Time Scales (time + scale)

Distribution by Scientific Domains
Distribution within Earth and Environmental Science

Kinds of Time Scales

  • annual time scale
  • decadal time scale
  • different time scale
  • evolutionary time scale
  • geological time scale
  • interannual time scale
  • longer time scale
  • microsecond time scale
  • multiple time scale
  • same time scale
  • seasonal time scale
  • short time scale
  • shorter time scale


  • Selected Abstracts


    Geologic Time Scale 2004 , why, how, and where next!

    LETHAIA, Issue 2 2004
    FELIX GRADSTEIN
    A Geologic Time Scale (GTS2004) is presented that integrates currently available stratigraphic and geochronologic information. The construction of Geologic Time Scale 2004 (GTS2004) incorporated different techniques depending on the data available within each interval. Construction involved a large number of specialists, including contributions by past and present subcommissions officers of the International Commission on Stratigraphy (ICS), geochemists working with radiogenic and stable isotopes, stratigraphers using diverse tools from traditional fossils to astronomical cycles to database programming, and geomathematicians. Anticipated advances during the next four years include formalization of all Phanerozoic stage boundaries, orbital tuning extended into the Cretaceous, standardization of radiometric dating methods and resolving poorly dated intervals, detailed integrated stratigraphy for all periods, and on-line stratigraphic databases and tools. The geochronological science community and the International Commission on Stratigraphy are focusing on these issues. The next version of the Geologic Time Scale is planned for 2008, concurrent with the planned completion of boundary-stratotype (GSSP) definitions for all international stages. [source]


    Evidence of the final drainage of the Baltic Ice Lake and the brackish phase of the Yoldia Sea in glacial varves from the Baltic Sea

    BOREAS, Issue 3 2002
    THOMAS ANDRÉN
    A clay-varve chronology based on 14 cross-correlated varve graphs from the Baltic Sea and a mean varve thickness curve has been constructed. This chronology is correlated with the Swedish Time Scale and covers the time span 11 530 to 10250 varve years BP. Two cores have been analysed for grain size, chemistry, content of diatoms and changes in colour by digital colour analysis. The final drainage of the Baltic Ice Lake is dated to c. 10800 varve years BP and registered in the cores analysed as a decrease in the content of clay. This event can be correlated with atmospheric ,14C content and might have resulted in an increase in these values recorded between 11 565 and 11 545 years BP. The results of the correlation between the varve chronology from the Baltic Sea, the Greenland GRIP ice core and the atmospheric ,14C record indicate that c. 760 years are missing in the Swedish Time Scale in the part younger than c. 10250 varve years BP. A change in colour from a brownish to grey varved glacial clay recorded c. 10770 varve years BP is found to be the result of oxygen deficiency due to an increase in the rate of sedimentation in the early Preboreal. The first brackish influence is recorded c. 10540 varve years BP in the northwestern Baltic Sea and some 90 years later in the eastern Gotland Basin. [source]


    The Logarithmic-To-Linear Shift: One Learning Sequence, Many Tasks, Many Time Scales

    MIND, BRAIN, AND EDUCATION, Issue 3 2009
    Robert S. Siegler
    ABSTRACT The relation between short-term and long-term change (also known as learning and development) has been of great interest throughout the history of developmental psychology. Werner and Vygotsky believed that the two involved basically similar progressions of qualitatively distinct knowledge states; behaviorists such as Kendler and Kendler believed that the two involved similar patterns of continuous growth; Piaget believed that the two were basically dissimilar, with only development involving qualitative reorganization of existing knowledge and acquisition of new cognitive structures. This article examines the viability of these three accounts in accounting for the development of numerical representations. A review of this literature indicated that Werner's and Vygotsky's position (and that of modern dynamic systems and information processing theorists) provided the most accurate account of the data. In particular, both changes over periods of years and changes within a single experimental session indicated that children progress from logarithmic to linear representations of numerical magnitudes, at times showing abrupt changes across a large range of numbers. The pattern occurs with representations of whole number magnitudes at different ages for different numerical ranges; thus, children progress from logarithmic to linear representations of the 0,100 range between kindergarten and second grade, whereas they make the same transition in the 0,1,000 range between second and fourth grade. Similar changes are seen on tasks involving fractions; these changes yield the paradoxical finding that young children at times estimate fractional magnitudes more accurately than adults do. Several different educational interventions based on this analysis of changes in numerical representations have yielded promising results. [source]


    A Predictive Coarse-Grained Model for Semiflexible Polymers in Specific Solvents

    MACROMOLECULAR THEORY AND SIMULATIONS, Issue 4 2010
    Sheng C. Shie
    Abstract A predictive CG model based on a conventional freely rotating chain was developed to describe semiflexible polymers on a relatively large length/time scale. Parameterization of the model requires only two material properties such as, the Kuhn length and coil density. The diameter of spherical "beads" employed in the model is used as an effective parameter that needs to be determined from preliminary data. Once determined for a particular solvent system, this parameter can then be used to model general solvent systems on a parameter-free basis. Comparison with SANS data on dilute conjugated polymer solutions reveals that the CG polymer model can well describe material properties ranging from local rodlike segments to bulk interchain aggregates. [source]


    Correlation of Density Pedestal Width and Neutral Penetration Length

    CONTRIBUTIONS TO PLASMA PHYSICS, Issue 1-3 2004
    X. Q. Xu
    Abstract Pedestal studies in DIII-D and MAST find good correlation between the width of the H-mode density barrier and the neutral penetration length [1, 2]. These results suggest that the width may be set by the combined effects of neutrals and plasma transport. This paper is a report on fluid simulations of boundary plasma using the BOUT code[3] with a neutral source added. Thus both neutral and plasma physics are treated. The plasma transport is self-consistently driven by boundary turbulence due to the resistive X-point mode, while neutrals are described by a simple analytic model. The plasma profiles are evolved on the same time scale as the turbulence for the given heat source from the core plasma and particle source from the neutrals. For prescribed neutral profiles, we find the formation of a density pedestal inside the separatrix in the L-mode even though the calculated plasma diffusion coefficients are almost radially constant and without the formation of a temperature pedestal. These results support the hypothesis that particle fueling can provide the dominant control for the size of the H-mode density barrier. The width of the density barrier decreases as the pedestal density increases which is also consistent with experimental data. (© 2004 WILEY-VCH Verlag GmbH & Co. KGaA, Weinheim) [source]


    An electro-optic monitor of the behavior of Chlamydomonas reinhardtii cilia

    CYTOSKELETON, Issue 2 2005
    Keith Josef
    Abstract The unicellular green alga Chlamydomonas reinhardtii steers through water with a pair of cilia (eukaryotic flagella). Long-term observation of the beating of its cilia with controlled stimulation is improving our understanding of how a cell responds to sensory inputs. Here we describe how to record ciliary motion continuously for long periods. We also report experiments on the network of intracellular signaling that connects the environment inputs with response outputs. Local spatial changes in ciliary response on the time scale of the underlying biochemical dynamics are observed. Near-infrared light monitors the cells held by a micropipette. This condition is tolerated well for hours, not interfering with ciliary beating or sensory transduction. A computer integrates the light stimulation of the eye of Chlamydomonas with the ciliary motion making possible long-term correlations. Measures of ciliary responses include the beating frequency, stroke velocity, and stroke duration of each cilium, and the relative phase of the cis and trans cilia. The stationarity and dependence of the system on light intensity was investigated. About 150,000,000 total beat cycles and up to 8 h on one cell have been recorded. Each beat cycle is resolved so that each asynchronous beat is detected. Responses extend only a few hundred milliseconds, but there is a persistence of momentary changes that last much longer. Interestingly, we see a response that is linear with absolute light intensity as well as different kinds of response that are clearly nonlinear, implying two signaling pathways from the cell body to the cilia. Cell Motil. Cytoskeleton 61:83,96, 2005. © 2005 Wiley-Liss, Inc. [source]


    Risk assessment for nonindigenous pests: 1.

    DIVERSITY AND DISTRIBUTIONS, Issue 5 2001
    Mapping the outputs of phenology models to assess the likelihood of establishment
    Abstract This paper demonstrates the use of phenology models mapped over the landscape as a tool in support of risk assessments for nonindigenous plant pests. Drawing on the relationship between pest development and temperature, the approach uses gridded sequential interpolated temperatures at a resolution of 1 km, linked with phenology models, to predict the potential for a pest to develop throughout the landscape. The potential for establishment of Colorado beetle (Leptinotarsa decemlineata) in England and Wales was used as an illustration. The likelihood of the pest completing a single generation during a 30-year period (1961,90) was computed. Summaries of phenology, based firstly upon point temperature series from weather stations and secondly upon temperatures interpolated across the landscape, were compared. The results revealed that the use of point data led to a 70% likelihood of over-estimating the area at risk from year to year. In the case of average long-term risk however, the point-based and landscape-wide distributions of establishment potential were similar. We demonstrate how the use of phenology models running on a daily time scale provides date based results, so allowing outputs to be tied in with periods in the cropping cycle. The application of daily data in computing the phenological results, unlike the main body of published work on pest risk assessment which uses averaged monthly data, reflects more fully the underlying variability and degrees of sensitivity of the pest to changes in weather. [source]


    The role of moisture cycling in the weathering of a quartz chlorite schist in a tropical environment: findings of a laboratory simulation

    EARTH SURFACE PROCESSES AND LANDFORMS, Issue 4 2005
    Tony Wells
    Abstract Long-term weathering of a quartz chlorite schist via wetting and drying was studied under a simulated tropical climate. Cubic rock samples (15 mm × 15 mm × 15 mm) were cut from larger rocks and subjected to time-compressed climatic conditions simulating the tropical wet season climate at the Ranger Uranium Mine in the Northern Territory, Australia. Fragmentation, moisture content and moisture uptake rate were monitored over 5000 cycles of wetting and drying. To determine the impact of climatic variables, five climatic regimes were simulated, varying water application, temperature and drying. One of the climatic regimes reproduced observed temperature and moisture variability at the Ranger Uranium Mine, but over a compressed time scale. It is shown that wetting and drying is capable of weathering quartz chlorite schist with changes expected over a real time period of decades. While wetting and drying alone does produce changes to rock morphology, the incorporation of temperature variation further enhances weathering rates. Although little fragmentation occurred in experiments, significant changes to internal pore structure were observed, which could potentially enhance other weathering mechanisms. Moisture variability is shown to lead to higher weathering rates than are observed when samples are subjected only to leaching. Finally, experiments were conducted on two rock samples from the same source having only subtle differences in mineralogy. The samples exhibited quite different weathering rates leading to the conclusion that our knowledge of the role of rock type and composition in weathering is insufficient for the accurate determination of weathering rates. Copyright © 2005 John Wiley & Sons, Ltd. [source]


