträgern ein stärkeres Bewusstsein für den Boden und ein höheres Engagement für den Bodenschutz angeregt werden. Im Beitrag wird ein Überblick über diese Bodengruppe gegeben und ein Standardprofil aus dem Kerngebiet der Schwarzerden aus Löss (Mitteldeutsches Trockengebiet) mit umfassenden Laboranalysen exemplarisch präsentiert. Schwarzerden entwickelten sich vorwiegend auf kalkreichen Lössen unter sommertrockenen Klimabedingungen in einer offenen parkähnlichen Landschaft mit Waldinseln. Die Entstehung der Schwarzerden setzte bereits im Spätglazial ein, und im Atlantikum waren sie voll entwickelt. Die weiträumig gleiche Mächtigkeit der Humushorizonte zeichnet primäre Substratunterschiede in der Lössdecke nach; sie sind nicht nur das Ergebnis einer Bioturbation. In Deutschland nehmen die Schwarzerden und schwarzerdeähnlichen Böden etwa 3 % der Bodenfläche bzw. 5 % (ca. 11.000 km2) der landwirtschaftlichen Nutzfläche ein. Die Ergebnisse des seit 1902 bestehenden Statischen Düngungsversuchs Bad Lauchstädt verdeutlichen den hohen Wert der Schwarzerden für Biomasseproduktion und Umwelt. Jeder Verlust durch Erosion oder Flächenentzug mindert die Erfüllung ökologischer Funktionen der Böden und ist dem Artenverlust von Tieren und Pflanzen gleichzustellen. In der Bodenpolitik müssen deshalb die Ergebnisse der Bodenforschung zum Erhalt und Schutz unserer Böden umfassender als bisher umgesetzt und Bodenwissenschaftler stärker in politische Entscheidungen eingebunden werden. Für die Akzeptanz und Umsetzung dieser Ziele in der Öffentlichkeit soll der ,Boden des Jahres" Impulse geben. [source] Holocene bipolar climate seesaw: possible subtle evidence from the deep North East Atlantic Ocean?,JOURNAL OF QUATERNARY SCIENCE, Issue 3 2010Mark A. Maslin Abstract The occurrence of a millennial-scale bipolar climate seesaw has been documented in detail for the last glacial period and Termination. There is, however, debate whether it occurs during interglacials and if it does what influence it could have on future climate. We present here new evidence from a North East Atlantic Ocean deep-sea core which supports the hypothesis for a Holocene bipolar climate seesaw. BENGAL Site 13078#16, from the Porcupine Abyssal Plain, is 4844,m deep and situated at the North Atlantic Deep Water and Antarctic Bottom Water (AABW) interface. Planktic foraminiferal fragment accumulation rate data at this site is an indicator of coarse carbonate dissolution, which is highly sensitive to the incursion of under-saturated AABW. Five dissolution peaks have been identified, which seem to occur approximately 500 a after each of the North Atlantic 'Bond' ice rafting pulses, suggesting a subsequent subtle shallowing of AABW. This indicates a possible lagged climatic link between North East Atlantic surface water conditions and AABW production in the Southern Ocean during the Holocene. This provides the first tentative evidence that there was a Holocene bipolar climate seesaw and that the deep ocean was involved. This study also suggests that extremely sensitive locations need to be sought as the Holocene bipolar climate seesaw seems to be very subtle compared with its glacial counterparts. Copyright © 2009 John Wiley & Sons, Ltd. [source] Sub-millennial climate shifts in the western Mediterranean during the last glacial period recorded in a speleothem from Mallorca, Spain,JOURNAL OF QUATERNARY SCIENCE, Issue 8 2008Edward J. Hodge Abstract Very few high-resolution and directly dated terrestrial archives of the last glacial period exist for the western Mediterranean region, yet this is a key locality for recording sub-millennial North Atlantic and Mediterranean climate change. Here, we present evidence of effective precipitation changes based on growth history and ,13C of calcite in a Mallorcan stalagmite that grew between 112 and 48,ka. Effective precipitation in Mallorca appears to have been sensitive to proximal sea surface temperature variations and at certain times, ca. 76,ka for example, changed rapidly from moist to arid conditions in only a few centuries. A sea-level highstand during Marine Isotope Stage (MIS) 5a interrupted growth. Regrowth started promptly after this, but effective precipitation decreased markedly for much of the later part of MIS 5a, and also for shorter periods correlative with Heinrich events H8 (ca. 90,ka) and H6 (ca. 65,ka), with growth ceasing during H5 (ca. 48,ka). Arid episodes in Mallorca appear to be expressions of extremely cold periods recorded further north in Europe and occur contemporaneously with rapid decreases in Greenland temperature. Copyright © 