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Geological Timescales (geological + timescale)
Selected AbstractsTHE HISTORICAL BIOGEOGRAPHY OF TWO CARIBBEAN BUTTERFLIES (LEPIDOPTERA: HELICONIIDAE) AS INFERRED FROM GENETIC VARIATION AT MULTIPLE LOCIEVOLUTION, Issue 3 2002Neil Davies Abstract Mitochondrial DNA and allozyme variation was examined in populations of two Neotropical butterflies, Heliconius charithonia and Dryas iulia. On the mainland, both species showed evidence of considerable gene flow over huge distances. The island populations, however, revealed significant genetic divergence across some, but not all, ocean passages. Despite the phylogenetic relatedness and broadly similar ecologies of these two butterflies, their intraspecific biogeography clearly differed. Phylogenetic analyses of mitochondrial DNA sequences revealed that populations of D. iulia north of St. Vincent are monophyletic and were probably derived from South America. By contrast, the Jamaican subspecies of H. charithonia rendered West Indian H. charithonia polyphyletic with respect to the mainland populations; thus, H. charithonia seems to have colonized the Greater Antilles on at least two separate occasions from Central America. Colonization velocity does not correlate with subsequent levels of gene flow in either species. Even where range expansion seems to have been instantaneous on a geological timescale, significant allele frequency differences at allozyme loci demonstrate that gene flow is severely curtailed across narrow ocean passages. Stochastic extinction, rapid (re)colonization, but low gene flow probably explain why, in the same species, some islands support genetically distinct and nonexpanding populations, while nearby a single lineage is distributed across several islands. Despite the differences, some common biogeographic patterns were evident between these butterflies and other West Indian taxa; such congruence suggests that intraspecific evolution in the West Indies has been somewhat constrained by earth history events, such as changes in sea level. [source] Recognition of southern Gondwanan palynomorphs at Gondwana's northern margin,and biostratigraphic correlation of Permian strata from SE Turkey and AustraliaGEOLOGICAL JOURNAL, Issue 2-3 2010Ellen Stolle Abstract This study focuses on the palynology of Guadalupian (Middle Permian) strata of SE Turkey, especially on late Wordian and earliest Capitanian deposits, which are dated by foraminifers and can be chronostratigraphically related to the geological timescale. Herein, palynological species, such as Altitriletes densus, Cymatiosphaera gondwanensis and Praecolpatites sinuosus, previously characteristic for Pakistan, Australia and Antarctica are recorded. Therefore, the Permian biozones of marine fauna and the palynology of SE Turkey and the rest of the Arabian area and Australia are compared and correlated. This long-distance, eastern Gondwana-wide biostratigraphical correlation, conducted for the first time in the Guadalupian epoch in this study, showed that Corisaccites alutas has a similar Last Occurrence Datum in SE Turkey and in Australia. The correlation also showed that in the late Wordian a number of species were present throughout eastern Gondwana, whereas the distribution of other certain species was influenced by provincialism. Hence, it may be concluded that certain species of parent plants probably co-occurred Gondwana-wide, while the distribution of others was dependant on climate. Copyright © 2010 John Wiley & Sons, Ltd. [source] Formal ratification of the Quaternary System/Period and the Pleistocene Series/Epoch with a base at 2.58 Ma,JOURNAL OF QUATERNARY SCIENCE, Issue 2 2010Philip L. Gibbard Abstract In June 2009, the Executive Committee of the International Union of Geological Sciences (IUGS) formally ratified a proposal by the International Commission on Stratigraphy to lower the base of the Quaternary System/Period to the Global Stratotype Section and Point (GSSP) of the Gelasian Stage/Age at Monte San Nicola, Sicily, Italy. The Gelasian until then had been the uppermost stage of the Pliocene Series/Epoch. The base of the Gelasian corresponds to Marine Isotope Stage 103, and has an astronomically tuned age of 2.58 Ma. A proposal that the base of the Pleistocene Series/Epoch be lowered to coincide with that of the Quaternary (the Gelasian GSSP) was also accepted by the IUGS Executive Committee. The GSSP at Vrica, Calabria, Italy, which had hitherto defined the basal boundary of both the Quaternary and the Pleistocene, remains available as the base of the Calabrian Stage/Age (now the second stage of the revised Pleistocene). In ratifying these proposals, the IUGS has acknowledged the distinctive qualities of the Quaternary by reaffirming it as a full system/period, correctly complied with the hierarchical requirements of the geological timescale by lowering the base of the Pleistocene to that of the