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Drier Conditions (drier + condition)
Selected AbstractsUnderground Vetch (Vicia sativa ssp. amphicarpa): A Potential Pasture and Forage Legume for Dry Areas in West AsiaJOURNAL OF AGRONOMY AND CROP SCIENCE, Issue 3 2003A. M. Abd El Moneim Abstract Subterranean vetch [Vicia sativa ssp. amphicarpa (Dorth.) Aschers & Graebn.] is native to disturbed grasslands of the Mediterranean basin where heavy grazing, seasonal drought and erosion act as strong selection forces. It produces two pod types, above-ground and 5 cm below the soil surface. Unlike subterranean clover (Trifolium subterranean L.), which buries its seeds after flowering above-ground, subterranean vetch flowers and forms pods beneath the soil surface on underground stems. The aerial pods are produced after vegetative development ceases, while the underground pods are produced in ontogeny. The ability of this unusual vetch to survive in marginal areas with low rainfall (about 250 mm year,1) and to produce nutritious herbage and pods is an important characteristic which helps address rehabilitation of degraded rangelands and increase feed production for small ruminants. Research at the International Center for Agricultural Research in the Dry Areas (ICARDA) during the 1988,93 growing seasons has assessed the herbage and seed productivity of underground vetch, its ability to grow in rotation with barley in marginal low-rainfall areas, and its capacity to regenerate after heavy grazing. Drier conditions in 1989 favoured earlier underground flowering; the number of underground pods was higher than that of aerial pods. Grain yield of barley (var. Atlas) was around 2.0 t ha,1 after underground vetch and only 1.2 t ha,1 after barley. Grazing underground vetch had no effect on the productivity of the succeeding barley crop. The aerial and underground pods serve two distinct functions; aerial pods increase dissemination within suitable habitats, while underground pods increase the probability of plant survival under adverse conditions such as drought and heavy grazing. Underground vetch has two potential uses, namely the rehabilitation of marginal areas and production in rotation with barley. [source] Origin and palaeo-environments of calcareous sediments in the Moshaweng dry valley, southeast BotswanaEARTH SURFACE PROCESSES AND LANDFORMS, Issue 6 2002S. Ringrose Abstract Quaternary sedimentation in the Moshaweng dry valley of southeastern Botswana is evaluated on the basis of geomorphological evolution and sedimentological analyses. Stratigraphic evidence reveals an upper surface (1095 m) containing abundant sil-calcrete, an intermediate surface (1085 m) in which sil-calcrete underlies nodular calcrete and lower (1075 m) surface in which sil-calcrete and nodular calcrete are interbedded. This subdivision is reflected in the geochemical composition of the sediments which show an overall trend of decreasing SiO2 content (and increasing CaCO3 content) with depth from the highest to the lowest surface levels. The calcretes and sil-calcretes represent modifications of pre-existing detrital Kalahari Group sand and basal Kalahari pebbles which thinned over a Karoo bedrock high. Modification took place during wet periods when abundant Ca++ -rich groundwater flowed along the structurally aligned valley system. With the onset of drier conditions, water table fluctuations led to the precipitation of nodular calcretes in the phreatic layer to a depth of about 20 m. A major geochemical change resulted in the preferential silicification of the nodular calcrete deposits. Conditions for silica mobilization may be related to drying-induced salinity and in situ geochemical differentiation brought about by pebble dissociation towards the top of the sediment pile. As calcretization and valley formation progressed to lower levels, silica release took place on a diminishing scale. Thermoluminescence dating infers a mid-Pleistocene age for sil-calcrete formation suggesting that valley evolution and original calcrete precipitation are much older. Late stage dissolution of CaCO3 from pre-existing surface calcretes or sil-calcretes led to the formation of pedogenic case-hardened deposits during a time of reduced flow through the Moshaweng system possibly during the upper or late Pleistocene. Copyright © 2002 John Wiley & Sons, Ltd. [source] Ecohydrology of a semi-arid forest: partitioning among water balance components and its implications for predicted precipitation changesECOHYDROLOGY, Issue 2 2010Naama Raz Yaseef Abstract The distribution of precipitation inputs into different hydrological components of water-limited forest ecosystems determines water availability to trees and consequently forest productivity. We constructed a complete hydrological budget of a semi-arid pine forest (285 mm annual precipitation) by directly measuring its main components: precipitation (P), soil water content, evapotranspiration (ET, eddy covariance), tree transpiration (sap flux), soil evaporation (soil chambers), and intercepted precipitation (calculated). Our results indicated that on average for the 4-year study period, ET accounted for 94% of P, varying between 100% when P < 250 mm and 85% when P > 300 mm (with indications for losses to subsurface flow and soil moisture storage in wetter years). Direct measurements of the components of the ET flux demonstrated that both transpiration and soil evaporation