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Fire Environment (fire + environment)
Selected AbstractsXylem root and shoot hydraulics is linked to life history type in chaparral seedlingsFUNCTIONAL ECOLOGY, Issue 1 2010Robert B. Pratt Summary 1.,Shrubs in fire prone chaparral communities have evolved different life history types in response to fire. A key to understanding the evolution of life history type differences is to understand how physiological traits are linked to differences in life history type. Vascular adaptations are important for delivering an efficient and stable water supply to evergreen chaparral shrub leaves. This study tested for a link between vascular physiology and life history type in chaparral shrubs. 2.,Chaparral shrub species along the south-western coast of North America survive wildfire by three different life histories. Non-sprouters are killed by fire and re-establish exclusively through germination of fire-stimulated seeds, facultative sprouters re-establish by a combination of vegetative sprouting and fire-stimulated seeds, and obligate sprouters re-establish exclusively by vegetative sprouting because their seeds do not survive fire. Non-sprouters and facultative sprouters establish seedlings in the open canopy post fire environment, whereas obligate sprouters establish seedlings in the shady understory of the mature chaparral canopy. 3.,Seedlings of nine species (Rhamnaceae) representing three each of the different life history types were grown in deep containers in a common garden under treatments of sun and shade. Hydraulic conductance was measured using a high-pressure flow meter for all organs, and a vacuum technique was used to measure conductance of fine and woody roots. We predicted that non-sprouters would exhibit greater hydraulic efficiency than the sprouting species, and that facultative sprouters would be more efficient than the shade tolerant obligate sprouters. 4.,Non-sprouters had the greatest hydraulic conductance per unit leaf and sapwood area at the whole seedling level, whereas facultative and obligate sprouters were not different. Comparing hydraulic conductance across major organs (from fine roots to leaves) showed that the hydraulic system was well coordinated. At the whole seedling level, the root system was more of a bottleneck than the shoot system. This pattern was consistent with high resistance extraxylary pathways in roots and differences in root architecture. 5.,The greater hydraulic efficiency of the non-sprouter life history type is attributed to its post-fire pioneering habit and may partially explain the relatively high speciation in the non-sprouters. Lower hydraulic efficiency is associated with a sprouting life history and greater shade tolerance. The seedling root systems represent a hydraulic bottleneck that may place roots under especially intense selection. [source] Life table response experiment analysis of the stochastic growth rateJOURNAL OF ECOLOGY, Issue 2 2010Hal Caswell Summary 1.,Life table response experiment (LTRE) analyses decompose treatment effects on a dependent variable (usually, but not necessarily, population growth rate) into contributions from differences in the parameters that determine that variable. 2.,Fixed, random and regression LTRE designs have been applied to plant populations in many contexts. These designs all make use of the derivative of the dependent variable with respect to the parameters, and describe differences as sums of linear approximations. 3.,Here, I extend LTRE methods to analyse treatment effects on the stochastic growth rate log ,s. The problem is challenging because a stochastic model contains two layers of dynamics: the stochastic dynamics of the environment and the response of the vital rates to the state of the environment. I consider the widely used case where the environment is described by a Markov chain. 4.,As the parameters describing the environmental Markov chain do not appear explicitly in the calculation of log ,s, derivatives cannot be calculated. The solution presented here combines derivatives for the vital rates with an alternative (and older) approach, due to Kitagawa and Keyfitz, that calculates contributions in a way analogous to the calculation of main effects in statistical models. 5.,The resulting LTRE analysis decomposes log ,s into contributions from differences in: (i) the stationary distribution of environmental states, (ii) the autocorrelation pattern of the environment, and (iii) the stage-specific vital rate responses within each environmental state. 6.,As an example, the methods are applied to a stage-classified model of the prairie plant Lomatium bradshawii in a stochastic fire environment. 7.,Synthesis. The stochastic growth rate is an important parameter describing the effects of environmental fluctuations on population viability. Like any growth rate, it responds to differences in environmental factors. Without a decomposition analysis there is no way to attribute differences in the stochastic growth rate to particular parts of the life cycle or particular aspects of the stochastic environment. The methods presented here provide such an analysis, extending the LTRE analyses already available for deterministic environments. [source] The influence of an upper-level frontal zone on the Mack Lake Wildfire environmentMETEOROLOGICAL APPLICATIONS, Issue 2 2007Tarisa Zimet Abstract Meteorological assessment of wildland fire danger has traditionally involved the identification of several synoptic weather types empirically determined to influence wildfire spread. Specifically, in the Great Lakes Region, high wildfire danger is often witnessed in association with northwesterly synoptic-scale flow aloft. Such synoptic-scale flow is regularly associated with the development of upper-level frontal zones also known as upper-level jet/front systems, which are often characterised by intrusions of stratospheric air into the troposphere. The notion that upper-frontal development can play an important role in promoting wildfire spread is advanced through interrogation of the output from a fine-scale numerical simulation of a