2008 John Wiley & Sons, Ltd. [source] Identification of the Fugloyarbanki tephra in the NGRIP ice core: a key tie-point for marine and ice-core sequences during the last glacial period,JOURNAL OF QUATERNARY SCIENCE, Issue 5 2008S. M. Davies Abstract A visible tephra horizon in the NGRIP ice core has been identified by geochemical analysis as the Fugloyarbanki Tephra, a widespread marker horizon in marine cores from the Faroe Islands area and the northern North Atlantic. An age of 26,740,±,390 yr b2k (1, uncertainty) is derived for this tephra according to the new Greenland Ice Core Chronology (GICC05) based on multi-parameter counting of annual layers. Detection of this tephra for the first time within the NGRIP ice core provides a key tie-point between marine and ice-core records during the transition between MIS 3 and 2. Identification of this volcanic event within the Greenland records demonstrates the future potential of using tephrochronology to precisely correlate palaeoarchives in widely separated localities that span the last glacial period, as well as providing a potential method for examining the extent of the radiocarbon marine reservoir effect at this time. Copyright © 2008 John Wiley & Sons, Ltd. [source] Late Quaternary deglaciation and climate history of the Larsemann Hills (East Antarctica)JOURNAL OF QUATERNARY SCIENCE, Issue 4 2004Elie Verleyen Abstract The Late Quaternary climate history of the Larsemann Hills has been reconstructed using siliceous microfossils (diatoms, chrysophytes and silicoflagellates) in sediment cores extracted from three isolation lakes. Results show that the western peninsula, Stornes, and offshore islands were ice-covered between 30,000,yr,BP and 13,500,cal.,yr,BP. From 13,500,cal.,yr,BP (shortly after the Antarctic Cold Reversal) the coastal lakes of the Larsemann Hills were deglaciated and biogenic sedimentation commenced. Between 13,500 and 11,500,cal.,yr,BP conditions were warmer and wetter than during the preceding glacial period, but still colder than today. From 11,500 to 9500,cal.,yr,BP there is evidence for wet and warm conditions, which probably is related to the early Holocene climate optimum, recorded in Antarctic ice cores. Between 9500 and 7400,cal.,yr,BP dry and cold conditions are inferred from high lake-water salinities, and low water levels and an extended duration of nearshore sea-ice. A second climate optimum occurred between 7400 and 5230,cal.,yr,BP when stratified, open water conditions during spring and summer characterised the marine coast of Prydz Bay. From 5230 until 2750,cal.,yr,BP sea-ice duration in Prydz Bay increased, with conditions similar to the present day. A short return to stratified, open water conditions and a reduction in nearshore winter sea-ice extent is evident between 2750 and 2200,cal.,yr,BP. Simultaneously, reconstructions of lake water depth and salinity suggests relatively humid and warm conditions on land between 3000 and 2000,cal.,yr,BP, which corresponds to a Holocene Hypsithermal reported elsewhere in Antarctica. Finally, dry conditions are recorded around 2000, between 760 and 690, and between 280 and 140,cal.,yr,BP. These data are consistent with ice-core records from Antarctica and support the hypothesis that lacustrine and marine sediments on land can be used to evaluate the effect of long-term climate change on the terrestrial environment. Copyright © 2004 John Wiley & Sons, Ltd. [source] Genetic structure of the European polecat (Mustela putorius) and its implication for conservation strategiesJOURNAL OF ZOOLOGY, Issue 1 2006C. Pertoldi Abstract During the last century, the European polecat Mustela putorius populations in most of Europe declined and survived in fragmented patches, because of habitat alterations and direct persecution. To assess the genetic consequences of the demographic decline and to describe the spatial pattern of genetic diversity, 250 polecats sampled at seven localities from five European countries , Poland, Denmark (southern Denmark and northern Denmark), Spain, Belgium (eastern and western) and the Netherlands , were screened by means of nine microsatellite loci. Genetic diversity estimated by mean expected heterozygosity (HE) and allelic richness (AR) were moderately high within populations [range: 0.50 (northern Denmark) ,HE,0.64 (Poland) and 1.33,AR,7.80] as compared with other carnivores and mustelids. Bottleneck tests suggested that polecat populations in southern Denmark and Poland have declined recently and populations from northern Denmark and the Netherlands have expanded recently, whereas the remaining