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Soil Depth (soil + depth)
Kinds of Soil Depth Selected AbstractsThe influence of groundwater on surface flow erosion processes during a rainstormEARTH SURFACE PROCESSES AND LANDFORMS, Issue 5 2002D. L. RockwellArticle first published online: 27 MAY 200 Abstract Surface erosion rates on a disturbed natural soil in a 10 m indoor flume increased by an order of magnitude when a water table developed at a 10 cm depth during simulated rainstorms. Erosion rate increases did not correlate well with surface hydraulic flow conditions, and all significant erosion increases began before the full soil depth was saturated, before the water table reached the soil surface, and before seepage was possible. Groundwater influenced erosion processes primarily by increasing unsaturated pore-water pressures and decreasing soil shear strength in surface rainflow, rather than through the direct entrainment of soil particles by seepage flow. There was no unique morphologic expression of the influence of groundwater during a rainstorm. Subsurface processes influencing surface erosion were detected only by appropriate subsurface instrumentation, which included micropiezometers, tensiometers and time domain reflectometry. Erosion rate increases occurred all along the slope, and were not concentrated at the base of slope due to a seepage zone. Soil depth was crucial to determining surface erosion increase. It is likely that confusing trends in surface flow erosion rates in past studies have occurred due to unrecorded groundwater development or an emphasis on seepage effects. Groundwater must be monitored along hillslopes under all moisture and soil conditions in order to avoid misleading and inconsistent conclusions derived solely from surface flow or seepage data. Copyright © 2002 John Wiley & Sons, Ltd. [source] Temporal coherence of aboveground net primary productivity in mesic grasslandsECOGRAPHY, Issue 3 2008Jana L. Heisler Synchrony in ecological variables over wide geographic areas suggests that large-scale environmental factors drive the structure and function of ecosystems and override more local-scale environmental variation. Described also as coherence, this phenomenon has been documented broadly in the ecological literature and has recently received increasing attention as scientists attempt to quantify the impacts of global changes on organisms and their habitats. Using a mesic grassland site in North America, we assessed coherence in ecosystem function by quantifying similarity in aboveground net primary production (ANPP) dynamics in 48 permanent sampling locations (PSLs) over a 16-yr period. Our primary objective was to characterize coherence across a broad geographic region (with similar ecosystem structure and function), and we hypothesized that precipitation and a similar fire frequency would strengthen coherence between PSLs. All 48 PSLs at our site (Konza Prairie Biological Station, Manhattan, KS, USA; KPBS) were exposed to a similar regional driver of ANPP (precipitation); however, local drivers (including differences in fire frequency and soil depth at different topographic positions) varied strongly among individual PSLs. For the purpose of this assessment, the watershed-level experimental design of KPBS was considered a model, which represented different fire management strategies across the Great Plains Region. Our analyses revealed a site-level (KPBS) coherence in ANPP dynamics of 0.53 for the period of 1984,1999. Annual fire enhanced coherence among PSLs to 0.76, whereas less frequent fire (fire exclusion or a 4-yr fire return interval) failed to further increase coherence beyond that of the KPBS site level. Soil depth also strongly influenced coherence among PSLs with shallow soils at upland sites showing strong coherence across fire regimes and annually burned uplands closely linked to annual precipitation dynamics. The lack of coherence in ecosystem function in PSLs with deep soils and low fire frequencies suggests that conservation and management efforts will need to be more location specific in such areas where biotic interactions may be more important than regional abiotic drivers. [source] Links between methane flux and transcriptional activities of methanogens and methane oxidizers in a blanket peat bogFEMS MICROBIOLOGY ECOLOGY, Issue 1 2010Thomas E. Freitag Abstract The relationship between biogeochemical process rates and microbial functional activity was investigated by analysis of the transcriptional dynamics of the key functional genes for methanogenesis (methyl coenzyme M reductase; mcrA) and methane oxidation (particulate methane monooxygenase; pmoA) and in situ methane flux at two peat soil field sites with contrasting net methane-emitting and -oxidizing characteristics. qPCR was used to quantify the abundances of mcrA and pmoA genes and transcripts at two soil depths. Total methanogen and methanotroph transcriptional dynamics, calculated from mcrA and pmoA gene : transcript abundance ratios, were similar at both sites and depths. However, a linear relationship was demonstrated between surface mcrA and pmoA transcript dynamics and surface flux rates at the methane-emitting and methane-oxidizing sites, respectively. Results indicate that methanotroph activity was at least partially substrate-limited at the methane-emitting site and by other factors at the methane-oxidizing site. Soil depth also contributed to the control of surface methane fluxes, but to a lesser extent. Small differences in the soil water content may have contributed to differences in methanogen and methanotroph activities. This study therefore provides a first insight into the regulation of in situ, field-level surface CH4 flux at the molecular level by an accurate reflection of gene : transcript abundance ratios for the key genes in methane generation and consumption. [source] Rainfall variability and hydrological and erosive response of an olive tree microcatchment under no-tillage with a spontaneous grass cover in SpainEARTH SURFACE PROCESSES AND LANDFORMS, Issue 7 2010E. V. Taguas Abstract Most studies on runoff and soil loss from olive orchards were performed on plots, despite the fact that measurements that examine a range of erosive processes on different scales are essential to evaluate the suitability of the use and soil management of this type of land. The main environmental limitations of much of the land used for olive orchards in the Mediterranean are the steep slopes and the shallow soil depth , and this was the case in the study area. Soil erosion and runoff over two hydrological years (2005,2006 and 2006,2007) were monitored in an olive orchard microcatchment of 6·1,ha under no-tillage with spontaneous grass in order to evaluate its hydrological and erosive behaviour. Moreover, soil parameters such as organic matter (%OM), bulk density (BD) and hydraulic saturated conductivity (Ks) were also examined in the microcatchment to describe management effects on hydrological balance and on erosive processes. In the study period, the results showed runoff coefficients of 6·0% in the first year and 0·9% in the second. The differences respond to the impact of two or three yearly maximum events which were decisive in the annual balances. On the event scale, although maximum rainfall intensity values had a big influence on peak flows and runoff, its importance on mean sediment concentrations and sediment discharges was difficult to interpret due to the likely control of grass cover on volume runoff and on soil protection. In the case of annual soil erosion, they were measured as 1·0,Mg,ha,1,yr,1 and 0·3,Mg,ha,1,yr,1. Both are lower than the tolerance values evaluated in Andalusia (Spain). These results support the implementation of no-tillage with spontaneous grass cover for sloping land, although