    Testing a model for predicting the timing and location of shallow landslide initiation in soil-mantled landscapes

    EARTH SURFACE PROCESSES AND LANDFORMS, Issue 9 2003
    M. Casadei
    Abstract The growing availability of digital topographic data and the increased reliability of precipitation forecasts invite modelling efforts to predict the timing and location of shallow landslides in hilly and mountainous areas in order to reduce risk to an ever-expanding human population. Here, we exploit a rare data set to develop and test such a model. In a 1·7 km2 catchment a near-annual aerial photographic coverage records just three single storm events over a 45 year period that produced multiple landslides. Such data enable us to test model performance by running the entire rainfall time series and determine whether just those three storms are correctly detected. To do this, we link a dynamic and spatially distributed shallow subsurface runoff model (similar to TOPMODEL) to an in,nite slope model to predict the spatial distribution of shallow landsliding. The spatial distribution of soil depth, a strong control on local landsliding, is predicted from a process-based model. Because of its common availability, daily rainfall data were used to drive the model. Topographic data were derived from digitized 1 : 24 000 US Geological Survey contour maps. Analysis of the landslides shows that 97 occurred in 1955, 37 in 1982 and ,ve in 1998, although the heaviest rainfall was in 1982. Furthermore, intensity,duration analysis of available daily and hourly rainfall from the closest raingauges does not discriminate those three storms from others that did not generate failures. We explore the question of whether a mechanistic modelling approach is better able to identify landslide-producing storms. Landslide and soil production parameters were ,xed from studies elsewhere. Four hydrologic parameters characterizing the saturated hydraulic conductivity of the soil and underlying bedrock and its decline with depth were ,rst calibrated on the 1955 landslide record. Success was characterized as the most number of actual landslides predicted with the least amount of total area predicted to be unstable. Because landslide area was consistently overpredicted, a threshold catchment area of predicted slope instability was used to de,ne whether a rainstorm was a signi,cant landslide producer. Many combinations of the four hydrological parameters performed equally well for the 1955 event, but only one combination successfully identi,ed the 1982 storm as the only landslide-producing storm during the period 1980,86. Application of this parameter combination to the entire 45 year record successfully identi,ed the three events, but also predicted that two other landslide-producing events should have occurred. This performance is signi,cantly better than the empirical intensity,duration threshold approach, but requires considerable calibration effort. Overprediction of instability, both for storms that produced landslides and for non-producing storms, appears to arise from at least four causes: (1) coarse rainfall data time scale and inability to document short rainfall bursts and predict pressure wave response; (2) absence of local rainfall data; (3) legacy effect of previous landslides; and (4) inaccurate topographic and soil property data. Greater resolution of spatial and rainfall data, as well as topographic data, coupled with systematic documentation of landslides to create time series to test models, should lead to signi,cant improvements in shallow landslides forecasting. Copyright © 2003 John Wiley & Sons, Ltd. [source]


    Novel coupling Rosenbrock-based algorithms for real-time dynamic substructure testing

    EARTHQUAKE ENGINEERING AND STRUCTURAL DYNAMICS, Issue 3 2008
    O. S. Bursi
    Abstract Real-time testing with dynamic substructuring is a novel experimental technique capable of assessing the behaviour of structures subjected to dynamic loadings including earthquakes. The technique involves recreating the dynamics of the entire structure by combining an experimental test piece consisting of part of the structure with a numerical model simulating the remainder of the structure. These substructures interact in real time to emulate the behaviour of the entire structure. Time integration is the most versatile method for analysing the general case of linear and non-linear semi-discretized equations of motion. In this paper we propose for substructure testing, L-stable real-time (LSRT) compatible integrators with two and three stages derived from the Rosenbrock methods. These algorithms are unconditionally stable for uncoupled problems and entail a moderate computational cost for real-time performance. They can also effectively deal with stiff problems, i.e. complex emulated structures for which solutions can change on a time scale that is very short compared with the interval of time integration, but where the solution of interest changes on a much longer time scale. Stability conditions of the coupled substructures are analysed by means of the zero-stability approach, and the accuracy of the novel algorithms in the coupled case is assessed in both the unforced and forced conditions. LSRT algorithms are shown to be more competitive than popular Runge,Kutta methods in terms of stability, accuracy and ease of implementation. Numerical simulations and real-time substructure tests are used to demonstrate the favourable properties of the proposed algorithms. Copyright © 2007 John Wiley & Sons, Ltd. [source]