populations did not show any sign of demographic change. Recent demographic changes could suggest that some of the populations are still not in equilibrium, which could partly explain the relatively high genetic variability observed in polecat populations despite the drastic decline in population size observed in several European countries. A significant heterozygote deficiency [FIS=0.19; 0.01,95% confidence interval (CI),0.32] suggests substructuring within the total European sample. Partitioning of the genetic variation among sampling locations (FST=0.14; 0.06,95% CI,0.23) and pairwise FST between localities (range: 0.01,FST,0.37) without any correlation with the geographic distances between localities were found, suggesting a recent divergence and a restriction of gene flow between populations and the action of genetic drift. An assignment test showed that the Polish and the northern Danish populations were the most unique, whereas the other populations were partially admixed. Factorial component analysis tests indicate a subdivision of the total sample into two distinct groups: one including the samples from Poland and the two Danish localities and the second group comprising the remaining localities investigated. The observed pattern of genetic differentiation is suggested to be due to two main routes of recolonization after the last glacial period. To compare the results obtained with microsatellite data, the most variable region of the mitochondrial DNA (d-loop) was sequenced and different phylogenetic reconstructions and genetic diversity analyses based on nucleotide (,) and haplotype diversity (h) measures within populations were performed using a subsample of populations. The lack of well-defined geographical structure, as well as the reduced level of mitochondrial DNA variability (,: 0.00274±0.00038; h: 0.876±0.028) that was found, has been previously reported in several studies on different carnivores and supports the hypothesis of post-glacial recolonization from southern or eastern refugees of Europe as suggested by the microsatellite data. Implications for conservation strategies of the polecat at the European level are discussed. [source] The distribution of mountain hares Lepus timidus in Europe: a challenge from brown hares L. europaeus?MAMMAL REVIEW, Issue 1 2003CARL-GUSTAF THULIN ABSTRACT 1.,Throughout the most recent glacial period (Weichsel), the mountain hare Lepus timidus had a continuous distribution in the tundra habitat south of the ice-rim. When the ice retreated, mountain hares colonized deglaciated land, and spread over northern Europe. 2.,Since the Weichsel, the mountain hare's distribution in Europe has been gradually reduced and at present comprises Ireland and the Scottish Highlands, high altitudes in the Alps, isolated forests in eastern Poland, most of Fennoscandia and from the Baltic countries eastwards through Russia. Declines during the last century have been observed in Sweden and Russia. 3.,This review defines and evaluates causes for this gradual reduction and fragmentation of the mountain hare's distribution, with special focus on interactions with brown hares Lepus europaeus. The relative importance of diseases, predation, cultivation and interactions with other herbivores than brown hares are discussed. 4.,A plausible cause of the possible permanent disappearance of mountain hares in Europe appears to be exclusion by interspecific competition and hybridization with, and/or epidemic diseases mediated by, the congeneric brown hare. [source] Wolves in the Great Lakes region: a phylogeographic puzzleMOLECULAR ECOLOGY, Issue 20 2010ETTORE RANDI Empirical studies demonstrate that natural hybridization in animals is more common than thought so far (Mallet 2005), particularly among species that originated recently through cycles of population contraction,expansion arising from climate changes over the last glacial period, the Pleistocene. In addition, the post-glacial global growth of human populations has fostered anthropogenic hybridization events, mediated by habitat changes, the persecution of large predators and the introduction of alien species (Allendorf et al. 2001). The Canis lineage shows cases of both natural and anthropogenic hybridization, exacerbating the controversy about the number of species that should be formally validated in the taxonomic lists, the evolutionary role of genetic introgression and the ways to manage hybrids with invading wild or domesticated populations. The study by Wheeldon et al. (2010), published in this issue of Molecular Ecology, adds a new piece to the intricate puzzle of evolution and taxonomy of Canis in North America. They show that sympatric wolves (C. lupus) and coyotes (C. latrans) are not (extensively) hybridizing in the western North American Great Lakes region (GLR). Widespread hybridization between coyotes and a genetically distinct, but closely related, wolf-like population (the eastern wolf) occurred in the northeastern regions of North America. In Wheeldon et al.'s (2010) opinion, these data should prove definitely that two different species of wolf (the western gray wolf C. lupus and the eastern wolf C. lycaon) and their hybrids are distributed across the GLR. [source] Phylogeography of ninespine sticklebacks (Pungitius pungitius) in North America: glacial refugia and the origins of adaptive traitsMOLECULAR ECOLOGY, Issue 18 2010JACLYN T. ALDENHOVEN Abstract The current geographical distribution of the ninespine stickleback (Pungitius pungitius) was shaped in large part by the glaciation events of the Pleistocene epoch (2.6 Mya,10 Kya). Previous efforts to elucidate the phylogeographical history of the ninespine stickleback in North America have focused on a limited set of morphological traits, some of which are likely subject to widespread convergent evolution, thereby potentially obscuring relationships among populations. In this study, we used genetic information from both mitochondrial DNA (mtDNA) sequences and nuclear microsatellite markers to determine the phylogenetic relationships among ninespine stickleback populations. We found that ninespine sticklebacks in North America probably dispersed from at least three glacial refugia,the Mississippi, Bering, and Atlantic refugia,not two as previously thought. However, by applying a molecular clock to our mtDNA data, we found that these three groups diverged long before the most recent glacial period. Our new phylogeny serves as a critical framework for examining the evolution of derived traits in this species, including adaptive phenotypes that evolved multiple times in different lineages. In particular, we inferred that loss of the pelvic (hind fin) skeleton probably evolved independently in populations descended from each of the three putative North American refugia. [source] Postglacial colonization of the Tibetan plateau inferred from the matrilineal genetic structure of the endemic red-necked snow finch, Pyrgilauda ruficollisMOLECULAR ECOLOGY, Issue 6 2005YAN HUA QU Abstract Most phylogeographical studies of postglacial colonization focus on high latitude locations in the Northern Hemisphere. Here, we studied the phylogeographical structure of the red-necked snow finch Pyrgilauda ruficollis, an endemic species of the Tibetan plateau. We analysed 879 base pairs (bp) of the mitochondrial cytochrome b gene and 529 bp of the control region in 41 birds from four regional groups separated by mountain ranges. We detected 34 haplotypes, 31 of which occurred in a single individual and only three of which were shared among sampling sites within regional groups or among regional groups. Haplotype diversity was high (h = 0.94); nucleotide diversity was low (ð = 0.00415) and genetic differentiation was virtually non-existent. Analyses of mismatch distributions and geographically nested clades yielded results consistent with contiguous range expansion, and the expansion times were estimated as 0.07,0.19 million years ago (Ma). Our results suggest that P. ruficollis colonized the Tibetan plateau after the extensive glacial period (0.5,0.175 Ma), expanding from the eastern margin towards the inner plateau. Thus, in contrast to many of the postglacial phylogeographical structures known at high latitudes, this colonization occurred without matrilineal population structuring. This might be due to the short glacial cycles typical of the Tibetan plateau, adaptation of P. ruficollis to cold conditions, or refugia and colonized habitat being semicontinuous and thus promoting population mixing. [source] Range expansions in the flightless longhorn cactus beetles, Moneilema gigas and Moneilema armatum, in response to Pleistocene climate changesMOLECULAR ECOLOGY, Issue 4 2005CHRISTOPHER IRWIN SMITH Abstract Pollen cores and plant and animal fossils suggest that global climate changes at the end of the last glacial period caused range expansions in organisms indigenous to the North American desert regions, but this suggestion has rarely been investigated from a population genetic perspective. In order to investigate the impact of Pleistocene climate changes and glacial/interglacial cycling on the distribution and population structure of animals in North