the reduced infiltration conditions determined by Ks in the first horizon suggest grass should be allowed to grow not only in spring but also in autumn. In addition, specific measurements to control gullies, which have formed in the terraced area in the catchment, should be included since it is expected that they could be the main sources of sediments. Copyright © 2010 John Wiley & Sons, Ltd. [source] A low-dimensional physically based model of hydrologic control of shallow landsliding on complex hillslopesEARTH SURFACE PROCESSES AND LANDFORMS, Issue 13 2008Ali Talebi Abstract Hillslopes have complex three-dimensional shapes that are characterized by their plan shape, profile curvature of surface and bedrock, and soil depth. To investigate the stability of complex hillslopes (with different slope curvatures and plan shapes), we combine the hillslope-storage Boussinesq (HSB) model with the infinite slope stability method. The HSB model is based on the continuity and Darcy equations expressed in terms of storage along the hillslope. Solutions of the HSB equation account explicitly for plan shape by introducing the hillslope width function and for profile curvature through the bedrock slope angle and the hillslope soil depth function. The presented model is composed of three parts: a topography model conceptualizing three-dimensional soil mantled landscapes, a dynamic hydrology model for shallow subsurface flow and water table depth (HSB model) and an infinite slope stability method based on the Mohr,Coulomb failure law. The resulting hillslope-storage Boussinesq stability model (HSB-SM) is able to simulate rain-induced shallow landsliding on hillslopes with non-constant bedrock slope and non-parallel plan shape. We apply the model to nine characteristic hillslope types with three different profile curvatures (concave, straight, convex) and three different plan shapes (convergent, parallel, divergent). In the presented model, the unsaturated storage has been calculated based on the unit head gradient assumption. To relax this assumption and to investigate the effect of neglecting the variations of unsaturated storage on the assessment of slope stability in the transient case, we also combine a coupled model of saturated and unsaturated storage and the infinite slope stability method. The results show that the variations of the unsaturated zone storage do not play a critical role in hillslope stability. Therefore, it can be concluded that the presented dynamic slope stability model (HSB-SM) can be used safely for slope stability analysis on complex hillslopes. Our results show that after a certain period of rainfall the convergent hillslopes with concave and straight profiles become unstable more quickly than others, whilst divergent convex hillslopes remain stable (even after intense rainfall). In addition, the relation between subsurface flow and hillslope stability has been investigated. Our analyses show that the minimum safety factor (FS) occurs when the rate of subsurface flow is a maximum. In fact, by increasing the subsurface flow, stability decreases for all hillslope shapes. Copyright © 2008 John Wiley & Sons, Ltd. [source] Soil production in heath and forest, Blue Mountains, Australia: influence of lithology and palaeoclimateEARTH SURFACE PROCESSES AND LANDFORMS, Issue 8 2005Marshall T. Wilkinson Abstract An Erratum has been published for this article in Earth Surfaces Processes and Landforms 25(13) 2005, 1683,1686. Recent determinations of soil production from in situ cosmogenic nuclides indicate that production decreases exponentially with soil depth. This contrasts with a long-held assumption that maximum soil production occurs under a soil cover of finite depth. Sites in the Blue Mountains, Australia, show a sharp decrease of soil depth where vegetation changes from forested plateau surfaces to heath-covered spurs, and bands of bare rock in the heath suggest that soil production depends on presence of a finite depth of soil. The substrate varies from hard ferruginized sandstone to soft saprolite. In situ 10Be determinations indicate that apparent rates of erosion and soil production are greater under the relatively thin heath soil than under the thicker forest soil but, in contrast to other studies, these sites do not show significant depth-dependence of apparent soil production. The pattern reflects both hardness variation in the rock substrate and the effect of Late Quaternary climatic change. Optically stimulated luminescence (OSL) dating indicates that soil ,30 cm depth is of Holocene age whereas the deeper soil is substantially older. The age-break coincides with a stone line interpreted as a former surface lag deposit. Assuming that pre-Holocene soil depths were 30 cm less than today, recalculated soil production tends to decrease with increasing depth. Soil production at this site requires soil cover but bare rock patches and vegetation comprise a shifting mosaic. In the long term, average rates of erosion and soil production decrease with increasing soil depth. Copyright © 2005 John Wiley & Sons, Ltd. [source] Testing a model for predicting the timing and location of shallow landslide initiation in soil-mantled landscapesEARTH SURFACE PROCESSES AND LANDFORMS, Issue 9 2003M. Casadei Abstract The growing availability of digital topographic data and the increased reliability of precipitation forecasts invite modelling efforts to predict the timing and location of shallow landslides in hilly and mountainous areas in order to reduce risk to an ever-expanding human population. Here, we exploit a rare data set to develop and test such a model. In a 1·7 km2 catchment a near-annual aerial photographic coverage records just three single storm events over a 45 year period that produced multiple landslides. Such data enable us to test model performance by running the entire rainfall time series and determine whether just those three storms are correctly detected. To do this, we link a dynamic and spatially distributed shallow subsurface runoff model (similar to TOPMODEL) to an in,nite slope model to predict the spatial distribution of shallow landsliding. The spatial distribution of soil depth, a strong control on local landsliding, is predicted from a process-based model. Because of its common availability, daily rainfall data were used to drive the model. Topographic data were derived from digitized 1 : 24 000 US Geological Survey contour maps. Analysis of the landslides shows that 97 occurred in 1955, 37 in 1982 and ,ve in 1998, although the heaviest rainfall was in 1982. Furthermore, intensity,duration analysis of available daily and hourly rainfall from the closest raingauges does not discriminate those three storms from others that did not generate failures. We explore the question of whether a mechanistic modelling approach is better able to identify landslide-producing storms. Landslide and soil production parameters were ,xed from studies elsewhere. Four hydrologic parameters characterizing the saturated hydraulic conductivity of the soil and underlying bedrock and its decline with depth were ,rst calibrated on the 1955 landslide record. Success was characterized as the most number of actual landslides predicted with the least amount of total area predicted to be unstable. Because landslide area was consistently overpredicted, a threshold catchment area of predicted slope instability was used to de,ne whether a rainstorm was a signi,cant landslide producer. Many combinations of the four hydrological parameters performed equally well for the 1955 event, but only one combination successfully identi,ed the 1982 storm as the only landslide-producing storm during the period 1980,86. Application of this parameter combination to the entire 45 year record successfully identi,ed the three events, but also predicted that two