    Shaking table tests on seismic response of steel braced frames with column uplift

    EARTHQUAKE ENGINEERING AND STRUCTURAL DYNAMICS, Issue 14 2006
    Mitsumasa Midorikawa
    Abstract Previous studies have suggested that rocking vibration accompanied by uplift motion might reduce the seismic damage to buildings subjected to severe earthquake motions. This paper reports on the use of shaking table tests and numerical analyses to evaluate and compare the seismic response of base-plate-yielding rocking systems with columns allowed to uplift with that of fixed-base systems. The study is performed using half-scale three-storey, 1 × 2 bay braced steel frames with a total height of 5.3 m. Base plates that yield due to column tension were installed at the base of each column. Two types of base plates with different thicknesses are investigated. The earthquake ground motion used for the tests and analyses is the record of the 1940 El Centro NS component with the time scale shortened by a factor of 1/,2. The maximum input acceleration is scaled to examine the structural response at various earthquake intensities. The column base shears in the rocking frames with column uplift are reduced by up to 52% as compared to the fixed-base frames. Conversely, the maximum roof displacements of the fixed and rocking frames are about the same. It is also noted that the effect of the vertical impact on the column associated with touchdown of the base plate is small because the difference in tensile and compressive forces is primarily due to the self-limiting tensile force in the column caused by yielding of the base plate. Copyright © 2006 John Wiley & Sons, Ltd. [source]


    Dimensional response analysis of yielding structures with first-mode dominated response

    EARTHQUAKE ENGINEERING AND STRUCTURAL DYNAMICS, Issue 10 2006
    Nicos Makris
    Abstract This paper introduces a new way of estimating the inelastic response of first-mode dominated structures with behaviour that can be approximated with the elastoplastic idealization. The proposed approach emerges from formal dimensional analysis and is liberated from the response of the elastic system. The application of the proposed method hinges upon the existence of a distinct time scale and a length scale that characterize the most energetic component of the ground shaking. Such time and length scales emerge naturally from the distinguishable pulses which dominate a wide class of strong earthquake records; they are directly related with the rise time and slip velocity of faulting, and can be formally extracted with validated mathematical models published in the literature. The most decisive feature of this work is that the inelastic response curves that result with the proposed approach assume similar shapes for different values of the normalized yield displacement. Because of this similarity the paper proposes a single inelastic response curve which offers directly the maximum inelastic displacement of the structure given the energetic pulse period and pulse amplitude of the ground shaking. When the proposed method is applied to MDOF structures it is not capable to estimate interstorey drifts nor is capable to capture the effects of negative stiffness which may result due to P-delta effect. Copyright © 2006 John Wiley & Sons, Ltd. [source]


    Representing genetic variation as continuous surfaces: an approach for identifying spatial dependency in landscape genetic studies

    ECOGRAPHY, Issue 6 2008
    Melanie A. Murphy
    Landscape genetics, an emerging field integrating landscape ecology and population genetics, has great potential to influence our understanding of habitat connectivity and distribution of organisms. Whereas typical population genetics studies summarize gene flow as pairwise measures between sampling localities, landscape characteristics that influence population genetic connectivity are often continuously distributed in space. Thus, there are currently gaps in both the ability to analyze genotypic data in a continuous spatial context and our knowledge of expected of landscape genetic structure under varying conditions. We present a framework for generating continuous "genetic surfaces", evaluate their statistical properties, and quantify statistical behavior of landscape genetic structure in a simple landscape. We simulated microsatellite genotypes under varying parameters (time since vicariance, migration, effective population size) and used ancestry (q) values from STRUCTURE to interpolate a genetic surface. Using a spatially adjusted Pearson's correlation coefficient to test the significance of landscape variable(s) on genetic structure we were able to detect landscape genetic structure on a contemporary time scale (,5 generations post vicariance, migration probability ,0.10) even when population differentiation was minimal (FST,0.00015). We show that genetic variation can be significantly correlated with geographic distance even when genetic structure is due to landscape variable(s), demonstrating the importance of testing landscape influence on genetic structure. Finally, we apply genetic surfacing to analyze an empirical dataset of black bears from northern Idaho USA. We find black bear genetic variation is a function of distance (autocorrelation) and habitat patch (spatial dependency), consistent with previous results indicating genetic variation was influenced by landscape by resistance. These results suggest genetic surfaces can be used to test competing hypotheses of the influence of landscape characteristics on genetic structure without delineation of categorical groups. [source]


    Plant species response to land use change ,Campanula rotundifolia, Primula veris and Rhinanthus minor