American desert communities, biogeographical patterns in the flightless, warm-desert cactus beetles, Moneilema gigas and Moneilema armatum, were examined using mitochondrial DNA (mtDNA) sequence data from the cytochrome oxidase I (COI) gene. Gene tree relationships between haplotypes were inferred using parsimony, maximum-likelihood, and Bayesian analysis. Nested clade analysis and coalescent modelling using the programs mdiv and fluctuate were used to identify demographically independent populations, and to test the hypothesis that Pleistocene climate changes caused recent range expansions in these species. A sign test was used to evaluate the probability of observing concerted population growth across multiple, independent populations. The phylogeographical and nested clade analyses reveal a history of northward expansion in both of these species, as well as a history of past range fragmentation, followed by expansion from refugia. The coalescent analyses provide highly significant evidence for independent range expansions from multiple refugia, but also identify biogeographical patterns that predate the most recent glacial period. The results indicate that widespread desert environments are more ancient than has been suggested in the past. [source] Involutions resulting from annual freeze,thaw cycles: a laboratory simulation based on observations in northeastern JapanPERMAFROST AND PERIGLACIAL PROCESSES, Issue 4 2007Yoshiko Ogino Abstract A pilot laboratory experiment using a reversed two-layer soil model simulated small-scale involutions formed in a seasonal frost environment during the last glacial period. At the modelled site, the interface between the upper aeolian sandy loam and the lower volcanic pumice constitutes small-scale involutions that display upward-extending tapered projections and downward-extending round hollows. Two scale-reduced laboratory models were subjected to three accelerated annual freeze,thaw cycles with monitoring of frost heave, soil temperature, moisture and pressure. Ice segregation near the layer interface induces upheaving of coarse pumice grains on freezing and earlier settlement of mobilised loam on thawing, resulting in deformation of the interface. A reconstructed 3-D interface displays mounds and depressions with a diameter of 15,20,cm and a height increasing with freeze, thaw alternations. The experimental results imply that the repetition of differential heave and soft-loam settlement promotes decimetre-scale involutions in near-saturated soils subject to deep seasonal frost penetration. Copyright © 2007 John Wiley & Sons, Ltd. [source] Late Cenozoic Geology and Paleo-environment Change in the Eastern Edge of Qinghai-Xizang PlateauACTA GEOLOGICA SINICA (ENGLISH EDITION), Issue 5 2008ZHAO Zhizhong Abstract There are late Cenozoic lacustrine deposits and loess and red clay and moraines in eastern edge of the Qinghai-Xizang Plateau. Various genetic sediments recorded rich information on late Cenozoic paleo-environment changes. Xigeda lacustrine formed during 4.2 Ma B.P.-2.6 Ma B.P. There were 9 periodic warm-cold alternations. Eolian deposition in western Sichuan began at 1.15 Ma B.P. The loess-soil sequences recorded successively 14 paleo-monsoon climate cycles. Laterite in Chengdu plain recorded 5 stages of paleoclimatic stages since 1.13 Ma B.P. There was an old glacial period of 4.3 Ma B.P. in eastern Qinghai-Xizang Plateau. During Quaternary, there are 5 extreme paleoclimatic events corresponding with 5 glaciations. [source] HISTORIC CYCLES OF FRAGMENTATION AND EXPANSION IN PARNASSIUS SMINTHEUS (PAPILIONIDAE) INFERRED USING MITOCHONDRIAL DNAEVOLUTION, Issue 1 2004Eric G. DeChaine Abstract Climate oscillations of the Quaternary drove the repeated expansion and contraction of ecosystems. Alpine organisms were probably isolated in sky island refugia during warm interglacials, such as now, and expanded their range by migrating down-slope during glacial periods. We used population genetic and phylogenetic approaches to infer how paleoclimatic events influenced the distribution of genetic variation in the predominantly alpine butterfly Parnassius smintheus. We sequenced a 789 bp region of cytochrome oxidase I for 385 individuals from 20 locations throughout the Rocky Mountains, ranging from southern Colorado to northern Montana. Analyses revealed at lease two centers of diversity in the northern and southern Rocky Mountains and strong population structure. Nested clade analysis suggested that the species experienced repeated cycles of population expansion and fragmentation. The estimated ages of these