other landslide-producing events should have occurred. This performance is signi,cantly better than the empirical intensity,duration threshold approach, but requires considerable calibration effort. Overprediction of instability, both for storms that produced landslides and for non-producing storms, appears to arise from at least four causes: (1) coarse rainfall data time scale and inability to document short rainfall bursts and predict pressure wave response; (2) absence of local rainfall data; (3) legacy effect of previous landslides; and (4) inaccurate topographic and soil property data. Greater resolution of spatial and rainfall data, as well as topographic data, coupled with systematic documentation of landslides to create time series to test models, should lead to signi,cant improvements in shallow landslides forecasting. Copyright © 2003 John Wiley & Sons, Ltd. [source] The influence of groundwater on surface flow erosion processes during a rainstormEARTH SURFACE PROCESSES AND LANDFORMS, Issue 5 2002D. L. RockwellArticle first published online: 27 MAY 200 Abstract Surface erosion rates on a disturbed natural soil in a 10 m indoor flume increased by an order of magnitude when a water table developed at a 10 cm depth during simulated rainstorms. Erosion rate increases did not correlate well with surface hydraulic flow conditions, and all significant erosion increases began before the full soil depth was saturated, before the water table reached the soil surface, and before seepage was possible. Groundwater influenced erosion processes primarily by increasing unsaturated pore-water pressures and decreasing soil shear strength in surface rainflow, rather than through the direct entrainment of soil particles by seepage flow. There was no unique morphologic expression of the influence of groundwater during a rainstorm. Subsurface processes influencing surface erosion were detected only by appropriate subsurface instrumentation, which included micropiezometers, tensiometers and time domain reflectometry. Erosion rate increases occurred all along the slope, and were not concentrated at the base of slope due to a seepage zone. Soil depth was crucial to determining surface erosion increase. It is likely that confusing trends in surface flow erosion rates in past studies have occurred due to unrecorded groundwater development or an emphasis on seepage effects. Groundwater must be monitored along hillslopes under all moisture and soil conditions in order to avoid misleading and inconsistent conclusions derived solely from surface flow or seepage data. Copyright © 2002 John Wiley & Sons, Ltd. [source] Temporal coherence of aboveground net primary productivity in mesic grasslandsECOGRAPHY, Issue 3 2008Jana L. Heisler Synchrony in ecological variables over wide geographic areas suggests that large-scale environmental factors drive the structure and function of ecosystems and override more local-scale environmental variation. Described also as coherence, this phenomenon has been documented broadly in the ecological literature and has recently received increasing attention as scientists attempt to quantify the impacts of global changes on organisms and their habitats. Using a mesic grassland site in North America, we assessed coherence in ecosystem function by quantifying similarity in aboveground net primary production (ANPP) dynamics in 48 permanent sampling locations (PSLs) over a 16-yr period. Our primary objective was to characterize coherence across a broad geographic region (with similar ecosystem structure and function), and we hypothesized that precipitation and a similar fire frequency would strengthen coherence between PSLs. All 48 PSLs at our site (Konza Prairie Biological Station, Manhattan, KS, USA; KPBS) were exposed to a similar regional driver of ANPP (precipitation); however, local drivers (including differences in fire frequency and soil depth at different topographic positions) varied strongly among individual PSLs. For the purpose of this assessment, the watershed-level experimental design of KPBS was considered a model, which represented different fire management strategies across the Great Plains Region. Our analyses revealed a site-level (KPBS) coherence in ANPP dynamics of 0.53 for the period of 1984,1999. Annual fire enhanced coherence among PSLs to 0.76, whereas less frequent fire (fire exclusion or a 4-yr fire return interval) failed to further increase coherence beyond that of the KPBS site level. Soil depth also strongly influenced coherence among PSLs with shallow soils at upland sites showing strong coherence across fire regimes and annually burned uplands closely linked to annual precipitation dynamics. The lack of coherence in ecosystem function in PSLs with deep soils and low fire frequencies suggests that conservation and management efforts will need to be more location specific in such areas where biotic interactions may be more important than regional abiotic drivers. [source] Relationships between spatial environmental heterogeneity and plant species diversity on a limestone pavementECOGRAPHY, Issue 6 2003Jeremy T. Lundholm No empirical studies have examined the relationship between diversity and spatial heterogeneity across unimodal species richness gradients. We determined the relationships between diversity and environmental factors for 144 0.18 m2 plots in a limestone pavement alvar in southern Ontario, Canada, including within-plot spatial heterogeneity in soil depth, microtopography and microsite composition. Species richness was unimodally related to mean soil depth and relative elevation. Microsite heterogeneity and soil depth heterogeneity were positively correlated with species richness, and the richness peaks of the unimodal gradients correspond to the maximally spatially heterogeneous plots. The best predictive models of species richness and evenness, however, showed that other factors, such as ramet density and flooding, are the major determinants of diversity in this system. The findings that soil depth heterogeneity had effects on diversity when the effects of mean soil depth were factored out, and that unimodal richness peaks were associated with high spatial heterogeneity in environmental factors represent significant contributions to our understanding of how spatial heterogeneity might contribute to diversity maintenance in plant communities. [source] Mechanistic assessment of hillslope transpiration controls of diel subsurface flow: a steady-state irrigation approachECOHYDROLOGY, Issue 2 2010H. R. Barnard Abstract Mechanistic assessment of how transpiration influences subsurface flow is necessary to advance understanding of catchment hydrology. We conducted a 24-day, steady-state irrigation experiment to quantify the relationships among soil moisture, transpiration and hillslope subsurface flow. Our objectives were to: (1) examine the time lag between maximum transpiration and minimum hillslope discharge with regard to soil moisture; (2) quantify the relationship between diel hillslope discharge and daily transpiration; and (3) identify the soil depth from which trees extract water for transpiration. An 8 × 20 m hillslope was irrigated at a rate of 3·6 mm h,1. Diel fluctuations in hillslope discharge persisted throughout the experiment. Pre-irrigation time lags between maximum transpiration and minimum hillslope discharge were 6·5 h, whereas lags during steady-state and post-irrigation conditions were 4 and 2 h, respectively. The greatest correlation between transpiration and hillslope discharge occurred during the post-irrigation period, when the diel reduction in hillslope discharge totalled 90% of total measured daily transpiration. Daily transpiration of trees within the irrigated area remained relatively constant throughout the experiment. Diel fluctuations in soil moisture were greatest