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Density Variations (density + variation)
Selected AbstractsDensity-dependent dispersal in birds and mammalsECOGRAPHY, Issue 3 2005Erik Matthysen Density-dependent dispersal can be caused by various mechanisms, from competition inducing individuals to emigrate (positive density-dependence) to social crowding effects impeding free movement (negative density-dependence). Various spatial population models have incorporated positively density-dependent dispersal algorithms, and recent theoretical models have explored the conditions for density-dependent dispersal (DD) to evolve. However, while the existence of DD is well documented in some taxa such as insects, there is no clear picture on its generality in vertebrates. Here I review the available empirical data on DD in birds and mammals, focusing mainly on variation in dispersal between years and on experimental density manipulations. Surprisingly few studies have explicitly focused on DD, and interpretation of the available data is often hampered by differences in approach, small sample sizes and/or statistical shortcomings. Positive DD was reported in 50 and 33% of the selected mammal and bird studies, respectively, while two studies on mammals (out of eight) reported negative DD. Among bird studies, DD was more often reported for emigration rates or long-distance recoveries than for average distances within finite study areas. Experimental studies manipulating densities (mainly on mammals) have consistently generated positive DD, typically showing reduced emigration in response to partial population removal. Studies that examined dispersal in relation to seasonal changes in density (small mammals only) have more often reported negative DD. Studies that compared dispersal between sites differing in density, also show a mixture of positive and negative DD. This suggests that dispersal changes in a more complex way with seasonal and spatial density variation than with annual densities, and/or that these results are confounded by other factors differing between seasons and sites, such as habitat quality. I conclude that both correlational and experimental studies support the existence of positive, rather than negative, density-dependent dispersal in birds and mammals. [source] Estrogen Receptor , Gene Polymorphisms and Peak Bone Density in Chinese Nuclear Families,JOURNAL OF BONE AND MINERAL RESEARCH, Issue 6 2003Yue-Juan Qin Abstract PBD is an important determinant of osteoporotic fractures. Few studies were performed to search for genes underlying PBD variation in Chinese populations. We tested linkage and/or association of the estrogen receptor , gene polymorphism with PBD in 401 Chinese nuclear families. This study suggests the ER-, gene may have some minor effects on PBM variation in the Chinese population. Low peak bone density (PBD) in adulthood is an important determinant of osteoporotic fractures in the elderly. PBD variation is mainly regulated by genetic factors. Extensive molecular genetics studies have been performed to search for genes underlying PBD variation, largely in whites. Few studies were performed in Chinese populations. In this study, we simultaneously test linkage and/or association of the estrogen receptor , (ER -,) gene polymorphism with PBD in 401 Chinese nuclear families (both parents plus their female children) of 1260 subjects, with the 458 children generally between 20 and 40 years of age. All the subjects were genotyped by polymerase chain reaction-restriction fragment length polymorphism (PCR-RFLP) at polymorphic PvuII and XbaI sites inside the ER -, gene. Bone mineral density was measured at the lumbar spine (L1-L4) and hip (femoral neck, trochanter, and intertrochanteric region). Raw bone mineral density values were adjusted by age, height, and weight as covariates. We detected marginally significant results for within-family association (transmission disequilibrium; p = 0.054) between the spine bone mineral density variation and the ER -, XbaI genotypes. For the hip bone mineral density variation, significant (p < 0.05) linkage results were generally found for the two intragenic markers. Analyses of the haplotypes defined by the two markers confer further evidence for linkage of the ER -, with the hip PBD variation. In conclusion, this study suggests that the ER -, gene may have minor effects on PBD variation in our Chinese population. [source] Energy density analysis of cluster size dependence of surface,molecule interactions (II): Formate adsorption onto a Cu(111) surfaceJOURNAL OF COMPUTATIONAL CHEMISTRY, Issue 8 2006Hiromi Nakai Abstract Adsorption of formate (HCOO) onto a Cu(111) surface has been treated theoretically using 18 kinds of Cun (6 , n , 56) clusters. The energy density analysis (EDA) proposed by Nakai has been adopted to examine surface,molecule interactions for different cluster sizes. EDA results for the largest model cluster Cu56 have shown that the adsorption-induced energy density variation in Cu atoms decays