events, assuming a molecular clock, corresponded with paleoclimatic data on habitat expansion and contraction over the past 400,000 years. We propose that alpine butterflies persisted in an archipelago of isolated sky islands during interglacials and that populations expanded and became more connected during cold glacial periods. An archipelago model implies that the effects of genetic drift and selection varied among populations, depending on their latitude, area, and local environment. Alpine organisms are sensitive indicators of climate change and their history can be used to predict how high-elevation ecosystems might respond to further climate warming. [source] Quantifying the erosional impact of the Fennoscandian ice sheet in the Torneträsk,Narvik corridor, northern Sweden, based on cosmogenic radionuclide dataGEOGRAFISKA ANNALER SERIES A: PHYSICAL GEOGRAPHY, Issue 3-4 2002Arjen P. Stroeven Despite spectacular landform evidence of a dominant role for glacial action in shaping landscapes under former northern hemisphere ice sheets, there is little quantitative evidence for rates and patterns of erosion associated with specific glaciations. Here we use cosmogenic nuclide data to assess rates of subglacial erosion underneath the Fennoscandian ice sheet. By testing whether there are remnant nuclide concentrations in samples taken from sites that include both relict areas and features and landscapes typically associated with vigorous glacial erosion, we can constrain the level and pattern of modification that resulted from the last glaciation. Cosmogenic 10Be and 36Cl data from the Torneträsk region confirm the temporal and spatial variability of glacial erosion suggested by geomorphological mapping. At some sites, glacial erosion estimates in what appear to be heavily scoured areas indicate erosion of only c. 2 ± 0.4 m of bedrock, based on cosmogenic nuclide inheritance. This implies that the generation of severely scoured terrain in this study area required multiple glaciations. The overall modification produced by ice sheets along glacial corridors may be more restricted than previously thought, or may have occurred preferentially during earlier Quaternary glacial periods. [source] Introductory perspective on the COREF ProjectISLAND ARC, Issue 4 2006Yasufumi Iryu Abstract Coral reefs are tropic to subtropic, coastal ecosystems comprising very diverse organisms. Late Quaternary reef deposits are fossil archives of environmental, tectonic and eustatic variations that can be used to reconstruct the paleoclimatic and paleoceanographic history of the tropic surface oceans. Reefs located at the latitudinal limits of coral-reef ecosystems (i.e. those at coral-reef fronts) are particularly sensitive to environmental changes , especially those associated with glacial,interglacial changes in climate and sealevel. We propose a land and ocean scientific drilling campaign in the Ryukyu Islands (the Ryukyus) in the northwestern Pacific Ocean to investigate the dynamic response of the corals and coral-reef ecosystems in this region to Late Quaternary climate and sealevel change. Such a drilling campaign, which we call the COREF (coral-reef front) Project, will allow the following three major questions to be evaluated: (i) What are the nature, magnitude and driving mechanisms of coral-reef front migration in the Ryukyus? (ii) What is the ecosystem response of coral reefs in the Ryukyus to Quaternary climate changes? (iii) What is the role of coral reefs in the global carbon cycle? Subsidiary objectives include (i) the timing of coral-reef initiation in the Ryukyus and its causes; (ii) the position of the Kuroshio current during glacial periods and its effects on coral-reef formation; and (iii) early carbonate diagenetic responses as a function of compounded variations in climate, eustacy and depositional mineralogies (subtropic aragonitic to warm-temperate calcitic). The geographic, climatic and oceanographic settings of the Ryukyu Islands provide an ideal natural laboratory to address each of these research questions. [source] Historical male-mediated introgression in horseshoe bats revealed by multilocus DNA sequence dataMOLECULAR ECOLOGY, Issue 7 2010XIUGUANG MAO Abstract Instances of hybridization between mammalian taxa in the wild are rarely documented. To test for introgression between sibling species of horseshoe bat (Rhinolophus yunanensis and R. pearsoni) and two subspecies of the latter (R. p. pearsoni and R. p. chinensis), we sequenced two mtDNA and two ncDNA markers in individuals sampled from multiple localities within their overlapping