at a depth of 0·9,1·2 m prior to irrigation and became more uniform throughout the soil profile during and post-irrigation. This study clearly demonstrates that when soil moisture is high, hillslope trees can be an important factor in diel fluctuations in stream discharge. We advance a conceptual model for the site whereby the relationship between transpiration and hillslope discharge is a function of soil moisture status and drainable porosity. Copyright © 2010 John Wiley & Sons, Ltd. [source] Woody plants modulate the temporal dynamics of soil moisture in a semi-arid mesquite savanna,ECOHYDROLOGY, Issue 1 2010Daniel L. Potts Abstract Climate variability and human activities interact to increase the abundance of woody plants in arid and semi-arid ecosystems worldwide. How woody plants interact with rainfall to influence patterns of soil moisture through time, at different depths in the soil profile and between neighboring landscape patches is poorly known. In a semi-arid mesquite savanna, we deployed a paired array of sensors beneath a mesquite canopy and in an adjacent open area to measure volumetric soil water content (,) every 30 min at several depths between 2004 and 2007. In addition, to quantify temporally dynamic variation in soil moisture between the two microsites and across soil depths we analysed , time-series using fast Fourier transforms (FFT). FFT analyses were consistent with the prediction that by reducing evaporative losses through shade and reducing rainfall inputs through canopy interception of small rainfall events, the mesquite canopy was associated with a decline in high-frequency (hour-to-hour and day-to-day) variation in shallow ,. Finally, we found that, in both microsites, high-frequency , variation declined with increasing soil depth as the influence of evaporative losses and inputs associated with smaller rainfall events declined. In this case, we argue that the buffering of shallow soil moisture against high-frequency variations can enhance nutrient cycling and alter the carbon cycle in dryland ecosystems. Copyright © 2009 John Wiley & Sons, Ltd. [source] Arsenic, lead, and other trace elements in soils contaminated with pesticide residues at the Hanford site (USA)ENVIRONMENTAL TOXICOLOGY, Issue 2 2003Jerry Yokel Abstract The primary purpose of this study was to characterize arsenic (As) and lead (Pb) concentrations in former orchard soils contaminated with lead arsenate pesticides at the Hanford site in Washington state (USA). Surface samples (n = 31) were collected from former orchard soils (in cultivation during the pre-Hanford period) at five locations at the 100 Areas and at one location at the Old Hanford Townsite (OHT). Another set of samples (n = 17) was collected over a soil depth interval of 10,50 cm at the four locations with the highest As and Pb surface concentrations. All samples were analyzed for 22 trace elements (including As and Pb) with inductively coupled plasma,atomic emission spectrometry (ICP,AES). The mean, standard deviation, and range for As in the surface soils were 30, 61, and 2.9,270 mg/kg dry wt, respectively. The corresponding statistics for Pb were 220, 460, and 6.5,1900 mg/kg dry wt, respectively. As and Pb concentrations in the surface soils were positively and significantly correlated (r = 0.91, Bonferroni p < 0.05). Descriptive statistics and bivariate correlations were also computed for other trace elements. As and Pb mean concentrations in the surface soils each differed significantly (p < 0.05) among Hanford locations, with the highest concentrations at the 100-H and 100-F Areas. Although both As and Pb mean concentrations decreased with soil depth, regression and correlation coefficients only, for Pb significantly differed from zero (b = ,0.0372, r = ,0.805, Bonferroni p < 0.05). Compared with data in the literature As and Pb concentrations found in this study exceeded background levels but were typical of orchard soils. Furthermore, mean As and Pb soil concentrations were in the range of various toxicological benchmarks derived for protection of human and ecological receptors. © 2003 Wiley Periodicals, Inc. Environ Toxicol 18: 104,114, 2003 [source] Presence and distribution of wastewater-derived pharmaceuticals in soil irrigated with reclaimed waterENVIRONMENTAL TOXICOLOGY & CHEMISTRY, Issue 2 2006Chad A. Kinney Abstract Three sites in the Front Range of Colorado, USA, were monitored from May through September 2003 to assess the presence and distribution of pharmaceuticals in soil irrigated with reclaimed water derived from urban wastewater. Soil cores were collected monthly, and 19 pharmaceuticals, all of which were detected during the present study, were measured in 5-cm increments of the 30-cm cores. Samples of reclaimed water were analyzed three times during the study to assess the input of pharmaceuticals. Samples collected before the onset of irrigation in 2003 contained numerous pharmaceuticals, likely resulting from the previous year's irrigation. Several of the selected pharmaceuticals increased in total soil concentration at one or more of the sites. The four most commonly detected pharmaceuticals were erythromycin, carbamazepine, fluoxetine, and diphenhydramine. Typical concentrations of the individual pharmaceuticals observed were low (0.02,15 ,g/kg dry soil). The existence of subsurface maximum concentrations and detectable concentrations at the lowest sampled soil depth might indicate interactions of soil components with pharmaceuticals during leaching through the vadose zone. Nevertheless, the present study demonstrates that reclaimed-water irrigation results in soil pharmaceutical concentrations that vary through the irrigation season and that some compounds persist for months after irrigation. [source] Quantifying dye tracers in soil profiles by image processingEUROPEAN JOURNAL OF SOIL SCIENCE, Issue 2 2000I. Forrer Summary Developing and testing models for solute transport in the field requires experimental data on the spreading of solutes in the soil. Obtaining such data is costly, and a substantial part of the total costs is in the preparation and chemical analysis of the tracing compounds in the gathered samples. We developed a cheap method to quantify the concentration of the mobile dye tracer Brilliant Blue FCF from digitized photographs of stained soil profiles, and we have tested it in the field. Soil sampling and chemical analyses were necessary only to establish a calibration relation between the dye content and the colour of the soil. The digital images were corrected for geometrical distortions, varying background brightness, and colour tinges, and then they were analysed to determine the soil colour at sampling points in the profiles. The resident concentration of the dye was modelled by polynomial regression with the primary colours red, green, blue and the soil depth as explanatory variables. Concentration maps of Brilliant Blue were then computed from the digitized images with a spatial resolution of 1 mm. Validation of the technique with independent data showed that the method predicted the concentration of the dye well, provided the corrected images contained only the colours included in the calibration. [source] Carbon monoxide uptake kinetics in unamended and long-term nitrogen-amended temperate forest soilsFEMS MICROBIOLOGY ECOLOGY, Issue 3 2006Alvarus S. K. Chan Abstract The effect of nitrogen (N) additions on the dynamics of carbon monoxide consumption in temperate forest soils is poorly understood. We measured soil CO profiles, potential rates of CO consumption and uptake kinetics in temperate hardwood and pine control plots and plots amended with 50 and 150 kg N ha,1 year,1 for more than 15 years. Soil profiles of CO concentrations were above atmospheric levels in the high-N plots of both stands, suggesting that in these