with distance from the adsorption site. Analysis of this decay, which can be carried out using the EDA technique, is important because it enables verification of the reliability of the model cluster used. In the case of formate adsorption onto the Cu(111) surface, it is found that at least a four-layer model cluster is necessary to treat the surface,molecule interaction with chemical accuracy. © 2006 Wiley Periodicals, Inc. J Comput Chem 27: 917,925, 2006 [source] Plastic changes in seed dispersal along ecological succession: theoretical predictions from an evolutionary modelJOURNAL OF ECOLOGY, Issue 2 2005OPHÉLIE RONCE Summary 1We use a deterministic model to explore theoretically the ecological and evolutionary relevance of plastic changes in seed dispersal along ecological succession. Our model describes the effect of changing disturbance regime, age structure, density and interspecific competition as the habitat matures, enabling us to seek the evolutionarily stable reaction norm for seed dispersal rate as a function of time elapsed since population foundation. 2Our model predicts that, in the context of ecological succession, selection should generally favour plastic strategies allowing plants to increase their dispersal rate with population age, contrary to previous predictions of models that have assumed genetically fixed dispersal strategies. 3More complex patterns can evolve showing periods with high production of dispersing seeds separated by periods of intense local recruitment. These patterns are due to the interaction of individual senescence with change in ecological conditions within sites. 4Evolution of plastic dispersal strategies affects the patterns of density variation with time since foundation and accelerates successional replacement. An interesting parallel can be drawn between the evolution of age-specific dispersal rates in successional systems and the evolution of senescence in age-structured populations. 5Seed dispersal plasticity could be a potential mechanism for habitat selection in plants and have implications for range expansion in invasive species because recently founded populations at the advancing front may show different patterns to those in the established range. [source] A finite element modified method of characteristics for convective heat transportNUMERICAL METHODS FOR PARTIAL DIFFERENTIAL EQUATIONS, Issue 3 2008Mofdi El-Amrani Abstract We propose a finite element modified method of characteristics for numerical solution of convective heat transport. The flow equations are the incompressible Navier-Stokes equations including density variation through the Boussinesq approximation. The solution procedure consists of combining an essentially non-oscillatory modified method of characteristics for time discretization with finite element method for space discretization. These numerical techniques associate the geometrical flexibility of the finite elements with the ability offered by modified method of characteristics to solve convection-dominated flows using time steps larger than its Eulerian counterparts. Numerical results are shown for natural convection in a squared cavity and heat transport in the strait of Gibraltar. Performance and accuracy of the method are compared to other published data. © 2007 Wiley Periodicals, Inc. Numer Methods Partial Differential Eq, 2008 [source] Spatio-temporal shifts in gradients of habitat quality for an opportunistic avian predatorECOGRAPHY, Issue 2 2003Fabrizio Sergio We used the conceptual framework of the theory of natural selection to study breeding habitat preferences by an opportunistic avian predator, the black kite Milvus migrans. In Europe, black kite populations are mostly found near large networks of aquatic habitats, usually considered optimal for foraging and breeding. We hypothesized that proximity to wetlands could vary among individuals and affect their fitness, and thus be subject to natural selection. We tested the hypothesis first on a population on Lake Lugano (Italian pre-Alps) which has been monitored for nine years, and then on seven other populations, each studied for four,five years, located along a continuum of habitat from large water bodies to scarce aquatic habitat of any kind. In the Lake Lugano population, black kite abundance was negatively related to distance to the lake in all the nine years of study, consistent with long-term natural selection. There was evidence of ongoing directional selection on strategic nest location in three of the years, and evidence of stabilizing selection in two years. In eight of the nine years the trend was for a linear increase in fitness with increasing proximity to the lake. At the population level, results were consistent with adaptive habitat choice in relation to the previous year's spatial variation in fitness: higher associations between fitness and distance to the lake (i.e. higher selection gradients) resulted in higher density variations in the following year, in turn related to the availability of fish, the main local