were significant in this dry forest (45% and 36% of ET, respectively). Comparison between ecosystem ET (eddy covariance measurements) and the sum of its measured components showed good agreement on annual scales, but up to 30% discrepancies (in both directions) on shorter timescales. The pulsed storm pattern, characteristics of semi-arid climates, was sufficient to maintain the topsoil layer wet during the whole wet season. Only less often and intensive storms resulted in infiltration to the root zone, increasing water availability for uptake by deeper roots. Our results indicate that climate change predictions that link reduced precipitation with increased storm intensity may have a smaller effect on water availability to forest ecosystems than reduced precipitation alone, which could help forests' survival and maintain productivity even under drier conditions. Copyright © 2009 John Wiley & Sons, Ltd. [source] Patterns of abundance of fire ants and native ants in a native ecosystemECOLOGICAL ENTOMOLOGY, Issue 4 2009KATHARINE L. STUBLE Abstract 1.,This correlational study examines the relationship between the red imported fire ant (Solenopsis invicta) and native ants in a longleaf pine savanna. Fire ants are frequently associated with a decline in native ants throughout the invaded range, but fire ant invasion is often coupled with habitat disturbance. Invasion of fire ants into the longleaf pine savanna provides an opportunity to examine the structure of the ant community in the absence of habitat disturbance. 2.,Pitfall trapping was conducted within the longleaf pine savanna as well as across a naturally occurring soil moisture gradient, in plots that had been artificially watered. 3.,Species richness did not vary as a function of fire ant density. There was an inverse relationship between native ant density and fire ant density, but this abundance pattern does not necessarily imply a causal link between fire ant invasion and native ant decline. For individual species, fire ant densities were negatively correlated with the densities of only two native ant species, including Solenopsis carolinensis, a native species that potentially limits the invasion of fire ants. Additionally, fire ants and native ants respond differently to soil moisture, with native ants favouring drier conditions than fire ants. 4.,The possible exclusion of fire ants by some native ants, as well as differences in habitat preferences, provide alternative explanations for the frequently observed negative correlation between fire ants and native ants. [source] What caused the mid-Holocene forest decline on the eastern Tibet-Qinghai Plateau?GLOBAL ECOLOGY, Issue 2 2010Ulrike Herzschuh ABSTRACT Aim, Atmospheric CO2 concentrations depend, in part, on the amount of biomass locked up in terrestrial vegetation. Information on the causes of a broad-scale vegetation transition and associated loss of biomass is thus of critical interest for understanding global palaeoclimatic changes. Pollen records from the north-eastern Tibet-Qinghai Plateau reveal a dramatic and extensive forest decline beginning c. 6000 cal. yr bp. The aim of this study is to elucidate the causes of this regional-scale change from high-biomass forest to low-biomass steppe on the Tibet-Qinghai Plateau during the second half of the Holocene. Location, Our study focuses on the north-eastern Tibet-Qinghai Plateau. Stratigraphical data used are from Qinghai Lake (3200 m a.s.l., 36°32,,37°15, N, 99°36,,100°47, E). Methods, We apply a modern pollen-precipitation transfer function from the eastern and north-eastern Tibet-Qinghai Plateau to fossil pollen spectra from Qinghai Lake to reconstruct annual precipitation changes during the Holocene. The reconstructions are compared to a stable oxygen-isotope record from the same sediment core and to results from two transient climate model simulations. Results, The pollen-based precipitation reconstruction covering the Holocene parallels moisture changes inferred from the stable oxygen-isotope record. Furthermore, these results are in close agreement with simulated model-based past annual precipitation changes. Main conclusions, In the light of these data and the model results, we conclude that it is not necessary to attribute the broad-scale forest decline to human activity. Climate change as a result of changes in the intensity of the East Asian Summer Monsoon in the mid-Holocene is the most parsimonious explanation for the widespread forest decline on the Tibet-Qinghai Plateau. Moreover, climate feedback from a reduced forest cover accentuates increasingly drier conditions in the area, indicating complex vegetation,climate interactions during this major ecological change. [source] A conceptual model for understanding rainfall variability in the West African Sahel on interannual and interdecadal timescalesINTERNATIONAL JOURNAL OF CLIMATOLOGY, Issue 14 2001S.E. Nicholson Abstract This article describes and validates a new conceptual model for understanding Sahel rainfall variability. This conceptual model provides a framework that can readily incorporate and synthesize the roles played by the oceans, the African landmass and local meteorological factors. The most important ,local' factors are the location of the African Easterly Jet (AEJ) and the associated shears. The position of the AEJ helps to distinguish between a ,wet mode' and a ,dry mode' in the Sahel, while other factors determine which