ranges. The interspecific mtDNA gene tree corresponded to the expected taxonomic divisions, and coalescent-based analyses suggested divergence occurred around 4 MYA. However, these relationships strongly conflicted with those recovered from two independent nuclear gene trees, in which R. yunanensis clustered with R. p. pearsoni to the exclusion of R. p. chinensis. This geographically widespread discordance is best explained by large-scale historical introgression of ncDNA from R. yunanensis to R. pearsoni by male-mediated exchange in mixed species colonies during Pleistocene glacial periods, when ranges may have contracted and overlapped more than at present. Further species tree,gene tree conflicts were detected between R. p. pearsoni and R. p. chinensis, also indicating past and/or current introgression in their overlapping regions. However, here the patterns point to asymmetric mtDNA introgression without ncDNA introgression. Analyses of coalescence times indicate this exchange has occurred subsequent to the divergence of these subspecies from their common ancestor. Our work highlights the importance of using multiple data sets for reconstructing phylogeographic histories and resolving taxonomic relationships. [source] Shifting distributions and speciation: species divergence during rapid climate changeMOLECULAR ECOLOGY, Issue 3 2007BRYAN C. CARSTENS Abstract Questions about how shifting distributions contribute to species diversification remain virtually without answer, even though rapid climate change during the Pleistocene clearly impacted genetic variation within many species. One factor that has prevented this question from being adequately addressed is the lack of precision associated with estimates of species divergence made from a single genetic locus and without incorporating processes that are biologically important as populations diverge. Analysis of DNA sequences from multiple variable loci in a coalescent framework that (i) corrects for gene divergence pre-dating speciation, and (ii) derives divergence-time estimates without making a priori assumptions about the processes underlying patterns of incomplete lineage sorting between species (i.e. allows for the possibility of gene flow during speciation), is critical to overcoming the inherent logistical and analytical difficulties of inferring the timing and mode of speciation during the dynamic Pleistocene. Estimates of species divergence that ignore these processes, use single locus data, or do both can dramatically overestimate species divergence. For example, using a coalescent approach with data from six loci, the divergence between two species of montane Melanoplus grasshoppers is estimated at between 200 000 and 300 000 years before present, far more recently than divergence estimates made using single-locus data or without the incorporation of population-level processes. Melanoplus grasshoppers radiated in the sky islands of the Rocky Mountains, and the analysis of divergence between these species suggests that the isolation of populations in multiple glacial refugia was an important factor in promoting speciation. Furthermore, the low estimates of gene flow between the species indicate that reproductive isolation must have evolved rapidly for the incipient species boundaries to be maintained through the subsequent glacial periods and shifts in species distributions. [source] Historical colonization and demography of the Mediterranean damselfish, Chromis chromisMOLECULAR ECOLOGY, Issue 13 2005VERA S. DOMINGUES Abstract The desiccation of the Mediterranean Sea during the Messinian Salinity Crisis 6.0,5.3 million years ago (Ma), caused a major extinction of the marine ichthyofauna of the Mediterranean. This was followed by an abrupt replenishment of the Mediterranean from the Atlantic after the opening of the Strait of Gibraltar. In this study, we combined demographic and phylogeographic approaches using mitochondrial and nuclear DNA markers to test the alternative hypotheses of where (Atlantic or Mediterranean) and when (before or after the Messinian Salinity Crisis) speciation occurred in the Mediterranean damselfish, Chromis chromis. The closely related geminate transisthmian pair Chromis multilineata and Chromis atrilobata was used as a way of obtaining an internally calibrated molecular clock. We estimated C. chromis speciation timing both by determining the time of divergence between C. chromis and its Atlantic sister species Chromis limbata (0.93,3.26 Ma depending on the molecular marker used, e.g. 1.23,1.39 Ma for the control region), and by determining the time of