prey. The progressive decline of inland pairs and increase in the density of lakeshore pairs caused a directional long-term trend of declining mean distance to the lake. Breeding near aquatic habitats was associated with higher foraging success, and higher frequency and biomass of prey deliveries to offspring. There was weak evidence of selection in other populations. The inland-wetland gradient of habitat quality may have been affected by predation risk, as estimated by density of a major predator of adults and nestlings, the eagle owl Bubo bubo. Behavioral decisions at the level of the individual probably translated into population effects on density and distribution at various spatial scales. Populations in optimal habitats showed higher density and produced six times as many young per unit space as those in sub-optimal habitats. [source] Modelling the hydraulic preferences of benthic macroinvertebrates in small European streamsFRESHWATER BIOLOGY, Issue 1 2007SYLVAIN DOLÉDEC Summary 1. Relating processes occurring at a local scale to the natural variability of ecosystems at a larger scale requires the design of predictive models both to orientate stream management and to predict the effects of larger scale disturbances such as climate changes. Our study contributes to this effort by providing detailed models of the hydraulic preferences of 151 invertebrate taxa, mostly identified at the species level. We used an extensive data set comprising 580 invertebrate samples collected using a Surber net from nine sites of second and third order streams during one, two or three surveys at each site. We used nested non-linear mixed models to relate taxon local densities to bed shear stresses estimated from FliesswasserStammTisch hemisphere numbers. 2. An average model by taxon, i.e. independent from surveys, globally explained 25% of the density variations of taxa within surveys. A quadratic relationship existed between the average preferences and the niche breadth of taxa, indicating that taxa preferring extreme hemisphere numbers had a reduced hydraulic niche breadth. A more complete model, where taxa preferences vary across surveys, globally explained 38% of the variation of taxa densities within surveys. Variations in preferences across surveys were weak for taxa preferring extreme hemisphere numbers. 3. There was a significant taxonomic effect on preferences computed from the complete model. By contrast, season, site, average hemisphere number within a survey and average density of taxa within a survey used as covariates did not consistently explain shifts in taxon hydraulic preferences across surveys. 4. The average hydraulic preferences of taxa obtained from the extensive data set were well correlated to those obtained from two additional independent data sets collected in other regions. The consistency of taxon preferences across regions supports the use of regional preference curves for estimating the impact of river management on invertebrate communities. By contrast, the hydraulic niche breadths of taxa computed from the different data sets were not related. [source] Snow density variations: consequences for ground-penetrating radarHYDROLOGICAL PROCESSES, Issue 7 2006A. Lundberg Abstract Reliable hydrological forecasts of snowmelt runoff are of major importance for many areas. Ground-penetrating radar (GPR) measurements are used to assess snowpack water equivalent for planning of hydropower production in northern Sweden. The travel time of the radar pulse through the snow cover is recorded and converted to snow water equivalent (SWE) using a constant snowpack mean density from the drainage basin studied. In this paper we improve the method to estimate SWE by introducing a depth-dependent snowpack density. We used 6 years measurements of peak snow depth and snowpack mean density at 11 locations in the Swedish mountains. The original method systematically overestimates the SWE at shallow depths (+25% for 0·5 m) and underestimates the SWE at large depths (,35% for 2·0 m). A large improvement was obtained by introducing a depth,density relation based on average conditions for several years, whereas refining this by using separate relations for individual years yielded a smaller improvement. The SWE estimates were substantially improved for thick snow covers, reducing the average error from 162 ± 23 mm to 53 ± 10 mm for depth range 1·2,2·0 m. Consequently, the introduction of a depth-dependent snow density yields substantial improvements of the accuracy in SWE values calculated from GPR data. Copyright © 2005 John Wiley & Sons, Ltd. [source] Observation of neutral density variations accompanying streamer progression across air gapsIEEJ TRANSACTIONS ON ELECTRICAL AND ELECTRONIC ENGINEERING, Issue 1 2009Tetsuo Fukuchi Member Abstract Neutral density variations accompanying progression of streamers across needle-to-rod air gaps of length 0.75 and 1 m were observed with the use of a laser shadowgraph system consisting of astronomical telescopes for beam expansion and