    ECOGRAPHY, Issue 1 2005
    Regina Lindborg
    Land use change is a crucial driver behind species loss at the landscape scale. Hence, from a conservation perspective, species response to habitat degradation or improvement of habitat quality, is important to examine. By using indicator species it may be possible to monitor long-term survival of local populations associated with land use change. In this study we examined three potential indicator (response) species for species richness and composition in Scandinavian semi-natural grassland communities: Campanula rotundifolia, Primula veris and Rhinanthus minor. With field inventories and experiments we examined their response to present land use, habitat degradation and improvement of local habitat quality. At the time scale examined, C. rotundifolia was the only species responding to both habitat degradation and improvement of habitat quality. Neither R. minor nor P. veris responded positively to habitat improvements although both responded rapidly to direct negative changes in habitat quality. Even though C. rotundifolia responded quickly to habitat degradation, it did not disappear completely from the sites. Instead, the population structure changed in terms of decreased population size and flowering frequency. It also showed an ability to form remnant populations which may increase resilience of local habitats. Although P. veris and especially R. minor responded rapidly to negative environmental changes and may be useful as early indicators of land use change, it is desirable that indicators respond to both degradation and improvement of habitat quality. Thus, C. rotundifolia is a better response species for monitoring effects of land use change and conservation measures, provided that both local and regional population dynamics are monitored over a long time period. [source]


    Extinction risk under coloured environmental noise

    ECOGRAPHY, Issue 2 2000
    Mikko Heino
    Positively autocorrelated red environmental noise is characterized by a strong dependence of expected sample variance on sample length. This dependence has to be taken into account when assessing extinction risk under red and white uncorrelated environmental noise. To facilitate a comparison between red and white noise, their expected variances can be scaled to be equal, but only at a chosen time scale. We show with a simple one-dimensional population dynamics model that the different but equally reasonable choices of the time scale yield qualitatively different results on the dependence of extinction risk on the colour of environmental noise: extinction risk might increase as well as decrease when the temporal correlation of noise increases. [source]


    Testing for criticality in ecosystem dynamics: the case of Amazonian rainforest and savanna fire

    ECOLOGY LETTERS, Issue 7 2010
    Salvador Pueyo
    Ecology Letters (2010) 13: 793,802 Abstract We test for two critical phenomena in Amazonian ecosystems: self-organized criticality (SOC) and critical transitions. SOC is often presented in the complex systems literature as a general explanation for scale invariance in nature. In particular, this mechanism is claimed to underlie the macroscopic structure and dynamics of terrestrial ecosystems. These would be inextricably linked to the action of fire, which is conceived as an endogenous ecological process. We show that Amazonian savanna fires display the scale-invariant features characteristic of SOC but do not display SOC. The same is true in Amazonian rainforests subject to moderate drought. These findings prove that there are other causes of scale invariance in ecosystems. In contrast, we do find evidence of a critical transition to a megafire regime under extreme drought in rainforests; this phenomenon is likely to determine the time scale of a possible loss of Amazonian rainforest caused by climate change. [source]


    Ecological and evolutionary consequences of niche construction for its agent

    ECOLOGY LETTERS, Issue 10 2008
    Grigoris Kylafis
    Abstract Niche construction can generate ecological and evolutionary feedbacks that have been underinvestigated so far. We present an eco-evolutionary model that incorporates the process of niche construction to reveal its effects on the ecology and evolution of the niche-constructing agent. We consider a simple plant,soil nutrient ecosystem in which plants have the ability to increase the input of inorganic nutrient as an example of positive niche construction. On an ecological time scale, the model shows that niche construction allows the persistence of plants under infertile soil conditions that would otherwise lead to their extinction. This expansion of plants' niche, however, requires a high enough rate of niche construction and a high enough initial plant biomass to fuel the positive ecological feedback between plants and their soil environment. On an evolutionary time scale, we consider that the rates of niche construction and nutrient uptake coevolve in plants while a trade-off constrains their values. Different evolutionary outcomes are possible depending on the shape of the trade-off. We show that niche construction results in an evolutionary feedback between plants and their soil environment such that plants partially regulate soil nutrient content. The direct benefit accruing to plants, however, plays a crucial role in the evolutionary advantage of niche construction. [source]


    Transient response of closed-loop MHD experimental facility

    ELECTRICAL ENGINEERING IN JAPAN, Issue 1 2007
    Hidemasa Takana
    Abstract Transient responses of a closed-loop MHD experimental facility from nonpower generation to power generation have been investigated by means of time-dependent quasi-one-dimensional numerical simulations. For the long-time continuous power generation experiment, the time required to obtain the steady state for the power generation is estimated to be approximately 20 hours. By increasing the electrical input power to the heater as an exponential function of time, the temperature increment of ceramics can be moderated. When the duration of the experiment is around 10 minutes, argon gas temperature at the exit of the heater hardly changes because of the large heat capacity of structure materials. It is found that the fluid disturbances are induced at the instant of the power generation and they propagate as they repeatedly reflect at the sudden change of duct shape. Since all of the induced disturbances attenuate approximately 0.4 second after the power generation, the time scale that the disturbances exist in the facility is estimated to be 1 second at most. © 2006 Wiley Periodicals, Inc. Electr Eng Jpn, 158(1): 46,52, 2007; Published online in Wiley InterScience (www.interscience.wiley.com). DOI 10.1002/eej.20237 [source]