documented explosive wildfire case; the Mack Lake Fire of May 1980. The Mack Lake case was characterised by a developing upper-level front embedded within a shortwave trough in the vicinity of the fire location. The upper-level front originated in northwesterly flow in central Canada as an upper-tropospheric ridge amplified over western North America. A thermally indirect circulation at the jet exit region both contributed to the intensification of the front and was associated with a maximum in quasi-geostrophic descent at mid-levels upstream of the fire region. The subsidence ushered dry air from the middle and upper-troposphere downward along sloping isentropes adiabatically warming and drying it along the way. A well-developed dry air intrusion associated with the operation of these processes extended to nearly the 750 hPa level far downstream from the actual upper-frontal zone supplying the fire environment with dry air that originated in the upper-troposphere/lower stratosphere. The organised subsidence was also responsible for downward advection of high momentum air from within the frontal zone into the fire environment, further influencing the wildfire spread. We conclude that upper-frontal processes, characteristic of northwesterly synoptic-scale flow, are likely a contributing factor to the prevalence of wildfire spread under such synoptic-scale conditions. Copyright © 2007 Royal Meteorological Society [source] Stage, age and individual stochasticity in demographyOIKOS, Issue 12 2009Hal Caswell Demography is the study of the population consequences of the fates of individuals. Individuals are differentiated on the basis of age or, in general, life cycle stages. The movement of an individual through its life cycle is a random process, and although the eventual destination (death) is certain, the pathways taken to that destination are stochastic and will differ even between identical individuals; this is individual stochasticity. A stage-classified demographic model contains implicit age-specific information, which can be analyzed using Markov chain methods. The living stages in the life cycles are transient states in an absorbing Markov chain; death is an absorbing state. This paper presents Markov chain methods for computing the mean and variance of the lifetime number of visits to any transient state, the mean and variance of longevity, the net reproductive rate R0, and the cohort generation time. It presents the matrix calculus methods needed to calculate the sensitivity and elasticity of all these indices to any life history parameters. These sensitivities have many uses, including calculation of selection gradients. It is shown that the use of R0 as a measure of fitness or an invasion exponent gives erroneous results except when R0=,=1. The Markov chain approach is then generalized to variable environments (deterministic environmental sequences, periodic environments, iid random environments, Markovian environments). Variable environments are analyzed using the vec-permutation method to create a model that classifies individuals jointly by the stage and environmental condition. Throughout, examples are presented using the North Atlantic right whale (Eubaleana glacialis) and an endangered prairie plant (Lomatium bradshawii) in a stochastic fire environment. [source] Characterization of the fire environments in central offices of the telecommunications industryFIRE AND MATERIALS, Issue 3 2003Archibald Tewarson Abstract Eight free burning and two sprinklered fire tests were performed with electrical cable trays and live digital switch racks in a large enclosure to simulate telecommunications central office (TCO) fires started by electrical overheating. Very-slow-growing (non-flaming), slower-growing (partially flaming) and low-intensity-faster-growing (flaming) fires releasing gray-white, gray, and black smoke, respectively, were observed in the tests. Under quiescent conditions present in the unvented enclosure fire tests for cables, very-slow-growing fires were detected in about 1452 s, whereas the slower-growing fires were detected in about 222 s by commercial fire detectors. Under ventilation conditions typical of TCOs, detection times were very similar for the five types of commercial TCOs fire detectors used in the tests. The average detection times for slower-growing fires (cable fires) and low-intensity-faster-growing fires (digital switch rack fires) were 242±17% and 249±11%s respectively. The TCO procedures to reduce smoke damage from fires (on fire detection, inlet ventilation flow is turned off and exhaust flow is turned on) were found to be beneficial. The extent of smoke damage decreased significantly with an increase in the exhaust flow rate. The chloride ion mass deposition suggested that equipment recovery would be possible in the smoke environment if the cable vapor concentration could be reduced below about 3 g/m3. The metal corrosion rate was found proportional to the 0.6th power of the smoke concentration, similar to that found for the corrosion of metal surfaces exposed to aqueous solutions of HCl and HNO3 and for acid rain with no protective layer at the surface. Sprinkler water was found to wash down the smoke deposits on the surfaces with little indication of corrosion enhancement. Copyright © 2003 John Wiley & Sons, Ltd. [source] Labour Market Regimes and Worker Recruitment and Retention in the European Union: Plant ComparisonsBRITISH JOURNAL OF INDUSTRIAL RELATIONS, Issue 4 2001Jane Morton We analyse worker recruitment and retention outcomes, contrasting production worker skill profiles in the UK subsidiaries of five multinationals with, in each case, a matched plant in another EU state. From personnel records we build up plant distributions of worker pay, age, schooling, prior experience and tenure. We find evidence of local management discretion, with differences in unit labour costs and in recruitment/retention outcomes between the matched subsidiaries. UK plants have a ,hire and fire' environment and a less skilled worker mix. Such outcomes can be linked to the UK's lower labour costs and less regulated labour market. [source] |