coalescence for C. chromis based on mitochondrial control region sequences (0.14,0.21 Ma). The time of speciation of C. chromis was always posterior to the replenishment of the Mediterranean basin, after the Messinian Salinity Crisis. Within the Mediterranean, C. chromis population structure and demographic characteristics revealed a genetic break at the Peloponnese, Greece, with directional and eastbound gene flow between western and eastern groups. The eastern group was found to be more recent and with a faster growing population (coalescent time = 0.09,0.13 Ma, growth = 485.3) than the western group (coalescent time = 0.13,0.20 Ma, growth = 325.6). Our data thus suggested a western origin of C. chromis, most likely within the Mediterranean. Low sea water levels during the glacial periods, the hydrographic regime of the Mediterranean and dispersal restriction during the short pelagic larval phase of C. chromis (18,19 days) have probably played an important role in C. chromis historical colonization. [source] Phylogeographic analysis of the red seaweed Palmaria palmata reveals a Pleistocene marine glacial refugium in the English ChannelMOLECULAR ECOLOGY, Issue 3 2005JIM PROVAN Abstract Phylogeography has provided a new approach to the analysis of the postglacial history of a wide range of taxa but, to date, little is known about the effect of glacial periods on the marine biota of Europe. We have utilized a combination of nuclear, plastid and mitochondrial genetic markers to study the biogeographic history of the red seaweed Palmaria palmata in the North Atlantic. Analysis of the nuclear rDNA operon (ITS1-5.8S-ITS2), the plastid 16S- trnI- trnA-23S-5S, rbcL- rbcS and rpl12- rps31- rpl9 regions and the mitochondrial cox2,3 spacer has revealed the existence of a previously unidentified marine refugium in the English Channel, along with possible secondary refugia off the southwest coast of Ireland and in northeast North America and/or Iceland. Coalescent and mismatch analyses date the expansion of European populations from approximately 128 000 bp and suggest a continued period of exponential growth since then. Consequently, we postulate that the penultimate (Saale) glacial maximum was the main event in shaping the biogeographic history of European P. palmata populations which persisted throughout the last (Weichselian) glacial maximum (c. 20 000 bp) in the Hurd Deep, an enigmatic trench in the English Channel. [source] History and evolution of the arctic flora: in the footsteps of Eric HulténMOLECULAR ECOLOGY, Issue 2 2003Richard J. Abbott Abstract A major contribution to our initial understanding of the origin, history and biogeography of the present-day arctic flora was made by Eric Hultén in his landmark book Outline of the History of Arctic and Boreal Biota during the Quarternary Period, published in 1937. Here we review recent molecular and fossil evidence that has tested some of Hultén's proposals. There is now excellent fossil, molecular and phytogeographical evidence to support Hultén's proposal that Beringia was a major northern refugium for arctic plants throughout the Quaternary. In contrast, most molecular evidence fails to support his proposal that contemporary east and west Atlantic populations of circumarctic and amphi-Atlantic species have been separated throughout the Quaternary. In fact, populations of these species from opposite sides of the Atlantic are normally genetically very similar, thus the North Atlantic does not appear to have been a strong barrier to their dispersal during the Quaternary. Hultén made no detailed proposals on mechanisms of speciation in the Arctic; however, molecular studies have confirmed that many arctic plants are allopolyploid, and some of them most probably originated during the Holocene. Recurrent formation of polyploids from differentiated diploid or more low-ploid populations provides one explanation for the intriguing taxonomic complexity of the arctic flora, also noted by Hultén. In addition, population fragmentation during glacial periods may have lead to the formation of new sibling species at the diploid level. Despite the progress made since Hultén wrote his book, there remain large gaps in our knowledge of the history of the arctic flora, especially about the origins of the founding stocks of this flora which first appeared in the Arctic at the end of the Pliocene (approximately 3 Ma). Comprehensive analyses of the molecular phylogeography of arctic taxa and their relatives together with detailed fossil studies are required to fill these gaps. [source] | ||||||||||||||||||||||||||||