    Adaptive responses to antimicrobial agents in biofilms

    ENVIRONMENTAL MICROBIOLOGY, Issue 8 2005
    Barbara Szomolay
    Summary Bacterial biofilms demonstrate adaptive resistance in response to antimicrobial stress more effectively than corresponding planktonic populations. We propose here that, in biofilms, reaction-diffusion limited penetration may result in only low levels of antimicrobial exposure to deeper regions of the biofilm. Sheltered cells are then able to enter an adapted resistant state if the local time scale for adaptation is faster than that for disinfection. This mechanism is not available to a planktonic population. A mathematical model is presented to illustrate. Results indicate that, for a sufficiently thick biofilm, cells in the biofilm implement adaptive responses more effectively than do freely suspended cells. Effective disinfection requires applied biocide concentration that increases quadratically or exponentially with biofilm thickness. [source]


    Enantiomeric composition of chiral polychlorinated biphenyl atropisomers in dated sediment cores

    ENVIRONMENTAL TOXICOLOGY & CHEMISTRY, Issue 2 2007
    Charles S. Wong
    Abstract ,Enantiomer fractions (EFs) of seven chiral poly chlorinated biphenyls (PCBs) were measured in dated sediment cores of Lake Hartwell (SC, USA) and Lake Ontario (USA) to detect, quantify, and gain insight regarding microbial reductive dechlorination of PCBs in lake sediments with high and low concentrations, respectively. Lake Hartwell sediments had high total PCBs (5,60 ,g/g), with significantly nonracemic EFs that generally were consistent with those from previous laboratory microcosm reductive dechlorination experiments using sediments from these sites. Thus, stereoselective reductive dechlorination had occurred in situ, including at total PCB concentrations of less than the threshold of approximately 30 to 80 ,g/g suggested as being necessary for reductive dechlorination. Enantiomer fractions of PCBs 91, 95, 132, and 136 in Lake Hartwell cores were significantly correlated both with concentrations of those individual congeners and with total PCB concentration for some sites. This result indicates that enantioselective microbial dechlorination activity increases with higher concentrations within sediments for these congeners. Enantiomer composition reversed with depth for PCBs 91, 132, and 176, suggesting that multiple microbial populations may be present within the same core that are enantioselectively dechlorinating PCBs. Such observations indicate that concentration and time are not the only factors affecting biotransformation, complicating prediction of enantioselectivity. Comparison of EFs with dates suggested biotransformation half-lives of approximately 30 years, which is on the same time scale as sequestration by burial. In contrast, Lake Ontario sediments (maximum total PCBs, 400 ng/g) had racemic or near-racemic amounts of most congeners throughout the core profile, which is consistent with achiral indicators suggesting no microbial biotransformation within Lake Ontario sediments. Thresholds for reductive dechlorination may exist, but they would be at concentrations of less than 30 to 80 ,g/g. [source]


    COALESCENCE IN A METAPOPULATION WITH RECURRENT LOCAL EXTINCTION AND RECOLONIZATION

    EVOLUTION, Issue 5 2003
    John R. Pannell
    Abstract Many species exist as metapopulations in balance between local population extinction and recolonization. The effect of these processes on average population differentiation, within-deme diversity, and specieswide diversity has been considered previously. In this paper, coalescent simulations of Slatkin's propagule-pool and migrant-pool models are used to characterize the distribution of neutral genetic diversity within demes (,s), diversity in the metapopulation a whole (TTT), the ratio FST= (,t,,S)/,T, Tajima's D statistic, and several ratios of gene-tree branch lengths. Using these distributions, power to detect differences in key metapopulation parameter values is determined under contrasting sampling regimes. The results indicate that it will be difficult to use sequence data from a single locus to detect a history of extinctions and recolonizations in a metapopulation because of high genealogical variance, the loss of diversity due to reductions in effective population size, and the fact that a genealogy of lineages from different demes under Slatkin's model differs from a neutral coalescent only in its time scale. Genetic indices of gene-tree shape that capture the effects of extinction/recolonization on both external branches and the length of the genealogy as a whole will provide the best indication of metapopulation dynamics if several lineages are sampled from several different demes. [source]


    THE EFFECTS OF SUBDIVISION ON THE GENETIC DIVERGENCE OF POPULATIONS AND SPECIES

    EVOLUTION, Issue 4 2000
    John Wakeley
    Abstract. An island model of migration is used to study the effects of subdivision within populations and species on sample genealogies and on between-population or between-species measures of genetic variation. The model assumes that the number of demes within each population or species is large. When populations (or species), connected either by gene flow or historical association, are themselves subdivided into demes, changes in the migration rate among demes alter both the structure of genealogies and the time scale of the coalescent process. The time scale of the coalescent is related to the effective size of the population, which depends on the migration rate among demes. When the migration rate among demes within populations is low, isolation (or speciation) events seem more recent and migration rates among populations seem higher because the effective size of each population is increased. This affects the probability of reciprocal monophyly of two samples, the chance that a gene tree of a sample matches the species tree, and relative likelihoods of different types of polymorphic sites. It can also have a profound effect on the estimation of divergence times. [source]


    The effect of a closed area and beach seine exclusion on coral reef fish catches

    FISHERIES MANAGEMENT & ECOLOGY, Issue 2 2001
    T. R. McClanahan
    Fish landing data from the Mombasa Marine National Park (MNP) and a marine reserve exploited by various gears were studied over a 5-yr period to determine the influence of the closed area and different gears in fisheries. The number fishing and boats per landing site was constant, but total and catch per unit effort progressively declined in all sites on an annual basis irrespective of the existence of a marine reserve, exclusion of the beach seines or use of gear. Differences between landing sites were most pronounced when analysed on a catch per area as opposed to the more standard catch per fisherman, suggesting compensation in human effort when catches decline. A marine reserve next to a closed area that excluded beach seines had the highest catch per area (5.5 kg ha,1 month,1) despite having the highest density of fishermen (0.07 ± 0.02 fishermen ha,1 month,1). The annual rate of decline in the catch was lower than the other sites at around 250 g day,1 compared with 310,400 g day,1 in the other sites. One landing site, which excluded beach seine landings for more than 20 yrs, had a high catch per area (,5.3 kg ha,1 month,1), but after experiencing a doubling in the effort of other gears (line, speargun and trap), the catch per fisherman and area were reduced. Environmental or habitat degradation and excessive effort remain the most likely explanation for the overall declines in catch from 1995 to 1999. Closed areas and beach seine exclusion have the potential to increase catch rates, but the first often reduces the total fishing area and possibly leads to a loss of total catch, at least on a time scale of less than 10 yrs. The exclusion of beach seines can lead to an increase in other gear types that can also cause reductions in catch. [source]


    Spatial and temporal variability of the Aleutian climate

    FISHERIES OCEANOGRAPHY, Issue 2005
    SERGEI N. RODIONOV
    Abstract The objective of this paper is to highlight those characteristics of climate variability that may pertain to the climate hypothesis regarding the long-term population decline of Steller sea lions (Eumetopias jubatus). The seasonal changes in surface air temperature (SAT) across the Aleutian Islands are relatively uniform, from 5 to 10°C in summer to near freezing temperatures in winter. The interannual and interdecadal variations in SAT, however, are substantially different for the eastern and western Aleutians, with the transition found at about 170°W. The eastern Aleutians experienced a regime shift toward a warmer climate in 1977, simultaneously with the basin-wide shift in the Pacific Decadal Oscillation (PDO). In contrast, the western Aleutians show a steady decline in winter SATs that started in the 1950s. This cooling trend was accompanied by a trend toward more variable SAT, both on the inter- and intra-annual time scale. During 1986,2002, the variance of winter SATs more than doubled compared to 1965,1985. At the same time in Southeast Alaska, the SAT variance diminished by half. Much of the increase in the intra-seasonal variability for the western Aleutians is associated with a warming trend in November and a cooling trend in January. As a result, the rate of seasonal cooling from November to January has doubled since the late 1950s. We hypothesize that this trend in SAT variability may have increased the environmental stress on the western stock of Steller sea lions and hence contributed to its decline. [source]


    Short-term variability in physical forcing in temperate reservoirs: effects on phytoplankton dynamics and sedimentary fluxes

    FRESHWATER BIOLOGY, Issue 1 2007
    ALEXANDRINE PANNARD
    Summary 1. The effects of wind events on phytoplankton dynamics were investigated in two temperate reservoirs. 2. Meteorological forcing, change in physical and chemical structure of the water column and biological responses of phytoplankton communities were followed for 3 weeks in three seasons. 3. Depending on the season, the phytoplankton response differed in response to nutrient and light conditions, and to the intensity of stratification and mixing. 4. We demonstrated that, on a time scale of a few days, wind events can modify phytoplankton dynamics, in terms of size structure and exported biomass. An increase of mixing favoured the largest size class and disadvantaged the smallest size class, while an increase in stratification had the opposite effects. The short-term change in size structure was reflected in the sedimentary fluxes but with a time lag. [source]


    Top-down control of phytoplankton: the role of time scale, lake depth and trophic state

    FRESHWATER BIOLOGY, Issue 12 2002
    JÜrgen Benndorf
    SUMMARY 1.,One of the most controversial issues in biomanipulation research relates to the conditions required for top-down control to cascade down from piscivorous fish to phytoplankton. Numerous experiments have demonstrated that Phytoplankton biomass Top-Down Control (PTDC) occurs under the following conditions: (i) in short-term experiments, (ii) shallow lakes with macrophytes, and (iii) deep lakes of slightly eutrophic or mesotrophic state. Other experiments indicate that PTDC is unlikely in (iv) eutrophic or hypertrophic deep lakes unless severe light limitation occurs, and (v) all lakes characterised by extreme nutrient limitation (oligo to ultraoligotrophic lakes). 2.,Key factors responsible for PTDC under conditions (i) to (iii) are time scales preventing the development of slow-growing inedible phytoplankton (i), shallow depth allowing macrophytes to become dominant primary producers (ii), and biomanipulation-induced reduction of phosphorus (P) availability for phytoplankton (iii). 3.,Under conditions (iv) and (v), biomanipulation-induced reduction of P-availability might also occur but is insufficient to alter the epilimnetic P-content enough to initiate effective bottom-up control (P-limitation) of phytoplankton. In these cases, P-loading is much too high (iv) or P-content in the lake much too low (v) to initiate or enhance P-limitation of phytoplankton by a biomanipulation-induced reduction of P-availability. However, PTDC may exceptionally result under condition (iv) if high mixing depth and/or light attenuation cause severe light limitation of phytoplankton. 4.,Recognition of the five different conditions reconciles previous seemingly contradictory results from biomanipulation experiments and provides a sound basis for successful application of biomanipulation as a tool for water management. [source]