Quaternary, and fully respected the historical and widespread current usage of both the terms ,Quaternary' and ,Pleistocene'. Copyright © 2009 John Wiley & Sons, Ltd. [source] Diversity patterns amongst herbivorous dinosaurs and plants during the Cretaceous: implications for hypotheses of dinosaur/angiosperm co-evolutionJOURNAL OF EVOLUTIONARY BIOLOGY, Issue 3 2009R. J. BUTLER Abstract Palaeobiologists frequently attempt to identify examples of co-evolutionary interactions over extended geological timescales. These hypotheses are often intuitively appealing, as co-evolution is so prevalent in extant ecosystems, and are easy to formulate; however, they are much more difficult to test than their modern analogues. Among the more intriguing deep time co-evolutionary scenarios are those that relate changes in Cretaceous dinosaur faunas to the primary radiation of flowering plants. Demonstration of temporal congruence between the diversifications of co-evolving groups is necessary to establish whether co-evolution could have occurred in such cases, but is insufficient to prove whether it actually did take place. Diversity patterns do, however, provide a means for falsifying such hypotheses. We have compiled a new database of Cretaceous dinosaur and plant distributions from information in the primary literature. This is used as the basis for plotting taxonomic diversity and occurrence curves for herbivorous dinosaurs (Sauropodomorpha, Stegosauria, Ankylosauria, Ornithopoda, Ceratopsia, Pachycephalosauria and herbivorous theropods) and major groups of plants (angiosperms, Bennettitales, cycads, cycadophytes, conifers, Filicales and Ginkgoales) that co-occur in dinosaur-bearing formations. Pairwise statistical comparisons were made between various floral and faunal groups to test for any significant similarities in the shapes of their diversity curves through time. We show that, with one possible exception, diversity patterns for major groups of herbivorous dinosaurs are not positively correlated with angiosperm diversity. In other words, at the level of major clades, there is no support for any diffuse co-evolutionary relationship between herbivorous dinosaurs and flowering plants. The diversification of Late Cretaceous pachycephalosaurs (excluding the problematic taxon Stenopelix) shows a positive correlation, but this might be spuriously related to poor sampling in the Turonian,Santonian interval. Stegosauria shows a significant negative correlation with flowering plants and a significant positive correlation with the nonflowering cycadophytes (cycads, Bennettitales). This interesting pattern is worthy of further investigation, and it reflects the decline of both stegosaurs and cycadophytes during the Early Cretaceous. [source] 55 Ice age kelp forests: climate-driven changes in kelp forest distribution since the last glacial maximumJOURNAL OF PHYCOLOGY, Issue 2003M. H. Graham Kelp forest distributions are constrained by the availability of rocky substrate within the depth range tolerable for growth and reproduction, which can vary over relatively short geological timescales (millennia) due to interactions between coastal bathymetry and climate-driven changes in eustatic sea level. Using GIS, a digital bathymetric map, sea level curves, and published kelp depth tolerances, I reconstructed changes in the size and distribution of giant kelp (Macrocystis pyrifera) forests in the Southern California Bight since the last glacial maximum. Reconstructions predicted that the total area of available kelp forest habitat for the California Channel Islands during the last glacial maximum (18.5 kyr BP; 628 square km) was greater than at present (382 square km) but less than at 16.5 kyr BP (1130 square km). Available kelp forest habitat along the southern California mainland also increased rapidly from 18.5 to 16.5 kyr BP but continued to increase with sea level rise. Differences in the effects of sea level rise on coastal geomorphology between the islands and mainland further constrained the extent of rocky substrate available to kelps. Given biomass and productivity estimates from present-day kelp forests, these reconstructions suggest more productive and spatially extensive island kelp forests near the last glacial maximum than at present, but the opposite pattern for the mainland. These climate-driven changes in kelp forest distribution and productivity likely had important historical impacts on the ecology and evolution of the present-day kelp ecosystem including kelp forest exploitation by early human inhabitants of southern California. [source] Benzene polycarboxylic acids,A ubiquitous class of compounds in soilsJOURNAL OF PLANT NUTRITION AND SOIL SCIENCE, Issue 5 2010Ludwig Haumaier Abstract Black carbon (BC) occurs ubiquitously in the environment. Its oxidation in the laboratory yields a suite of benzene polycarboxylic acids (BPCAs), suggesting similar oxidation products in soils. Since