forest soils the balance between consumption and production may be shifted so that either production is increased or consumption decreased. Highest rates of CO consumption were measured in the organic horizon and decreased with soil depth. In the N-amended plots, CO consumption increased in all but one soil depth of the hardwood stand, but decreased in all soil depths of the pine stand. CO enzyme affinities increased with soil depth in the control plots. However, enzyme affinities in the most active soil depths (organic and 0,5 cm mineral) decreased in response to low levels of N in both stands. In the high-N plots, affinities dramatically-increased in the hardwood stand, but decreased in the organic horizon and increased slightly in the 0,5 cm mineral soil in the pine stand. These findings indicate that long-term N addition either by fertilization or deposition may alter the size, composition and/or physiology of the community of CO consumers so that their ability to act as a sink for atmospheric CO has changed. This change could have a substantial effect on the lifetime of greenhouse gases such as CH4 and therefore the future of Earth's climate. [source] Soil organic carbon contents in long-term experimental grassland plots in the UK (Palace Leas and Park Grass) have not changed consistently in recent decadesGLOBAL CHANGE BIOLOGY, Issue 7 2009D. W. HOPKINS Abstract A recent report of widespread declines in soil organic C (SOC) in the UK over the 10,25 years until the early 2000s has focussed attention on the importance of resampling previously characterized sites to assess long-term trends in SOC contents and the importance of soils as a potentially volatile and globally significant reservoir of terrestrial C. We have used two sets of long-term experimental plots which have been under constant and known management for over a century and for which historical data exist that allow comparison over recent decades to determine what, if any, changes in SOC content have occurred. The plots used are the Palace Leas (PL) Meadow Hay Plots in north-east England (UK) established in 1897, and from the Park Grass (PG) Continuous Hay experiment established in 1856 at Rothamsted in south-east England. Collectively, these plots represent the only grassland sites in the UK under long-term management where changes in SOC over several decades can be assessed, and are probably unique in the world. The plots have received different manure and fertilizer treatment and have been under known management for at least 100 years. In 1982, total SOC contents were determined for the 0,27 cm layer of six of the PL plots using measurements of SOC concentrations, bulk density and soil depth. In 2006, the same six PL plots were resampled and SOC contents determined again. Four of the plots showed no net change in SOC content, but two plots showed net loss of SOC of 15% and 17% (amounting to decreases of 18 and 15 t C ha,1) since 1982. However, these differences in total SOC content were in a similar range to the variations in bulk density (6,31%) with changing soil water content. In 1959, the soil masses and SOC concentrations to 23 cm depth were measured on six PG plots with fertilizer and manure treatments corresponding closely with those measured on PL. In 2002, the SOC concentrations on the same plots were measured again. On three of the PG plots, SOC concentrations had declined by 2,10%, but in the other three it had increased by 4,8% between 1959 and 2002. If it is assumed that the soil bulk density had not changed over this period, the losses of SOC from the top soils ranged range from 10 to 3 t C ha,1, while the gains ranged from 4 to 7 t C ha,1. When the differences with time in SOC contents for the six PL and the six PG plots were examined using paired t -tests, that is, regarding the plots as two sets of six replicate permanent grasslands, there were no significant differences between 1982 and 2006 for the PL plots or between 1959 and 2002 for the PG plots. Thus, these independent observations on similar plots at PL and PG indicate there has been no consistent decrease in SOC stocks in surface soils under old, permanent grassland in England in recent decades, even though meteorological records for both sites indicate significant warming of the soil and air between 1980 and 2000. Because the potential influences of changes in management or land use have been definitively excluded, and measured rather than derived bulk densities have been used to convert from SOC concentrations to SOC amounts, our observations question whether for permanent grassland in England, losses in SOC in recent decades reported elsewhere can be attributed to widespread environmental change. [source] Conversion of hardwood forests to spruce and pine plantations strongly reduced soil methane sink in GermanyGLOBAL CHANGE BIOLOGY, Issue 6 2003WERNER BORKEN Abstract Well-drained forest soils are thought to be a significant sink for atmospheric methane. Recent research suggests that land use change reduces the soil methane sink by diminishing populations of methane oxidizing bacteria. Here we report soil CH4 uptake from ,natural' mature beech forests and from mature pine and spruce plantations in two study areas of Germany with distinct climate and soils. The CH4 uptake rates of both beech forests at Solling and Unterlüß were about two,three times the CH4 uptake rates of the adjacent pine and spruce plantations, indicating a strong impact of forest type on the soil CH4 sink. The CH4 uptake rates of sieved mineral soils from our study sites confirmed the tree species effect and indicate that methanotrophs were mainly reduced in the 0,5 cm mineral soil depth. The reasons for the reduction are still unknown. We found no site effect between Solling and Unterlüß, however, CH4 uptake rates from Solling were significantly higher at the same effective CH4 diffusivity. This potential site effect was masked by higher soil water contents at Solling. Soil pH (H2O) explained 71% of the variation in CH4 uptake rates of sieved mineral soils from the 0,5 cm depth, while cation exchange capacity, soil organic carbon, soil nitrogen and total phosphorous content were not correlated with CH4 uptake rates. Comparing 1998,99, annual CH4 uptake rates increased by 69,111% in the beech and spruce stands and by 5,25% in the pine stands, due primarily to differences in growing season soil moisture. Cumulative CH4 uptake rates from November throughout April were rather constant in both years. The CH4 uptake rates of each stand were separately predicted using daily average soil matric potential and a previously developed empirical model. The model results revealed that soil matric potential explains 53,87% of the temporal variation in CH4 uptake. The differences between measured and predicted annual CH4 uptake rates were less than 10%, except for the spruce stand at Solling in 1998 (17%). Based on data from this study and from the literature, we calculated a total reduction in the soil CH4 sink of 31% for German forests due in part to conversion of deciduous to coniferous forests. [source] Predicting and quantifying the structure of tropical dry forests in South Florida and the Neotropics using spaceborne imageryGLOBAL ECOLOGY, Issue 3 2006Thomas W. Gillespie ABSTRACT Aim, This research examines environmental theories and remote sensing methods that have been hypothesized to be associated with tropical dry forest structure. Location, Tropical dry forests of South Florida and the Neotropics. Methods, Field measurements of stand density, basal area and tree height were collected from 22 stands in South Florida and 30 stands in the Neotropics. In South Florida, field measurements were compared to climatic (temperature, precipitation, hurricane disturbance) and edaphic (rockiness, soil depth) variables, spectral indices (NDVI, IRI, MIRI) from Landsat 7 ETM+, and estimates of tree height from the Shuttle Radar Topography Mission (SRTM) and the National Elevation Dataset (NED). Environmental variables associated with tropical dry forest structure in South Florida were compared to tropical dry forest in other Neotropical sites. Results, There were significant correlations among temperature and precipitation, and stand density and tree height in South Florida. There were significant correlations between (i) stand density and mean NDVI and standard deviation of NDVI, (ii) MIRI and stand density, basal area and mean tree height, and (iii) estimates of tree height from SRTM with maximum tree height. In the Neotropics, there were no relationships between temperature or precipitation and tropical dry forest structure, however, Neotropical sites that experience hurricane disturbance had significantly shorter tree heights and higher stand densities. Main conclusions, It is possible to predict and quantify the forest structure characteristics of tropical dry forests using climatic data, Landsat 7 ETM+ imagery and SRTM data in South Florida. However, results based on climatic data are region-specific and not necessarily transferable between tropical dry forests at a continental spatial scale. Spectral indices from Landsat 7 ETM+ can be used to quantify forest structure characteristics, but SRTM data are currently not transferable to other regions. Hurricane disturbance has a significant impact on forest structure in the Neotropics. [source] Evaluating interactions between soil drainage and seedling performance in a restoration of Pinus sylvestris woodland, ScotlandGLOBAL ECOLOGY, Issue 2 2001M. D. Crowell Abstract 1,This paper evaluates the role of soil drainage in tree seedling performance at a site being restored from Calluna vulgaris moorland to Pinus sylvestris woodland, in Glen Affric, Scotland. The investigation focuses on the relationships between height of planted seedlings, type of ground vegetation and drainage conditions. 2,Slope, aspect, and soil depth were assessed as potential surrogates for direct measures of soil drainage, all of which were derived from digital terrain data. 3,Six variables related to drainage were recorded at 58 seedling locations and used in a factor analysis to understand links between soil moisture conditions, topographic variables and soil depth characteristics. 4,Factor analysis generated two factors that accounted for 70.5% of the variance in the correlation matrix of these variables: Factor 1 correlated strongly with variables that controlled peat accumulation and Factor 2 correlated strongly with topographic controls upon drainage patterns. 5,These two factors explained a significant amount of the variance in height of the Pinus seedlings planted at these locations. Significant differences were found between the factor scores associated with different types of ground vegetation, as well as between the seedling heights observed at locations with different vegetation types. 6,Multiple regressions were developed that indicated that slope, aspect, and soil depth were significant as independent variables in models where soil moisture content and aerobic soil depth were the dependent variables. [source] Root and rhizome systems of perennial grasses grown in Inner Mongolian grassland, ChinaGRASSLAND SCIENCE, Issue 4 2009Min Ao Abstract The root and rhizome systems of dominant perennial grasses in Inner Mongolian grassland were clarified. We surveyed the vertical distribution of root and rhizome biomass in the natural stands, and the changes of under-ground biomass and the branching pattern of rhizomes for transplanted plants in a container experiment. Most roots of Leymus chinensis, Bromus inermis, Elymus dahuricus and Agropyron cristatum were distributed in the soil depth of 0,10 cm. Roots of E. dahuricus and A. cristatum were distributed in a shallower soil layer, but those of L. chinensis and B. inermis were distributed in a deeper soil layer. Biomass of above-ground parts increased with growth, resulting in a decreasing ratio of under-ground parts to total biomass. Rhizomes of L. chinensis and B. inermis were distributed in the soil depth of 0,10 cm, but E. dahuricus and A. cristatum did not have rhizomes. L. chinensis had longer rhizomes and new ramets were produced away from their mother plant. B. inermis had many short rhizomes and produced daughter plants near their mother plant. [source] Power function decay of hydraulic conductivity for a TOPMODEL-based infiltration routineHYDROLOGICAL PROCESSES, Issue 18 2006Jun Wang Abstract TOPMODEL rainfall-runoff hydrologic concepts are based on soil saturation processes, where soil controls on hydrograph recession have been represented by linear, exponential, and power function decay with soil depth. Although these decay formulations have been incorporated into baseflow decay and topographic index computations, only the linear and exponential forms have been incorporated into infiltration subroutines. This study develops a power function formulation of the Green and Ampt infiltration equation for the case where the power n = 1 and 2. This new function was created to represent field measurements in the New York City, USA, Ward Pound Ridge drinking water supply area, and provide support for similar sites reported by other researchers. Derivation of the power-function-based Green and Ampt model begins with the Green and Ampt formulation used by Beven in deriving an exponential decay model. Differences between the linear, exponential, and power function infiltration scenarios are sensitive to the relative difference between rainfall rates and hydraulic conductivity. Using a low-frequency 30 min design storm with 4·8 cm h,1 rain, the n = 2 power function formulation allows for a faster decay of infiltration and more rapid generation of runoff. Infiltration excess runoff is rare in most forested watersheds, and advantages of the power function infiltration routine may primarily include replication of field-observed processes in urbanized areas and numerical consistency with power function decay of baseflow and topographic index distributions. Equation development is presented within a TOPMODEL-based Ward Pound Ridge rainfall-runoff simulation. Copyright © 2006 John Wiley & Sons, Ltd. [source] Soil moisture dynamics in an eastern Amazonian tropical forestHYDROLOGICAL PROCESSES, Issue 12 2006Rogério D. Bruno Abstract We used frequency-domain reflectometry to make continuous, high-resolution measurements for 22 months of the soil moisture to a depth of 10 m in an Amazonian rain forest. We then used these data to determine how soil moisture varies on diel, seasonal and multi-year timescales, and to better understand the quantitative and mechanistic relationships between soil moisture and forest evapotranspiration. The mean annual precipitation at the site was over 1900 mm. The field capacity was approximately 0·53 m3 m,3 and was nearly uniform with soil depth. Soil moisture decreased at all levels during the dry season, with the minimum of 0·38 m3 m,3 at 3 m beneath the surface. The moisture in the upper 1 m showed a strong diel cycle with daytime depletion due to evapotranspiration. The moisture beneath 1 m declined during both day and night due to the combined effects of evapotranspiration, drainage and a nighttime upward movement of water. The depth of active water withdrawal changed markedly over the year. The upper 2 m of soil supplied ,56% of the water used for evapotranspiration in the wet season and ,28% of the water used in the dry season. The zone of active water withdrawal extended to a depth of at least 10 m. The day-to-day rates of moisture withdrawal from the upper 10 m of soil during rain-free periods agreed well with simultaneous measurements of whole-forest evapotranspiration