of two spatial patterns prevail during years of the dry regime. We test the paradigm by contrasting selected circulation parameters for the years 1958,1967 (representing the wet mode) and 1968,1997 (representing the dry mode). In doing so, we have identified several changes in the general atmospheric circulation that have accompanied the shift to drier conditions. The AEJ is further southward and more intense, the Inter-tropical Convergence Zone (ITCZ) is further south, the Tropical Easterly Jet (TEJ) is weaker, the equatorial westerlies are shallower and weaker, the southwesterly monsoon flow is weaker, and the relative humidity is lower (but not consistently so). The results of this study suggest that the key factor controlling the occurrence of the ,wet Sahel' mode versus the ,dry' mode is the presence of deep, well-developed equatorial westerlies. These displace the AEJ northward into Sahelian latitudes and increase the shear instabilities. The westerlies appear to be at least partially responsible for the well-known association between a weaker AEJ and wetter conditions in the Sahel, because the thermal wind induced by the Sahara/Atlantic temperature gradient is imposed upon a westerly basic state. Since one of the strongest contrasts between the ,wet Sahel' and ,dry Sahel' modes is the strength of the TEJ, the TEJ probably also plays a pivotal role in rainfall variability. In the dry mode, the equatorial westerlies are poorly developed and the core of the AEJ lies well to the south of the Sahel. The dry mode consists of two basic spatial patterns, depending on whether the Guinea Coast Region is anomalously wet or dry (the well-known dipole and no-dipole patterns, respectively). Which occurs is determined by other factors acting to reduce the intensity of the rainbelt. One of the relevant factors appears to be sea-surface temperatures (SSTs) in the Gulf of Guinea. Copyright © 2001 Royal Meteorological Society [source] Phylogeographic analysis detects congruent biogeographic patterns between a woodland agamid and Australian wet tropics taxa despite disparate evolutionary trajectoriesJOURNAL OF BIOGEOGRAPHY, Issue 8 2010Danielle L. Edwards Abstract Aim, To test the congruence of phylogeographic patterns and processes between a woodland agamid lizard (Diporiphora australis) and well-studied Australian wet tropics fauna. Specifically, to determine whether the biogeographic history of D. australis is more consistent with a history of vicariance, which is common in wet tropics fauna, or with a history of dispersal with expansion, which would be expected for species occupying woodland habitats that expanded with the increasingly drier conditions in eastern Australia during the Miocene,Pleistocene. Location, North-eastern Australia. Methods, Field-collected and museum tissue samples from across the entire distribution of D. australis were used to compile a comprehensive phylo-geographic dataset based on c. 1400 bp of mitochondrial DNA (mtDNA), incorporating the ND2 protein-coding gene. We used phylogenetic methods to assess biogeographic patterns within D. australis and relaxed molecular clock analyses were conducted to estimate divergence times. Hierarchical Shimodaira,Hasegawa tests were used to test alternative topologies representing vicariant, dispersal and mixed dispersal/vicariant biogeographic hypotheses. Phylogenetic analyses were combined with phylogeographic analyses to gain an insight into the evolutionary processes operating within D. australis. Results, Phylogenetic analyses identified six major mtDNA clades within D. australis, with phylogeographic patterns closely matching those seen in many wet tropics taxa. Congruent phylogeographic breaks were observed across the Black Mountain Corridor, Burdekin and St Lawrence Gaps. Divergence amongst clades was found to decrease in a north,south direction, with a trend of increasing population expansion in the south. Main conclusions, While phylogeographic patterns in D australis reflect those seen in many rain forest fauna of the wet tropics, the evolutionary processes underlying these patterns appear to be very different. Our results support a history of sequential colonization of D. australis from north to south across major biogeographic barriers from the late Miocene,Pleistocene. These patterns are most likely in response to expanding woodland habitats. Our results strengthen the data available for this iconic region in Australia by exploring the understudied woodland habitats. In addition, our study shows the importance of thorough investigations of not only the biogeographic patterns displayed by species but also the evolutionary processes underlying such patterns. [source] ORIGINAL ARTICLE: Towards an understanding of the Holocene distribution of Fagus sylvatica L.JOURNAL OF BIOGEOGRAPHY, Issue 1 2007Thomas Giesecke Abstract Aim, Understanding the driving forces and mechanisms of changes in past plant distribution and abundance will help assess the biological consequences of future climate change scenarios. The aim of this paper is to investigate whether modelled patterns of climate parameters 6000 years ago can account for the European distribution of Fagus sylvatica at that time. Consideration is also given to the role of non-climatic parameters as driving forces of the Holocene spread and population expansion of F. sylvatica. Location, Europe. Methods, European distributions were simulated using a