only for a few soils the occurrence of BPCAs in the free form has been documented, screening for them in a broad range of contrasting soils was conducted. They were extracted from soil samples with 0.5 M NaOH and quantified using gas chromatography,mass spectrometry. As expected, BPCAs turned out to be as ubiquitous as BC. They were detected not only in every soil sample investigated so far, but also in samples from drill cores up to a depth of 10 m and in recently deposited calcareous tufa. The concentrations covered a range similar to that of some phenolic acids. The range exceeded those reported for low-molecular-weight aliphatic acids or simple sugars in soils. The distribution of BPCAs in soil profiles indicated a considerable potential of translocation within, and export from, soil, in particular of benzene hexacarboxylic (mellitic) acid. Mellitic acid may therefore be present in almost any geochemical sample affected by seepage water from soils. Its high water solubility and strong metal-complexing ability suggest it may be involved in metal-transport processes, at least on geological timescales. [source] A Bayesian approach to inverse modelling of stratigraphy, part 1: methodBASIN RESEARCH, Issue 1 2009Karl Charvin ABSTRACT The inference of ancient environmental conditions from their preserved response in the sedimentary record still remains an outstanding issue in stratigraphy. Since the 1970s, conceptual stratigraphic models (e.g. sequence stratigraphy) based on the underlying assumption that accommodation space is the critical control on stratigraphic architecture have been widely used. Although these methods considered more recently other possible parameters such as sediment supply and transport efficiency, they still lack in taking into account the full range of possible parameters, processes, and their complex interactions that control stratigraphic architecture. In this contribution, we present a new quantitative method for the inference of key environmental parameters (specifically sediment supply and relative sea level) that control stratigraphy. The approach combines a fully non-linear inversion scheme with a ,process,response' forward model of stratigraphy. We formulate the inverse problem using a Bayesian framework in order to sample the full range of possible solutions and explicitly build in prior geological knowledge. Our methodology combines Reversible Jump Markov chain Monte Carlo and Simulated Tempering algorithms which are able to deal with variable-dimensional inverse problems and multi-modal posterior probability distributions, respectively. The inverse scheme has been linked to a forward stratigraphic model, BARSIM (developed by Joep Storms, University of Delft), which simulates shallow-marine wave/storm-dominated systems over geological timescales. This link requires the construction of a likelihood function to quantify the agreement between simulated and observed data of different types (e.g. sediment age and thickness, grain size distributions). The technique has been tested and validated with synthetic data, in which all the parameters are specified to produce a ,perfect' simulation, although we add noise to these synthetic data for subsequent testing of the inverse modelling approach. These tests addressed convergence and computational-overhead issues, and highlight the robustness of the inverse scheme, which is able to assess the full range of uncertainties on the inferred environmental parameters and facies distributions. [source] Apatite fission track thermochronology: an overview of its potential and limitations in geomorphologyBASIN RESEARCH, Issue 2 2000Gunnell The specific time window of apatite fission track thermochronology (AFTT) places it in a unique position to offer continuous time,temperature baseline histories for relatively stable shield, rift and passive margin environments, spanning several geological eras. Fission tracks anneal partially at temperatures between ,110 °C and ,60 °C and thereby provide information on residence times within specific levels of the crust. Most samples collected from these terrains, however, are usually found to have cooled out of the apatite partial annealing zone (APAZ) by the mid-Cenozoic at the latest. Owing to the stability, at geological timescales, of tracks at temperatures <60 °C (i.e. within depths of 0,2.5 km at normal geothermal gradients), a significant loss of resolution must therefore be reckoned with at shallow, although geomorphologically crucial, crustal depths. Indeed, the Neogene and Quaternary are understood to have been most influential in generating the scenery of today, and a use of radiometric and stratigraphic techniques in a nested, multisystem approach can assist in bridging the resolution gap. This paper examines and illustrates, mostly with original examples, the uses and limitations of AFTT in addressing the response of Earth surface systems to event patterns in global