made by the eddy covariance technique. The forest at the site was well adapted to the normal cycle of wet and dry seasons, and the dry season had only a small effect on the rates of land,atmosphere water vapour exchange. Copyright © 2006 John Wiley & Sons, Ltd. [source] Soil water content and yield variability in vineyards of Mediterranean northeastern Spain affected by mechanization and climate variabilityHYDROLOGICAL PROCESSES, Issue 11 2006M. C. Ramos Abstract The objective of this paper was to analyse the combined influence of the Mediterranean climate variability (particularly the irregular rainfall distribution throughout the year) and the land transformations carried out in vineyards of northeastern Spain on soil water content evolution and its influence on grape production. The study was carried out in a commercial vineyard located in the Anoia,Alt Penedès region (Barcelona province, northeastern Spain), which was prepared for mechanization with important land transformations. Two plots were selected for the study: one with low degree of transformation of the soil profile, representing a non-disturbed situation, and the second one in which more than 3 m were cut in the upper part of the plot and filled in the lower part, representing the disturbed situation. Soil water content was evaluated at three positions along the slope in each plot and at three depths (0,20, 20,40, 40,60 cm) during the period 1999,2001, years with different rainfall characteristics, including extreme events and long dry periods. Rainfall was recorded in the experimental field using a pluviometer linked to a data-logger. Runoff rates and yield were evaluated at the same positions. For the same annual rainfall, the season of the year in which rainfall is recorded and its intensity are critical for water availability for crops. Soil water content varies within the plot and is related to the soil characteristics existing at the different positions of the landscape. The differences in soil depth created by soil movements in the field mechanization give rise to significant yield reductions (up to 50%) between deeper and shallow areas. In addition, for the same annual rainfall, water availability for crops depends on its distribution over the year, particularly in soils with low water-storage capacity. The yield was strongly affected in years with dry or very dry winters. Copyright © 2005 John Wiley & Sons, Ltd. [source] The effects of log erosion barriers on post-fire hydrologic response and sediment yield in small forested watersheds, southern California,HYDROLOGICAL PROCESSES, Issue 15 2001Peter M. Wohlgemuth Abstract Wildfire usually promotes flooding and accelerated erosion in upland watersheds. In the summer of 1999, a high-severity wildfire burned a series of mixed pine/oak headwater catchments in the San Jacinto Mountains of southern California. Log erosion barriers (LEBs) were constructed across much of the burned area as an erosion control measure. We built debris basins in two watersheds, each about 1 ha in area, one with LEBs, the other without, to measure post-fire hydrologic response and sediment yield and to evaluate the effectiveness of the LEBs. The watersheds are underlain by granitic bedrock, producing a loamy sand soil above large extents of weathered bedrock and exposed core stones (tors) on the surface. Measured soil water-repellency was similar over the two catchments. Rain gauges measured 348 mm of precipitation in the first post-fire year. The ephemeral stream channels experienced surface flow after major rainstorms, and the source of the water was throughflow exfiltration at the slope/channel interface. Post-fire overland flow produced some rilling, but hillslope erosion measured in silt fences away from any LEBs was minor, as was sediment accumulation behind the LEBs. Stream channels in the catchments exhibited minor net scour. Water yield was much greater in the LEB-treated watershed. This resulted in 14 times more sediment yield by weight than the untreated watershed. Average soil depths determined by augering were nearly double in the catchment without the LEBs compared with the treated watershed. This suggests that differences in water and sediment yield between the two catchments are due to the twofold difference in the estimated soil water-holding capacity in the untreated watershed. It appears that the deeper soils in the untreated watershed were able to retain most of the precipitation, releasing less water to the channels and thereby reducing erosion and sediment yield. Thus, the test of LEB effectiveness was inconclusive in this study, because soil depth and soil water-holding capacity may have masked their performance. Published in 2001 John Wiley & Sons, Ltd. [source] A conceptual model of preferential flow systems in forested hillslopes: evidence of self-organizationHYDROLOGICAL PROCESSES, Issue 10 2001Roy C. Sidle Abstract Preferential flow paths are known to be important conduits of subsurface stormflow in forest hillslopes. Earlier research on preferential flow paths focused on vertical transport; however, lateral transport is also evident in steep forested slopes underlain by bedrock or till. Macropores consisting of decayed and live roots, subsurface erosion, surface bedrock fractures, and animal burrows form the basis of a ,backbone' for lateral preferential flow in such sites. Evidence from field studies in Japan indicates that although individual macropore segments are generally <0·5 m in length, they have a tendency to self-organize into larger preferential flow systems as sites become wetter. Staining tests show clear evidence of interconnected macropore flow segments, including: flow within decayed root channels and subsurface erosion cavities; flow in small depressions of the bedrock substrate; fracture flow in weathered bedrock; exchange between macropores and mesopores; and flow at the organic horizon,mineral soil interface and in buried pockets of organic material and loose soil. Here we develop a three-dimensional model for preferential flow systems based on distributed attributes of macropores and potential connecting nodes (e.g. zones of loose soil and buried organic matter). We postulate that the spatially variable and non-linear preferential flow response observed at our Japan field site, as well as at other sites, is attributed to discrete segments of macropores connecting at various nodes within the regolith. Each node is activated by local soil water conditions and is influenced strongly by soil depth, permeability, pore size, organic matter distribution, surface and substrate topography, and possibly momentum dissipation. This study represents the first attempt to characterize the spatially distributed nature of preferential flow paths at the hillslope scale and presents strong evidence that these networks exhibit complex system behaviour. Copyright © 2001 John Wiley & Sons, Ltd. [source] Steady infiltration from buried point source into heterogeneous cross-anisotropic unsaturated soilINTERNATIONAL JOURNAL FOR NUMERICAL AND ANALYTICAL METHODS IN GEOMECHANICS, Issue 10 2004G. J. Chen Abstract The paper presents the analytical solution for the steady-state infiltration from a buried point source into two types of heterogeneous cross-anisotropic unsaturated half-spaces. In the first case, the heterogeneity of the soil is modelled by an exponential relationship between the hydraulic conductivity and the soil depth. In the second case, the heterogeneous soil is represented by a multilayered half-space where each layer is homogeneous. The hydraulic conductivity varies exponentially with moisture potential and this leads to the linearization of the Richards equation governing unsaturated flow. The analytical solution is obtained by using the Hankel integral transform. For the multilayered case, the