physiologically-based bioclimatic model (STASH) driven by three different atmospheric general circulation model (AGCM) outputs for 6000 years ago. Results, The three simulations generally showed F. sylvatica to have potentially been as widespread 6000 years ago as it is today, which gives a profound mismatch with pollen-based reconstructions of the F. sylvatica distribution at that time. The results indicate that drier conditions during the growing season 6000 years ago could have caused a restriction of the range in the south. Poorer growth conditions with consequently reduced competitive ability were modelled for large parts of France. Main conclusions, Consideration of the entire European range of F. sylvatica showed that no single driving force could account for the observed distributional limits 6000 years ago, or the pattern of spread during the Holocene. Climatic factors, particularly drought during the growing season, are the likely major determinants of the potential range. Climatic factors are regionally moderated by competition, disturbance effects and the intrinsically slow rate of population increase of F. sylvatica. Dynamic vegetation modelling is needed to account for potentially important competitive interactions and their relationship with changing climate. We identify uncertainties in the climate and pollen data, as well as the bioclimatic model, which suggest that the current study does not identify whether or not climate determined the distribution of F. sylvatica 6000 years ago. Pollen data are better suited for comparison with relative abundance gradients rather than absolute distributional limits. These uncertainties from a study of the past, where we have information about plant distribution and abundance, argue for extreme caution in making forecasts for the future using equilibrium models. [source] Pollen-based reconstructions of biome distributions for Australia, Southeast Asia and the Pacific (SEAPAC region) at 0, 6000 and 18,000 14C yr BPJOURNAL OF BIOGEOGRAPHY, Issue 9 2004Elizabeth J. Pickett Abstract Aim, This paper documents reconstructions of the vegetation patterns in Australia, Southeast Asia and the Pacific (SEAPAC region) in the mid-Holocene and at the last glacial maximum (LGM). Methods, Vegetation patterns were reconstructed from pollen data using an objective biomization scheme based on plant functional types. The biomization scheme was first tested using 535 modern pollen samples from 377 sites, and then applied unchanged to fossil pollen samples dating to 6000 ± 500 or 18,000 ± 1000 14C yr bp. Results, 1. Tests using surface pollen sample sites showed that the biomization scheme is capable of reproducing the modern broad-scale patterns of vegetation distribution. The north,south gradient in temperature, reflected in transitions from cool evergreen needleleaf forest in the extreme south through temperate rain forest or wet sclerophyll forest (WSFW) and into tropical forests, is well reconstructed. The transitions from xerophytic through sclerophyll woodlands and open forests to closed-canopy forests, which reflect the gradient in plant available moisture from the continental interior towards the coast, are reconstructed with less geographical precision but nevertheless the broad-scale pattern emerges. 2. Differences between the modern and mid-Holocene vegetation patterns in mainland Australia are comparatively small and reflect changes in moisture availability rather than temperature. In south-eastern Australia some sites show a shift towards more moisture-stressed vegetation in the mid-Holocene with xerophytic woods/scrub and temperate sclerophyll woodland and shrubland at sites characterized today by WSFW or warm-temperate rain forest (WTRF). However, sites in the Snowy Mountains, on the Southern Tablelands and east of the Great Dividing Range have more moisture-demanding vegetation in the mid-Holocene than today. South-western Australia was slightly drier than today. The single site in north-western Australia also shows conditions drier than today in the mid-Holocene. Changes in the tropics are also comparatively small, but the presence of WTRF and tropical deciduous broadleaf forest and woodland in the mid-Holocene, in sites occupied today by cool-temperate rain forest, indicate warmer conditions. 3. Expansion of xerophytic vegetation in the south and tropical deciduous broadleaf forest and woodland in the north indicate drier conditions across mainland Australia at the LGM. None of these changes are informative about the degree of cooling. However the evidence from the tropics, showing lowering of the treeline and forest belts, indicates that conditions were between 1 and 9 °C (depending on elevation) colder. The encroachment of tropical deciduous broadleaf forest and woodland into lowland evergreen broadleaf forest implies greater aridity. Main conclusions, This study provides the first continental-scale reconstruction of mid-Holocene and LGM vegetation patterns from Australia, Southeast Asia and the Pacific (SEAPAC region) using an objective biomization scheme. These data will provide a benchmark for evaluation of palaeoclimate simulations within the framework of the Palaeoclimate Modelling Intercomparison Project. [source] Australian biogeographical connections and the phylogeny of large genera in the plant family MyrtaceaeJOURNAL OF BIOGEOGRAPHY, Issue 7 2003Pauline Y. Ladiges Abstract Aim To compare the phylogeny of