reduction, and an acousto-optic laser deflector for high-speed imaging. The system had a spatial resolution of about 1 mm over an annular observation region of outer diameter 28 cm and inner diameter 11 cm, and a temporal resolution in the order of microseconds. The system was also used to observe neutral density variations accompanying shock waves which resulted from a spark discharge. Copyright © 2009 Institute of Electrical Engineers of Japan. Published by John Wiley & Sons, Inc. [source] Efficient implementation of the shock-fitting algorithm for the Lighthill,Whitham,Richards traffic flow modelINTERNATIONAL JOURNAL FOR NUMERICAL METHODS IN ENGINEERING, Issue 4 2008Wenqin Chen Abstract This paper firstly presents the existence and uniqueness properties of the intersection time between two neighboring shocks or between a shock and a characteristic for the analytical shock-fitting algorithm that was proposed to solve the Lighthill,Whitham,Richards (LWR) traffic flow model with a linear speed,density relationship in accordance with the monotonicity properties of density variations along a shock, which have greatly improved the robustness of the analytical shock-fitting algorithm. Then we discuss the efficient evaluation of the measure of effectiveness (MOE) of the analytical shock-fitting algorithm. We develop explicit expressions to calculate the MOE,which is the total travel time that is incurred by travelers, within the space-time region that is encompassed by the shocks and/or characteristic lines. A numerical example is used to illustrate the effectiveness and efficiency of the proposed method compared with the numerical solutions that are obtained by a fifth-order weighted essentially non-oscillatory scheme. Copyright © 2007 John Wiley & Sons, Ltd. [source] The direct simulation Monte Carlo method using unstructured adaptive mesh and its applicationINTERNATIONAL JOURNAL FOR NUMERICAL METHODS IN FLUIDS, Issue 4 2002J.-S. Wu Abstract The implementation of an adaptive mesh-embedding (h-refinement) scheme using unstructured grid in two-dimensional direct simulation Monte Carlo (DSMC) method is reported. In this technique, local isotropic refinement is used to introduce new mesh where the local cell Knudsen number is less than some preset value. This simple scheme, however, has several severe consequences affecting the performance of the DSMC method. Thus, we have applied a technique to remove the hanging node, by introducing the an-isotropic refinement in the interfacial cells between refined and non-refined cells. Not only does this remedy increase a negligible amount of work, but it also removes all the difficulties presented in the originals scheme. We have tested the proposed scheme for argon gas in a high-speed driven cavity flow. The results show an improved flow resolution as compared with that of un-adaptive mesh. Finally, we have used triangular adaptive mesh to compute a near-continuum gas flow, a hypersonic flow over a cylinder. The results show fairly good agreement with previous studies. In summary, the proposed simple mesh adaptation is very useful in computing rarefied gas flows, which involve both complicated geometry and highly non-uniform density variations throughout the flow field. Copyright © 2002 John Wiley & Sons, Ltd. [source] Effects of air pollution on natural enemies of the leaf beetle Melasoma lapponicaJOURNAL OF APPLIED ECOLOGY, Issue 2 2000Elena L. Zvereva Summary 1. ,Air pollution might have differential effects on herbivores and their natural enemies, thus changing population dynamics. Therefore, from 1993 to 1998 we studied mortality caused by parasitoids and predators to the willow-feeding leaf beetle Melasoma lapponica in the impact zone of the Severonikel nickel,copper smelter (Kola Peninsula, north-western Russia). 2. ,Densities of M. lapponica were very low at clean forest sites (below five beetles per 10-min count) but higher in polluted areas (10,340 beetles per count). There were, however, variations between study years. 3. ,Egg predation, mainly by syrphid larvae and zoophagous bugs, was higher at relatively clean sites (55·3%) than at polluted sites (22·2%). Similarly, predation on larvae by zoophagous bugs and wood ants was higher at clean sites (68·4%) than at polluted sites (32·9%). 4. ,In contrast to predation, mortality caused by the parasitoid flies Megaselia opacicornis (Phoridae) and Cleonice nitidiuscula (Tachinidae) was lower at clean sites (12·3%) than at polluted sites (35·3%). Total parasitism levels increased significantly with pollution load. 5. ,Total mortality caused by natural enemies was higher at clean sites (93·7%) than at polluted sites (79·4%) due to higher predation rates, which may partly explain increased leaf beetle density within the smelter's impact zone. The effects of predators in clean forests were confirmed by the extinction of adults of M. lapponica introduced to one of the forest sites. 