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Alpine Tundra (alpine + tundra)
Selected AbstractsThe Deciduous Forest , Boreal Forest EcotoneGEOGRAPHY COMPASS (ELECTRONIC), Issue 7 2010David Goldblum Ecotones have been subject to significant attention over the past 25 years as a consensus emerged that they might be uniquely sensitive to the effects of climate change. Most ecotone field studies and modeling efforts have focused on transitions between forest and non-forest biomes (e.g. boreal forest to Arctic tundra, forest to prairie, subalpine forests to alpine tundra) while little effort has been made to evaluate or simply understand forest,forest ecotones, specifically the deciduous forest , boreal forest ecotone. Geographical shifts and changes at this ecotone because of anthropogenic factors are tied to the broader survival of both the boreal and deciduous forest communities as well as global factors such as biodiversity loss and dynamics of the carbon cycle. This review summarizes what is known about the location, controlling mechanisms, disturbance regimes, anthropogenic impacts, and sensitivity to climate change of the deciduous forest , boreal forest ecotone. [source] Eastern Beringian biogeography: historical and spatial genetic structure of singing voles in AlaskaJOURNAL OF BIOGEOGRAPHY, Issue 8 2010Marcelo Weksler Abstract Aim Pleistocene climatic cycles have left marked signatures in the spatial and historical genetic structure of high-latitude organisms. We examine the mitochondrial (cytochrome b) genetic structure of the singing vole, Microtus miurus (Rodentia: Cricetidae: Arvicolinae), a member of the Pleistocene Beringian fauna, and of the insular vole, Microtus abbreviatus, its putative sister species found only on the St Matthew Archipelago. We reconstruct the phylogenetic and phylogeographical structure of these taxa, characterize their geographical partitioning and date coalescent and cladogenetic events in these species. Finally, we compare the recovered results with the phylogenetic, coalescent and spatial genetic patterns of other eastern Beringian mammals and high-latitude arvicoline rodents. Location Continental Alaska (alpine and arctic tundra) and the St Matthew Archipelago (Bering Sea). Methods We generated and analysed cytochrome b sequences of 97 singing and insular voles (M. miurus and M. abbreviatus) from Alaska. Deep evolutionary structure was inferred by phylogenetic analysis using parsimony, maximum likelihood and Bayesian approaches; the geographical structure of genetic diversity was assessed using analysis of molecular variance and network analysis; ages of cladogenetic and coalescent events were estimated using a relaxed molecular clock model with Bayesian approximation. Results Regional nucleotide diversity in singing voles is higher than in other high-latitude arvicoline species, but intra-population diversity is within the observed range of values for arvicolines. Microtus abbreviatus specimens are phylogenetically nested within M. miurus. Molecular divergence date estimates indicate that current genetic diversity was formed in the last glacial (Wisconsinan) and previous interglacial (Sangamonian) periods, with the exception of a Middle Pleistocene split found between samples collected in the Wrangell Mountains region and all other singing vole samples. Main conclusions High levels of phylogenetic and spatial structure are observed among analysed populations. This pattern is consistent with that expected for a taxon with a long history in Beringia. The spatial genetic structure of continental singing voles differs in its northern and southern ranges, possibly reflecting differences in habitat distribution between arctic and alpine tundra. Our phylogenetic results support the taxonomic inclusion of M. miurus in its senior synonym, M. abbreviatus. [source] Surviving climate changes: high genetic diversity and transoceanic gene flow in two arctic,alpine lichens, Flavocetraria cucullata and F. nivalis (Parmeliaceae, Ascomycota)JOURNAL OF BIOGEOGRAPHY, Issue 8 2010József Geml Abstract Aim, We examined genetic structure and long-distance gene flow in two lichenized ascomycetes, Flavocetraria cucullata and Flavocetraria nivalis, which are widespread in arctic and alpine tundra. Location, Circumpolar North. Methods, DNA sequences were obtained for 90 specimens (49 for F. cucullata and 41 for F. nivalis) collected from various locations in Europe, Asia and North America. Sequences of the nuclear internal transcribed spacer (ITS) + 5.8S ribosomal subunit gene region were generated for 89 samples, and supplemented by beta-tubulin (BTUB) and translation elongation factor 1-alpha gene (EF1) sequences for a subset of F. cucullata specimens. Phylogenetic, nonparametric permutation methods and coalescent analyses were used to assess population divergence and to estimate the extent and direction of migration among continents. Results, Both F. cucullata and F. nivalis were monophyletic, supporting their morphology-based delimitation, and had high and moderately high intraspecific genetic