the eucalypt and melaleuca groups with geological events and ages of fossils to discover the time frame of clade divergences. Location Australia, New Caledonia, New Guinea, Indonesian Archipelago. Methods We compare published molecular phylogenies of the eucalypt and melaleuca groups of the plant family Myrtaceae with geological history and known fossil records from the Cretaceous and Cenozoic. Results The Australasian eucalypt group includes seven genera, of which some are relictual rain forest taxa of restricted distribution and others are species-rich and widespread in drier environments. Based on molecular and morphological data, phylogenetic analyses of the eucalypt group have identified two major clades. The monotypic Arillastrum endemic to New Caledonia is related in one clade to the more species-rich Angophora, Corymbia and Eucalyptus that dominate the sclerophyll vegetation of Australia. Based on the time of rifting of New Caledonia from eastern Gondwana and the age of fossil eucalypt pollen, we argue that this clade extends back to the Late Cretaceous. The second clade includes three relictual rain forest taxa, with Allosyncarpia from Arnhem Land the sister taxon to Eucalyptopsis of New Guinea and the eastern Indonesian archipelago, and Stockwellia from the Atherton Tableland in north-east Queensland. As monsoonal, drier conditions evolved in northern Australia, Arnhem Land was isolated from the wet tropics to the east and north during the Oligocene, segregating ancestral rain forest biota. It is argued also that the distribution of species in Eucalyptopsis and Eucalyptus subgenus Symphyomyrtus endemic in areas north of the stable edge of the Australian continent, as far as Sulawesi and the southern Philippines, is related to the geological history of south-east Asia-Australasia. Colonization (dispersal) may have been aided by rafting on micro-continental fragments, by accretion of arc terranes onto New Guinea and by land brought into closer proximity during periods of low sea-level, from the Late Miocene and Pliocene. The phylogenetic position of the few northern, non-Australian species of Eucalyptus subgenus Symphyomyrtus suggests rapid radiation in the large Australian sister group(s) during this time frame. A similar pattern, connecting Australia and New Caledonia, is emerging from phylogenetic analysis of the Melaleuca group (Beaufortia suballiance) within Myrtaceae, with Melaleuca being polyphyletic. Main conclusion The eucalypt group is an old lineage extending back to the Late Cretaceous. Differentiation of clades is related to major geological and climatic events, including rifting of New Caledonia from eastern Gondwana, development of monsoonal and drier climates, collision of the northern edge of the Australian craton with island arcs and periods of low sea level. Vicariance events involve dispersal of biota. [source] Hunter-gatherer response to late Holocene climatic variability in northern and central AustraliaJOURNAL OF QUATERNARY SCIENCE, Issue 6 2010Alan N. Williams Abstract Sum probability analysis of 1275 radiometric ages from 608 archaeological sites across northern and central Australia demonstrates a changing archaeological signature that can be closely correlated with climate variability over the last 2 ka. Results reveal a marked increase in archaeological records across northern and central Australia over the last 2 ka, with notable declines in western and northern Australia between ca. AD 700 and 1000 and post-AD 1500 , two periods broadly coeval with the Medieval Climatic Anomaly and the Little Ice Age as they have been documented in the Asia,Pacific region. Latitudinal and longitudinal analysis of the dataset suggests the increase in archaeological footprint was continent wide, while the declines were greatest from 9 to 20° S, 110 to 135° E and 143 to 150° E. The change in the archaeological data suggests that, combined with an increase in population over the late Holocene, a disruption or reorganisation of pre-European resource systems occurred across Australia between ca. AD 700 and 1000 and post-AD 1500. These archaeological responses can be broadly correlated with transitions of the El Nińo,Southern Oscillation (ENSO) mean state on a multi-decadal to centennial timescale. The latter involve a shift towards the La Nińa-like mean state with wetter conditions in the Australian region between AD 700 and 1150. A transition period in ENSO mean state occurred across Australia during AD 1150,1300, with persistent El Nińo-like and drier conditions to ca. AD 1500, and increasing ENSO variability post-AD 1500 to the present. Copyright © 2010 John Wiley & Sons, Ltd. [source] Holocene climate change in the eastern Mediterranean region: a comparison of stable isotope and pollen data from Lake Gölhisar, southwest Turkey,JOURNAL OF QUATERNARY SCIENCE, Issue 4 2007Warren J. Eastwood Abstract Stable isotope and pollen data from Gölhisar Gölü, a small intramontane lake located in southwest Turkey, provide complementary records of Holocene climate change. Modern oxygen and hydrogen isotope water data are used as a means of comparing present-day isotope composition of the lake water to the past oxygen isotope composition of the lake water as calculated from 18O/16O ratios in calcite precipitated in the summer months. Despite the lake system being chemically dilute, the modern isotope data clearly establish that the lake water is evaporated in