    Water Dynamics in Ionomer Membranes by Field-Cycling NMR Relaxometry,

    FUEL CELLS, Issue 1 2006
    J.-C. Perrin
    Abstract The water dynamics of two types of ionomer membranes, the Nafion® and sulfonated polyimides, have been investigated by field-cycling nuclear magnetic relaxation. This technique, applied to samples prepared at different water content, allows to characterize the proton motion at the time scale of the microsecond. The polyimides appear to behave as standard porous materials, whereas in the Nafion®, we have observed two different dynamical regimes related to a complex swelling process. [source]


    Human and climatic impact on late Quaternary deposition in the Sparta Basin Piedmont: Evidence from alluvial fan systems

    GEOARCHAEOLOGY: AN INTERNATIONAL JOURNAL, Issue 7 2003
    Richard J. J. Pope
    The evolution of four alluvial fans in the Evrotas Valley, southern Greece, was examined using a combination of geomorphological and sedimentological techniques. Source material for the fans is derived largely from the Taygetos Mountains, and thus the fan deposits provide proxy evidence for erosion of upland landscapes. Stratigraphic sequences exposed in the fanhead trenches suggest a progressive change in depositional style down-fan. Within the St. Johns, North Anogia, and North Xilocambi fans, debris flow deposits are gradually replaced by gravel-dominated hyperconcentrated flow deposits and then fine-grained hyperconcentrated flow deposits. Within the Kalivia Sokas fan, gravel-dominated hyperconcentrated flow deposits give way to fine-grained hyperconcentrated flow deposits, and finally to fluvial gravels. Mineral magnetic studies combined with thermoluminescence dating suggest that sedimentation also occurred over a similar time scale. Deposition cycles during the late Pleistocene appear to be climatically driven, with proximal and medial fan segments developing during stadial phases of the Riss/Würm and Würm, respectively. Distal segments aggraded during the Holocene. During interstadial episodes of the late Pleistocene, fan entrenchment occurred. Holocene accretion is likely to be related to human activity and appears to be concentrated in the early/middle Helladic and the Hellenistic periods, when population levels, indicated by increased numbers of archaeological sites, were rising. © 2003 Wiley Periodicals, Inc. [source]


    Growth of the Yellow River delta over the past 800 years, as influenced by human activities

    GEOGRAFISKA ANNALER SERIES A: PHYSICAL GEOGRAPHY, Issue 1 2003
    Xu Jiongxin
    Abstract Based on measurements made over the past 50 years and data extracted from historical documents, a study has been made of the effect of human activities on the growth of the Yellow River delta over a time scale of 102 -103 years. During the period studied, the Yellow River emptied into the Yellow Sea and Bohai Sea, successively, and the deltas it created are known as the Abandoned and Modern Yellow River deltas. The growth of the Abandoned Yellow River delta can be divided into two stages. The first extended from AD 1194 to 1578, during which the growth was rather slow. The second extended from AD 1579 to 1855, when the growth was greatly accelerated. The curve showing the temporal variation of the growth of the Modern Yellow River mouth can be fitted by three straight lines with different slopes. Accordingly, its growth from 1855 to present can be divided into three stages, and the inflection points associated with three straight lines may be regarded as threshold points, reflecting marked changes in the nature and extent of human activities. On this basis, a three-stage descriptive model has been proposed to describe the process of river mouth extension in response to changing human activities. [source]


    A high-resolution relative time scale for the Viséan Stage (Carboniferous) of the Kulm Basin (Rhenish Mountains, Germany)

    GEOLOGICAL JOURNAL, Issue 3 2009
    Dieter Korn
    Abstract The Viséan (Carboniferous) sedimentary succession of the basinal Kulm facies (Rhenish Mountains) was investigated in detail in order to achieve a high-resolution stratigraphic subdivision and correlation. Additionally, the ranges of fossil index taxa (ammonoids), fossil marker beds, volcaniclastic horizons and sedimentary features (e.g. colour changes) were integrated in the correlation. As a result, a comprehensive database was compiled, which contains 190 stratigraphic events of the Viséan interval of this area. Several sections are almost completely composed of shales, which are regarded to represent a slow but constant basinal background sedimentation of the Kulm facies. The thickness of lithological homogeneous sections thus indicates an approximately linear record of time and the average thicknesses of biozones and positions of stratigraphic events can easily be calculated from the compiled database. The result is an approximately time-linear biostratigraphic scale for the Viséan Stage of the Kulm Basin. Given a numerical length of the Viséan Stage of ca. 19,Ma, 190 stratigraphic events give a mean resolution of 100,000 years. This is unique in Palaeozoic stratigraphy. Copyright © 2008 John Wiley & Sons, Ltd. [source]