6. ,Although some individual sources of mortality appeared to be density dependent (direct or inverse), the joint effect of all natural enemies was not. 7. ,Our data show that a decrease in predation can contribute to increased leaf beetle density at polluted sites. However, the overall effects of natural enemies in this case were not sufficient to account for all density variations between sites. To our knowledge this is the first study to assess how pollution affects the partitioning of mortality in herbivorous insects between predators and parasitoids. [source] Energy density of anchovy Engraulis encrasicolus in the Bay of BiscayJOURNAL OF FISH BIOLOGY, Issue 3 2009J. Dubreuil The energy density (ED) of anchovy Engraulis encrasicolus in the Bay of Biscay was determined by direct calorimetry and its evolution with size, age and season was investigated. The water content and energy density varied seasonally following opposite trends. The ED g,1 of wet mass (MW) was highest at the end of the feeding season (autumn: c. 8 kJ g,1MW) and lowest in late winter (c. 6 kJ g,1MW). In winter, the fish lost mass, which was partially replaced by water, and the energy density decreased. These variations in water content and organic matter content may have implications on the buoyancy of the fish. The water content was the major driver of the energy density variations for a MW basis. A significant linear relationship was established between ED g,1 (y) and the per cent dry mass (MD; x): y =,4·937 + 0·411x. In the light of the current literature, this relationship seemed to be not only species specific but also ecosystem specific. Calibration and validation of fish bioenergetics models require energy content measurements on fish samples collected at sea. The present study provides a first reference for the energetics of E. encrasicolus in the Bay of Biscay. [source] Integrated 3-D model from gravity and petrophysical data at the Bosumtwi impact structure, GhanaMETEORITICS & PLANETARY SCIENCE, Issue 4-5 2007Hernan UGALDE A vast amount of geoscience data is available from the pre-site surveys and the actual drilling phase. A 3-D gravity model was constructed and calibrated with the available data from the two ICDP boreholes, LB-07A and LB-08A. The 3-D gravity model results agree well with both the sediment thickness and size of the central uplift revealed by previously collected seismic data, and with the petrophysical data from the LB-08A and LB-07A core materials and the two borehole logs. Furthermore, the model exhibits lateral density variations across the structure and refines the results from previous 2.5-D modeling. An important new element of the 3-D model is that the thickness of the intervals comprising polymict lithic impact breccia and suevite, monomict lithic breccia and fractured basement is much smaller than that predicted by numerical modeling. [source] The 1999 Leonid meteor storm: verification of rapid activity variations by observations at three sitesMONTHLY NOTICES OF THE ROYAL ASTRONOMICAL SOCIETY, Issue 3 2000W. Singer We report observations of an unpredicted fine structure in the activity profile of the Leonid meteor storm of 1999 November 18. Our observations were obtained at three widely separated sites (on the Iberian peninsula, in Germany, and in northern Sweden) and with two totally different techniques (video cameras and meteor radars). The observations clearly show quasi-periodic variations of the meteor rate with temporal separations of individual maxima in the 6- to 9-min range. These temporal variations translate into spatial variations within the dust trail with scales between 10 000 and 30 000 km, depending in which reference frame or direction one chooses to compare. The times for the central three maxima as observed at the three sites agree within 2 min of each other after application of the appropriate topocentric time corrections. We consider a number of potential causes for the observed density variations within the meteor stream. [source] Parsec-scale jet in the distant gigahertz-peaked spectrum quasar PKS 0858,279ASTRONOMISCHE NACHRICHTEN, Issue 2-3 2009Y. Y. KovalevArticle first published online: 13 FEB 200 Abstract The high redshift GPS quasar PKS 0858,279 exhibits the following properties which make the source unusual. Our RATAN-600 monitoring of 1,22 GHz spectrum has detected broad-band radio variability with high amplitude and relatively short time scale. In the same time, the milliarcsecond scale structure observed in a snapshot VLBA survey turned out to be very resolved which is not expected from the fast flux density variations. We performed 1.4,22 GHz VLBA observations of this quasar in 2005,2007. It has revealed a core-jet morphology. A high Doppler factor , is suggested for the jet, its nature is discussed in this report on the basis of the multi-frequency VLBA and RATAN data collected. Synchrotron self-absorption was confirmed to be dominating at low frequencies, the magnetic field strength of the dominating jet feature is estimated of an order of 0.1, mG (© 2009 WILEY-VCH Verlag GmbH & Co. KGaA, Weinheim) [source] | ||||||||||||||||