relation to its spring input, suggesting that the palaeo data can be interpreted primarily in terms of changing precipitation/evaporation ratios. ,18O and ,13C values from authigenic calcite through the Holocene show predominantly negative values indicating climatic conditions wetter than today. Particularly notable are low (depleted) isotope values during the earliest Holocene (ca. 10,600,8800 cal. yr. BP), a period for which pollen data imply drier conditions than at present. This divergence between pollen-inferred and stable isotope palaeoclimate data is found in other east Mediterranean lake sediment records, and suggests that vegetation may have taken several millennia to reach climatic equilibrium at the start of the Holocene. Isotopic fluctuations during the early-to-mid Holocene (8800,5100 cal. yr. BP) suggest oscillations between aridity and humidity. Higher ,18O and ,13C values for the second half of the Holocene indicate generally drier conditions than during the period before ca.5100 cal. yr BP although there is some evidence for increased humidity coinciding with pollen evidence for increasing human impact and intensification of agriculture, notably during the so-called Bey,ehir Occupation Phase (Classical and early Byzantine periods). The modern trend towards aridity started about 1300 yr ago. Copyright © 2006 John Wiley & Sons, Ltd. [source] Tephra-linked peat humification records from Irish ombrotrophic bogs question nature of solar forcing at 850 cal. yr BC,JOURNAL OF QUATERNARY SCIENCE, Issue 1 2006G. Plunkett Abstract This paper investigates evidence for palaeoclimatic changes during the period ca. 1500,500,cal.,yr BC through peat humification studies on seven Irish ombrotrophic bogs. The sites are well-correlated by the identification of three mid-first millennium BC tephras, which enable the humification records at specific points in time to be directly compared. Phases of temporarily increased wetness are suggested at ca. 1300,1250,cal.,yr BC, ca. 1150,1050 cal.,yr BC, ca. 940,cal.,yr BC and ca. 740,cal.,yr BC. The last of these is confirmed to be synchronous at five sites, suggesting external forcing on a regional scale. The timing of this wet-shift is constrained by two closely dated tephras and is demonstrated to be distinct from the widely reported changes to cooler/wetter conditions associated with a solar minimum at 850,760,cal.,yr BC, at which time the Irish sites appear instead to experience drier conditions. The results suggest the possibility of either non-uniform responses to solar forcing in northwest Europe at this time, or the existence of unrelated climate events in the early first millennium BC. The findings caution against the correlation of loosely dated palaeoclimate data if the effects of forcing mechanisms are to be understood. Copyright © 2006 John Wiley & Sons, Ltd. [source] A late Holocene record of arid events from the Cuzco region, PeruJOURNAL OF QUATERNARY SCIENCE, Issue 6 2003Alex Chepstow-Lusty Abstract The small recently infilled lake basin of Marcacocha (13°13,S, 72°12,W, 3355,m) in the Cuzco region of Peru has a morphology and location that renders it extremely sensitive to environmental change. A record of vegetation, human impact and climatic change during the past 4200,yr has been obtained from a highly organic core taken from the centre of the basin. Sustained arid episodes that affected the Peruvian Andes may be detectable using the proxy indicator of sedge (Cyperaceae) pollen abundances. As the lake-level was lowered during sustained drier conditions, the local catchment was colonised by Cyperaceae, whereas during lake floods, they retreated or were submerged and pollen production was correspondingly reduced. Drier episodes during prehistoric times occurred around 900,bc, 500,bc, ad 100 and ad 550, with a longer dry episode occurring from ad 900 to 1800. Evidence from the independently derived Quelccaya ice-core record and the archaeological chronology for the Cuzco region appears to support the climatic inferences derived from the sedge data. Many of these aridity episodes appear to correspond with important cultural changes in the Cuzco region and elsewhere in the Central Andes. Copyright © 2003 John Wiley & Sons, Ltd. [source] Spatial patterns of desert annuals in relation to shrub effects on soil moistureJOURNAL OF VEGETATION SCIENCE, Issue 2 2010J. Li Abstract Questions: What are the effects of a shrub (Haloxylon ammodendron) on spatial patterns of soil moisture in different seasons? How does productivity of understorey annuals respond to these effects? Are such effects always positive for annuals under shrubs? Location: South Gurbantunggut Desert, northwest China. Methods: Using geostatistics, we explored seasonal patterns of topsoil moisture in a 12 × 9-m plot over the growing season. To determine spatial patterns of understorey annuals in response to H. ammodendron presence, biomass of annuals was recorded in four 0.2 × 5.0-m transects from the centre of a shrub to the space between shrubs (interspace). We also investigated vertical distribution of root biomass for annuals and soil moisture dynamics across soil profiles in shrub-canopied areas and interspaces. Results: Topsoil moisture changed from autocorrelation in the wet spring to random structure in the dry season, while soil moisture below 20 cm was higher in shrub-canopied areas. Across all microhabitats, soil moisture in upper soil layers was