diversity, respectively. Clades within each species contained specimens from both North America and Eurasia. We found only weak genetic differentiation among North American and Eurasian populations, and evidence for moderate to high transoceanic gene flow. Main conclusions, Our results suggest that both F. cucullata and F. nivalis have been able to migrate over large distances in response to climatic fluctuations. The high genetic diversity observed in the Arctic indicates long-term survival at high latitudes, whereas the estimated migration rates and weak geographic population structure suggest a continuing long-distance gene flow between continents that has prevented pronounced genetic differentiation. The mode of long-distance dispersal is unknown, but wind dispersal of conidia and/or ascospores is probably important in the open arctic landscapes. The high genetic diversity and efficient long-distance dispersal capability of F. cucullata and F. nivalis suggest that these species, and perhaps other arctic lichens as well, will be able to track their potential niche in the changing Arctic. [source] Post-fire tree establishment patterns at the alpine treeline ecotone: Mount Rainier National Park, Washington, USAJOURNAL OF VEGETATION SCIENCE, Issue 1 2009Kirk M. Stueve Abstract Questions: Does tree establishment: (1) occur at a treeline depressed by fire, (2) cause the forest line to ascend upslope, and/or (3) alter landscape heterogeneity? (4) What abiotic and biotic local site conditions are most important in structuring establishment patterns? (5) Does the abiotic setting become more important with increasing upslope distance from the forest line? Location: Western slopes of Mount Rainier, USA. Methods: We performed classification analysis of 1970 satellite imagery and 2003 aerial photography to delineate establishment. Local site conditions were calculated from a LIDAR-based DEM, ancillary climate data, and 1970 tree locations in a GIS. We used logistic regression on a spatially weighted landscape matrix to rank variables. Results: Considerable establishment after 1970 caused forest line elevation to increase over 150 m in specific locations. Landscape heterogeneity increased with distance from the 1970 forest line. At a broad spatial context, we found establishment was most common near existing trees (0-50 m) and at low elevations (1250-1350 m). Slope aspect (W, NW, N, NE, and E), slope angle (40-60°), and other abiotic factors emerged as important predictors of establishment with increasing upslope distance from the forest line to restricted spatial extents. Conclusions: Favorable climatic conditions likely triggered widespread tree establishment. Readily available seed probably enhanced establishment rates near sexually mature trees, particularly in the less stressful environment at low elevations. The mass effect of nearly ubiquitous establishment in these areas may have obscured the importance of the abiotic setting to restricted spatial extents. Topographic variability apparently produced favorable sites that facilitated opportunistic establishment with increasing upslope distance from the forest line, thereby enabling additional trees to invade the alpine tundra. [source] Establishing a missing link: warm summers and winter snow cover promote shrub expansion into alpine tundra in ScandinaviaNEW PHYTOLOGIST, Issue 4 2010Martin Hallinger Summary ,Shrub expansion in alpine and arctic areas is a process with possibly profound implications for ecosystem functioning. The recent shrub expansion has been mainly documented by remote sensing techniques, but the drivers for this process largely remain hypotheses. ,Here, we outline a dendrochronological method, adapted to shrubs, to address these hypotheses and then present a mechanism for the current shrub expansion by linking recent climate change to shrub growth performance in northern Sweden. ,A pronounced increase in radial and vertical growth during recent decades along an elevational gradient from treeline to shrubline indicates an ongoing shrub expansion. Age distribution of the shrub population indicates the new colonization of shrubs at high elevations. ,Shrub growth is correlated with warm summers and winter snow cover and suggests the potential for large-scale ecosystem changes if climate change continues as projected. [source] Alpine flora dynamics , a critical review of responses to climate change in the Swedish Scandes since the early 1950sNORDIC JOURNAL OF BOTANY, Issue 4 2010Leif Kullman Reports about changes of alpine plant species richness over the past 60 years in the Swedish Scandes are reviewed, synthesized and updated with data from recent reinventories. Methodologically, this endeavour is based on resurveys of the floristic composition on the uppermost 20 m of four high-mountain summits. The key finding is that the species pool has increased by 60,170% since the 1950s and later. Some of the invading species are new to the alpine tundra, with more silvine and thermophilic properties than the extant alpine flora. Not a single species of the original flora