tectonics, the controls of lithology and structure on denudation rates, the origin and evolution of passive margin escarpments, the mass-balanced reconstruction of palaeoelief, the use of apatites as tracers for understanding provenance in sediment routing systems, and the tempo (or episodicity) of denudation as postulated by W. M. Davis' canons of the ,geographical cycle'. Alongside efforts towards standardizing the supply of analytically robust AFT results in the laboratory, a more standardized geoscientific interpretation of AFT data is also desirable in order to build a consistent world base of geomorphic rates which can be made available to , and used with confidence by , non-AFTT specialists. [source] Testing coevolutionary hypotheses over geological timescales: interactions between Cretaceous dinosaurs and plantsBIOLOGICAL JOURNAL OF THE LINNEAN SOCIETY, Issue 1 2010RICHARD J. BUTLER Testing coevolutionary scenarios over extended geological timescales is fraught with difficulties. Most tests rely on comparisons of temporal variations in taxonomic diversity for the groups of interest: however, this approach typically excludes spatiotemporal data. Here, we apply a quantitative method that incorporates the spatiotemporal distributions of the proposed coevolving groups using a Geographical Information System. Distributional data for Cretaceous dinosaur and plant groups were mapped onto palaeogeographical reconstructions in a series of time-slices. Within each time-slice, palaeocontinental surfaces were divided into a series of grids, each of which was scored as present, absent or inapplicable (unsampled) for each group. Distributions were compared statistically to determine whether the putative coevolving groups co-occurred within grid squares more or less frequently than expected by chance. Pairwise comparisons were made between herbivorous dinosaur clades and major plant groups (e.g. cycads, angiosperms) on a global scale. Only three nonrepeated associations of marginal significance were recovered, demonstrating that, in general, current knowledge of the spatiotemporal distributions of these groups provides little support for coevolutionary hypotheses. The Geographical Information System methods used are readily applicable to many other questions whose answers are reliant on a detailed knowledge of organismal distributions in time and space. © 2010 The Linnean Society of London, Biological Journal of the Linnean Society, 2010, 100, 1,15. [source] Testing co-evolutionary hypotheses over geological timescales: interactions between Mesozoic non-avian dinosaurs and cycadsBIOLOGICAL REVIEWS, Issue 1 2009Richard J. Butler Abstract The significance of co-evolution over ecological timescales is well established, yet it remains unclear to what extent co-evolutionary processes contribute to driving large-scale evolutionary and ecological changes over geological timescales. Some of the most intriguing and pervasive long-term co-evolutionary hypotheses relate to proposed interactions between herbivorous non-avian dinosaurs and Mesozoic plants, including cycads. Dinosaurs have been proposed as key dispersers of cycad seeds during the Mesozoic, and temporal variation in cycad diversity and abundance has been linked to dinosaur faunal changes. Here we assess the evidence for proposed hypotheses of trophic and evolutionary interactions between these two groups using diversity analyses, a new database of Cretaceous dinosaur and plant co-occurrence data, and a geographical information system (GIS) as a visualisation tool. Phylogenetic evidence suggests that the origins of several key biological properties of cycads (e.g. toxins, bright-coloured seeds) likely predated the origin of dinosaurs. Direct evidence of dinosaur,cycad interactions is lacking, but evidence from extant ecosystems suggests that dinosaurs may plausibly have acted as seed dispersers for cycads, although it is likely that other vertebrate groups (e.g. birds, early mammals) also played a role. Although the Late Triassic radiations of dinosaurs and cycads appear to have been approximately contemporaneous, few significant changes in dinosaur faunas coincide with the late Early Cretaceous cycad decline. No significant spatiotemporal associations between particular dinosaur groups and cycads can be identified , GIS visualisation reveals disparities between the spatiotemporal distributions of some dinosaur groups (e.g. sauropodomorphs) and cycads that are inconsistent with co-evolutionary hypotheses. The available data provide no unequivocal support for any of the proposed co-evolutionary interactions between cycads and herbivorous dinosaurs , diffuse co-evolutionary scenarios that are proposed to operate over geological timescales are plausible, but such hypotheses need to be firmly grounded on direct evidence of interaction and may be difficult to support given the patchiness of the fossil record. [source] | |||||||||||||