higher than in deeper soil layers during the spring wet season, but lower during summer drought. Topsoil was close to air-dry during the dry season and developed a ,dry sand layer' that reduced evaporative loss of soil water from deeper layers recharged by snowmelt in spring. Aboveground biomass of understorey annuals was lowest adjacent to shrub stems and peaked at the shrub margin, forming a ,ring' of high herbaceous productivity surrounding individual shrubs. To acclimate to drier conditions, annuals in interspaces invested more root biomass in deeper soil with a root/shoot ratio (R/S) twice that in canopied areas. Conclusions: Positive and negative effects of shrubs on understorey plants in arid ecosystems are commonly related to nature of the environmental stress and tested species. Our results suggest there is also microhabitat-dependence in the Gurbantunggut Desert. Soil water under H. ammodendron is seasonally enriched in topsoil and deeper layers. Understorey annuals respond to the effect of shrubs on soil water availability with lower R/S and less root biomass in deeper soil layers and develop a ,ring' of high productivity at the shrub patch margin where positive and negative effects of shrubs are balanced. [source] Radial growth responses to drought of Pinus sylvestris and Quercus pubescens in an inner-Alpine dry valleyJOURNAL OF VEGETATION SCIENCE, Issue 6 2007Pascale Weber Abstract Question: Lower montane treeline ecotones such as the inner Alpine dry valleys are regarded as sensitive to climate change. In the dry Valais valley (Switzerland) the composition of the widespread, low altitude Pinus forests is shifting towards a mixed deciduous state. The sub-boreal P. sylvestris shows high mortality rates, whereas the deciduous sub-mediterranean Quercus pubescens is spreading. These species may act as early indicators of climate change. We evaluate this hypothesis by focusing on their differences in drought tolerance, which are hardly known, but are likely to be crucial in the current forest shift and also for future forest development. Methods: We used dendroecological methods to detect species-specific patterns in the growth response to drought. The relationship between radial growth of 401 trees from 15 mixed stands and drought was analysed by calculating response functions using yearly tree-ring indices and monthly drought indices. PCA was applied to the response ratios to discover spatial patterns of drought response. Results: A species-specific response to moisture as well as a sub-regional differentiation of the response patterns were found. While Quercus showed a response mainly to the conditions of the previous autumn and those of current spring, Pinus did not start responding before May, but showed responses throughout the whole summer. Quercus may restrict physiological activity to moist periods; growth of Pinus was much more dependent on prior growth. Conclusions: Given that the climate is changing towards (1) longer summer drought periods, (2) higher mean temperatures and (3) shifted seasonally of moisture availability, Quercus may benefit from adapting better to drier conditions. Pinus may increasingly face problems related to drought stress as it depends on summer moisture and has a smaller adaptive capacity due to its long-lived photosynthetic tissue. [source] Effects of altered water regimes on forest root systemsNEW PHYTOLOGIST, Issue 1 2000J. D. JOSLIN How ecosystems adapt to climate changes depends in part on how individual trees allocate resources to their components. A review of research using tree seedlings provides some support for the hypothesis that some tree species respond to exposure to drought with increases in root,shoot ratios but little change in total root biomass. Limited research on mature trees over moderately long time periods (2,10 yr), has given mixed results with some studies also providing evidence for increases in root: shoot ratios. The Throughfall Displacement Experiment (TDE) was designed to simulate both an increase and a decrease of 33% in water inputs to a mature deciduous forest over a number of years. Belowground research on TDE was designed to examine four hypothesized responses to long-term decreases in water availability; (1) increases in fine-root biomass, (2) increases in fine root,foliage ratio, (3) altered rates of fine-root turnover (FRT), and (4) depth of rooting. Minirhizotron root elongation data from 1994 to 1998 were examined to evaluate the first three hypotheses. Differences across treatments in net fine-root production (using minirhizotron root elongation observations as indices of biomass production) were small and not significant. Periods of lower root production in the dry treatment were compensated for by higher growth during favorable periods. Although not statistically significant, both the highest production (20 to 60% higher) and mortality (18 to 34% higher) rates were found in the wet treatment, resulting in the highest index of FRT. After 5 yr, a clear picture of stand fine-root-system response to drought exposure has yet to emerge in this forest ecosystem. Our results provide little support for either an increase in net fine-root production or a shift towards an increasing root,shoot ratio with long-term drought exposure. One possible explanation for higher FRT rates in the wet treatment could be a positive relationship between FRT and nitrogen and other nutrient availability, as