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Shallow Water (shallow + water)
Terms modified by Shallow Water Selected AbstractsThe effects of low summer flow on wild salmon (Salmo salar), trout (Salmo trutta) and grayling (Thymallus thymallus) in a small streamFRESHWATER BIOLOGY, Issue 12 2009W. D. RILEY Summary 1.,The effects of an experimentally imposed low summer flow on habitat use, displacement and survival of wild populations of juvenile salmon (Salmo salar), trout (Salmo trutta) and grayling (Thymallus thymallus) were investigated in a chalk stream. The habitat use and mobility of the fish in response to reduced flow was determined using passive integrated transponder-tag detector systems. 2.,Habitat use was compared to that available under different flow regimes. These consisted of an initial control phase of normal summer flow, an abrupt step change to 21 days of low flow, followed by a second control phase when normal summer flow was reinstated. First year (0+) salmon showed little change in their preferred substratum during low flow, whilst 1+ salmon increased their use of gravel and reduced that of mud during the day. