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Basal Zone (basal + zone)
Selected AbstractsEffects of the Little Ice Age on avalanche boulder tongues in the French Alps (Massif des Ecrins)EARTH SURFACE PROCESSES AND LANDFORMS, Issue 5 2004Vincent Jomelli Abstract Lichens of the subspecies Rhizocarpon geographicum s.l were measured on 25 avalanche boulder tongues in the Massif des Ecrins to elucidate the Little Ice Age history of avalanche activity. Results show: (1) an increase of lichen size from the median to the distal zone of deposits, and a decrease from the edges to the centre; (2) three types of lichen settlement. From the uppermost to the median zone, lichens are absent, because avalanche activity is very active. Down-slope, lichens occur in two different zones: the median zone is colonized by 5,20 mm size lichens on sides of blocks protected from the abrasional action of avalanches, while in the distal zone lichen diameters are largest (>30 mm) and occur on all sides of the blocks. The spatial distribution of the lichens and their size according to elevation make it possible to distinguish different phases during which avalanche activity has increased. At high elevation, the avalanche activity was at a maximum before ad 1650 and between ad 1730 and 1830. During these two periods avalanches had suf,cient magnitude to reach the basal zone of the deposits. At low elevation since ad 1650 the magnitude and frequency of avalanches have declined. Copyright © 2004 John Wiley & Sons, Ltd. [source] Solifluction processes on permafrost and non-permafrost slopes: results of a large-scale laboratory simulationPERMAFROST AND PERIGLACIAL PROCESSES, Issue 4 2008Charles Harris Abstract We present results of full-scale physical modelling of solifluction in two thermally defined environments: (a) seasonal frost penetration but no permafrost, and (b) a seasonally thawed active layer above cold permafrost. Modelling was undertaken at the Laboratoire M2C, Université de Caen-Basse Normandie, Centre National de la Recherche Scientifique, France. Two geometrically similar slope models were constructed using natural frost-susceptible test soil. In Model 1 water was supplied via a basal sand layer during freezing. In Model 2 the basal sand layer contained refrigerated copper tubing that maintained a permafrost table. Soil freezing was from the top down in Model 1 (one-sided freezing) but from the top down and bottom up (two-sided freezing) in Model 2. Thawing occurred from the top down as a result of positive air temperatures. Ice segregation in Model 1 decreased with depth, but in Model 2, simulated rainfall led to summer frost heave associated with ice segregation at the permafrost table, and subsequent two-sided freezing increased basal ice contents further. Thaw consolidation in Model 1 decreased with depth, but in Model 2 was greatest in the ice-rich basal layer. Soil shear strain occurred during thaw consolidation and was accompanied by raised pore water pressures. Displacement profiles showed decreasing movement rates with depth in Model 1 (one-sided freezing) but ,plug-like' displacements of the active layer over a shearing basal zone in Model 2 (two-sided active layer freezing). Volumetric transport rates were approximately 2.8 times higher for a given rate of surface movement in the permafrost model compared with the non-permafrost model. Copyright © 2008 John Wiley & Sons, Ltd. [source] Embryo development of Corticium candelabrum (Demospongiae: Homosclerophorida)INVERTEBRATE BIOLOGY, Issue 3 2007Sonia De Caralt Abstract. Corticium candelabrum is a homosclerophorid sponge widespread along the rocky Mediterranean sublittoral. Scanning and transmission electron microscopy were used to describe the gametes and larval development. The species is hermaphroditic. Oocytes and spermatocytes are clearly differentiated in April. Embryos develop from June to July when the larvae are released spontaneously. Spermatic cysts originate from choanocyte chambers and spermatogonia from choanocytes by choanocyte mitosis. Oocytes have a nucleolate nucleus and a cytoplasm filled with yolk granules and some lipids. Embryos are surrounded by firmly interlaced follicular cells from the parental tissue. A thin collagen layer lies below the follicular cells. The blastocoel is formed by migration of blastomeres to the morula periphery. Collagen is spread through the whole blastocoel in the embryo, but is organized in a dense layer (basal lamina) separating cells from the blastocoel in the larva. The larva is a typical cinctoblastula. The pseudostratified larval epithelium is formed by ciliated cells. The basal zone of the ciliated cells contains lipid inclusions and some yolk granules; the intermediate zone is occupied by the nucleus; and the apical zone contains abundant electron-lucent vesicles and gives rise to cilia with a single cross-striated rootlet. Numerous paracrystalline structures are contained in vacuoles within both apical and basal zones of the ciliated cells. Several slightly differentiated cell types are present in different parts of the larva. Most cells are ciliated, and show ultrastructural particularities depending on their location in the larvae (antero-lateral, intermediate, and posterior regions). A few smaller cells are non-ciliated. Several features of the C. candelabrum larva seem to support the previously proposed paraphyletic position of homoscleromorphs with respect to the other demosponges. [source] Long-distance transport of gases in plants: a perspective on internal aeration and radial oxygen loss from rootsPLANT CELL & ENVIRONMENT, Issue 1 2003T. D. COLMER ABSTRACT Internal transport of gases is crucial for vascular plants inhabiting aquatic, wetland or flood-prone environments. Diffusivity of gases in water is approximately 10 000 times slower than in air; thus direct exchange of gases between submerged tissues and the environment is strongly impeded. Aerenchyma provides a low-resistance internal pathway for gas transport between shoot and root extremities. By this pathway, O2 is supplied to the roots and rhizosphere, while CO2, ethylene, and methane move from the soil to the shoots and atmosphere. Diffusion is the mechanism by which gases move within roots of all plant species, but significant pressurized through-flow occurs in stems and rhizomes of several emergent and floating-leaved wetland plants. Through-flows can raise O2 concentrations in the rhizomes close to ambient levels. In general, rates of flow are determined by plant characteristics such as capacity to generate positive pressures in shoot tissues, and resistance to flow in the aerenchyma, as well as environmental conditions affecting leaf-to-air gradients in humidity and temperature. O2 diffusion in roots is influenced by anatomical, morphological and physiological characteristics, and environmental conditions. Roots of many (but not all) wetland species contain large volumes of aerenchyma (e.g. root porosity can reach 55%), while a barrier impermeable to radial O2 loss (ROL) often occurs in basal zones. These traits act synergistically to enhance the amount of O2 diffusing to the root apex and enable the development of an aerobic rhizosphere around the root tip, which enhances root penetration into anaerobic substrates. The barrier to ROL in roots of some species is induced by growth in stagnant conditions, whereas it is constitutive in others. An inducible change in the resistance to O2 across the hypodermis/exodermis is hypothesized to be of adaptive significance to plants inhabiting transiently waterlogged soils. Knowledge on the anatomical basis of the barrier to ROL in various species is scant. Nevertheless, it has been suggested that the barrier may also impede influx of: (i) soil-derived gases, such as CO2, methane, and ethylene; (ii) potentially toxic substances (e.g. reduced metal ions) often present in waterlogged soils; and (iii) nutrients and water. Lateral roots, that remain permeable to O2, may be the main surface for exchange of substances between the roots and rhizosphere in wetland species. Further work is required to determine whether diversity in structure and function in roots of wetland species can be related to various niche habitats. [source] | ||||||||||