treatments have apparently resulted in increased immobilization of nutrients in the forest floor litter under drier conditions. Such hypotheses point to the continued need to study the interactions of water stress, nutrient availability and carbon-fixation efficiency in future long-term studies. [source] Stratigraphic and environmental implications of a large ice-wedge cast at Tjćreborg, DenmarkPERMAFROST AND PERIGLACIAL PROCESSES, Issue 1 2004Else Kolstrup Abstract Exceptionally large ice-wedge casts and composite-wedge casts occur together with involutions (cryoturbations) in a gravel pit near Tjćreborg, western Jutland. The filling reveals distinctly different, vertically-orientated sedimentary units, suggesting discrete events. Variations in wedge structure and infill between the different exposures suggest differences with alternating pools and drier conditions over the former wedges. In an attempt to date and correlate crack development a perusal of local glacial history is given and optically-stimulated-luminescence (OSL) dates are presented. A review of Saalian and early Weichselian wedge casts and deep involutions in other areas in northwest Europe is provided for correlation. It is suggested that the cracking in Tjćreborg took place during the Saale or/and early Weichselian. The existence of such old wedges shows that the present land surface has probably existed since the Saalian, leaving the ground available for cracking and infilling during succeeding periods of permafrost. Copyright © 2004 John Wiley & Sons, Ltd. [source] Reproduction of an early-flowering Mediterranean mountain narrow endemic (Armeria caespitosa) in a contracting mountain islandPLANT BIOLOGY, Issue 4 2009R. García-Camacho Abstract Reproduction at population lower edges is important for plant species persistence, especially in populations on contracting high-mountain islands. In this context, the ability of plants to reproduce in different microhabitats seems to be important to guarantee seed production in stressful environments, such as Mediterranean high mountains. We hypothesised that the warmer and drier conditions at the lower edge would be deleterious for the reproduction of Armeria caespitosa, an early-flowering plant. In addition, reproductive plasticity along this mountain gradient may also be microhabitat-dependent. We studied factors affecting the reproductive success of A. caespitosa, an endemic of the Spanish Sistema Central. We considered a complex set of predictors, including phenology, plant size and environmental factors at different scales using generalised estimating equations and generalised linear models. Microhabitat, together with the position in the altitudinal gradient and inter-annual variability affected the reproduction of A. caespitosa. In addition, individuals with longer flowering periods (duration of flowering) had significantly lower fruit set and a higher number of unviable seeds; delayed flowering peaks favoured the production of both viable and unviable fruits. Microhabitat variability over an altitudinal range is relevant for the reproduction of A. caespitosa, and is more important at the lower edge of the altitudinal range, where the species faces the most adverse conditions. In addition, the ability to reproduce in different microhabitats might increase the chances of the species to cope with environmental uncertainties under on-going climate warming. Finally, reproduction of this early-flowering plant is constrained by summer drought, which might shape its reproductive phenology. [source] Vegetation transition following drainage in a high-latitude hyper-oceanic ecosystemAPPLIED VEGETATION SCIENCE, Issue 2 2010Anna Maria Fosaa Abstract Questions: How does draining affect the composition of vegetation? Are certain functional groups favoured? Can soil parameters explain these differences? Location: Central Faroe Islands, treeless islands in the northern boreal vegetation zone. Since 1987, an area of 21 km2 at 100,200 m a.s.l. was drained in order to provide water for hydro-electric production. Method: Vegetation and soil of a drained area and a control, undrained neighbouring area of approximately the same size were sampled in 2007. Six sites were sampled in each area. The vegetation was classified with cluster analysis. Results: Four plant communities were defined in the area: Calluna vulgaris,Empetrum nigrum,Vaccinium myrtillus heath, Scirpus cespitosus,Eriophorum angustifolium blanket mire, Carex bigelowii,Racomitrium lanuginosum moss-heath, Narthecium ossifragum,Carex panacea mire. Heath was more extensively distributed within, and was the dominant community of the drained area, whereas moss-heath was more extensive in the undrained area. Blanket mire and mire had approximately the same distribution in both areas. For the blanket mire, species composition indicated drier conditions in the drained than in the undrained area. The drained area had higher frequencies of woody species and lichens, grasses had finer roots and available soil phosphate was considerably higher, whereas the undrained area had higher frequencies of grasses and sedges. Conclusion: The dominant plant communities were different in the two areas, which indicated that the blanket mire was drying in the drained area. Higher concentration of soil phosphate in the drained area also indicated increased decomposition of organic soils owing to desiccation. [source] | ||||||||||||||||