has disappeared from any of the summits. This circumstance is discussed in perspective of widespread expectations of pending temperature-driven extinction of alpine species in an alleged future warmer climate. These progressive changes coincided with distinct warming (summer and winter) since the late 1980s. During a short cooler period (1974,1994), the species numbers decreased and the upper elevational limits of some ground cover species descended. Thus, discernible responses, concurrent with both warming and cooling intervals, sustain a strong causal link between climate variability and alpine plant species richness. Methodologically, plot-less revisitation studies of the present kind are beset with substantial uncertainties, which may overstate floristic changes over time. However, it is argued here that carefully executed and critically interpreted, no other method can equally effectively sense the earliest phases of plant invasions into alpine vegetation. [source] Fertilization effects on species density and primary productivity in herbaceous plant communitiesOIKOS, Issue 3 2000Laura Gough Fertilization experiments in plant communities are often interpreted in the context of a hump-shaped relationship between species richness and productivity. We analyze results of fertilization experiments from seven terrestrial plant communities representing a productivity gradient (arctic and alpine tundra, two old-field habitats, desert, short- and tall-grass prairie) to determine if the response of species richness to experimentally increased productivity is consistent with the hump-shaped curve. In this analysis, we compared ratios of the mean response in nitrogen-fertilized plots to the mean in control plots for aboveground net primary productivity (ANPP) and species density (D; number of species per plot of fixed unit area). In general, ANPP increased and plant species density decreased following nitrogen addition, although considerable variation characterized the magnitude of response. We also analyzed a subset of the data limited to the longest running studies at each site (,4 yr), and found that adding 9 to 13 g N m,2 yr,1 (the consistent amount used at all sites) increased ANPP in all communities by approximately 50% over control levels and reduced species density by approximately 30%. The magnitude of response of ANPP and species density to fertilization was independent of initial community productivity. There was as much variation in the magnitude of response among communities within sites as among sites, suggesting community-specific mechanisms of response. Based on these results, we argue that even long-term fertilization experiments are not good predictors of the relationship between species richness and productivity because they are relatively small-scale perturbations whereas the pattern of species richness over natural productivity gradients is influenced by long-term ecological and evolutionary processes. [source] Population genetic structure of rock ptarmigan in the ,sky islands' of French Pyrenees: implications for conservationANIMAL CONSERVATION, Issue 2 2009N. Bech Abstract Expected consequences of global warming include habitat reduction in many cool climate species. Rock ptarmigan is a Holarctic grouse that inhabits arctic and alpine tundra. In Europe, the Pyrenean ptarmigan inhabits the southern edge of the species' range and since the last glacial maximum its habitat has been severely fragmented and is restricted to high-alpine zones or ,sky islands'. A recent study of rock ptarmigan population genetic in Europe found that the Pyrenean ptarmigan had very low genetic diversity compared with that found in the Alps and Scandinavia. Habitat fragmentation and reduced genetic diversity raises concerns about the viability of ptarmigan populations in the Pyrenees. However, information on population structuring and gene flow across the Pyrenees, which is essential for designing a sound management plan, is absent. In this study, we use seven microsatellites and mitochondrial control region sequences to investigate genetic variation and differentiation among five localities across the Pyrenees. Our analyses reveal the presence of genetic differentiation among all five localities and a significant isolation-by-distance effect that is likely the result of short dispersal distances and high natal and breeding philopatry of Pyrenean ptarmigan coupled with severe habitat fragmentation. Furthermore, analysis of molecular variance, principal component analysis and Bayesian analysis of genetic structuring identified the greatest amount of differentiation between the eastern and main parts of the Pyrenean chain separated by the Sčgre Valley. Our data also show that the Canigou massif may host an isolated population and requires special conservation attention. We propose a management plan which includes the translocation of birds. If a sky island structure affects genetic divergence in rock ptarmigan, it may also affect the genetic structure of other sky island species having low dispersal abilities. [source] | |||||||||||||