combination of a special forward and backward transfer matrix techniques makes the numerical evaluation of the solution very accurate and efficient. The correctness of both formulations is validated by comparison with alternative solutions for two different cases. The results from typical cases are presented to illustrate the influence on the flow field of the cross-anisotropic hydraulic conductivity, the soil heterogeneity and the depth of the source. Copyright © 2004 John Wiley & Sons, Ltd. [source] Drip Irrigation Frequency: The Effects and Their Interaction with Nitrogen Fertilization on Sandy Soil Water Distribution, Maize Yield and Water Use Efficiency Under Egyptian ConditionsJOURNAL OF AGRONOMY AND CROP SCIENCE, Issue 3 2008S. E. El-Hendawy Abstract Irrigation frequency is one of the most important factors in drip irrigation scheduling that affects the soil water regime, the water and fertilization use efficiency and the crop yield, although the same quantity of water is applied. Therefore, field experiments were conducted for 2 years in the summer season of 2005 and 2006 on sandy soils to investigate the effects of irrigation frequency and their interaction with nitrogen fertilization on water distribution, grain yield, yield components and water use efficiency (WUE) of two white grain maize hybrids (Zea mays L.). The experiment was conducted by using a randomized complete block split-split plot design, with four irrigation frequencies (once every 2, 3, 4 and 5 days), two nitrogen levels (190 and 380 kg N ha,1), and two maize hybrids (three-way cross 310 and single cross 10) as the main-plot, split-plot, and split-split plot treatments respectively. The results indicate that drip irrigation frequency did affect soil water content and retained soil water, depending on soil depth. Grain yield with the application of 190 kg N ha,1 was not statistically different from that at 380 kg N ha,1 at the irrigation frequency once every 5 days. However, the application of 190 kg N ha,1 resulted in a significant yield reduction of 25 %, 18 % and 9 % in 2005 and 20 %, 13 % and 6 % in 2006 compared with 380 kg N ha,1 at the irrigation frequencies once every 2, 3 and 4 days respectively. The response function between yield components and irrigation frequency treatments was quadratic in both growing seasons except for 100-grain weight, where the function was linear. WUE increased with increasing irrigation frequency and nitrogen levels, and reached the maximum values at once every 2 and 3 days and at 380 kg N ha,1. In order to improve the WUE and grain yield for drip-irrigated maize in sandy soils, it is recommended that irrigation frequency should be once every 2 or 3 days at the investigated nitrogen levels of 380 kg N ha,1 regardless of maize varieties. However, further optimization with a reduced nitrogen application rate should be aimed at and will have to be investigated. [source] Spatio-temporal variation in fruit production and seed predation in a perennial herb influenced by habitat quality and population sizeJOURNAL OF ECOLOGY, Issue 2 2008Jon Ågren Summary 1In patchily distributed plant species, seed production is likely to be influenced both by local abiotic factors affecting plant size and conditions for fruit maturation, and by population characteristics affecting the intensity of interactions with mutualists and antagonists. However, the relative importance of these effects is poorly known. 2We used multiple regression and path models to examine the importance of abiotic factors (sun exposure, soil depth) and population characteristics (size, density and connectivity) for variation in flower and fruit production and intensity of seed predation among 39 populations of the long-lived herb Vincetoxicum hirundinaria in three consecutive years. In addition, we manipulated water availability in a field experiment and recorded short-term and long-term effects on fruit output, and conducted a supplemental hand-pollination experiment. 3Flower production varied little, while fruit initiation, fruit abortion and fruit predation varied considerably among years. Sun exposure and soil depth affected fruit production per plant indirectly and positively through their effects on flower number. Population density affected fruit production negatively through its effect on flower number. Both fruit initiation and the proportion of fruits attacked by the tephritid fly Euphranta connexa were related positively to population size. 4The number of full-size fruits per plant was related positively to sun exposure and population size in two years each, and related negatively to population density in one year. However, because of seed predation, the number of intact mature fruits was related significantly to population characteristics in only one of three years. 5The field experiments showed that both shortage of water and insufficient pollination may limit fruit set in V. hirundinaria. 6Synthesis. These results demonstrate that the relative importance of local abiotic conditions and population characteristics may vary considerably along the chain of events from flower formation to intact fruit, and also among years. They further show that, at least in species with a naturally patchy distribution, connectivity may be relatively unimportant for variation in reproductive output compared to effects of habitat quality, population size and density. [source] Grassland diversity related to the Late Iron Age human population densityJOURNAL OF ECOLOGY, Issue 3 2007MEELIS PÄRTEL Summary 1Species-rich semi-natural grasslands in Europe developed during prehistoric times and have endured due to human activity. At the same time, intensive grassland management or changes in land use may result in species extinction. As a consequence, plant diversity in semi-natural calcareous grasslands may be related to both historical and current human population density. 2We hypothesize that current vascular plant diversity in semi-natural calcareous grasslands is positively correlated with the Late Iron Age (c. 800,1000 years ago) density of human settlements (indicated by Late Iron Age fortresses and villages) due to enhancement of grassland extent and species dispersal, and negatively correlated with current human population density due to habitat loss and deterioration. 3We described the size of the community vascular plant species pool, species richness per 1 m2 and the relative richness (richness divided by the size of the species pool) in 45 thin soil, calcareous (alvar) grasslands in Estonia. In addition to historical and current human population density we considered simultaneously the effects of grassland area, connectivity to other alvar grasslands, elevation above sea level (indicating grassland age), soil pH, soil N, soil P, soil depth, soil depth heterogeneity, geographical east,west gradient, precipitation and spatial autocorrelation. 4Both the size of the community species pool and the species richness are significantly correlated with the Late Iron Age human population density. In addition, species richness was unimodally related to the current human population density. The relative richness (species ,packing density') was highest in the intermediate current human population densities, indicative of moderate land-use intensity. 5Community species pool size decreased non-linearly with increasing soil N, and was highest at intermediate elevation. Small-scale richness was greater when sites were well connected and when the elevation was intermediate. Spatial autocorrelation was also significant for both species pool size and small-scale richness. 6In summary, human land-use legacy from prehistoric times is an important aspect in plant ecology, which could be an important contributor to the current variation in biodiversity. [source] | |||||||||||||||||||||||||||||||||||||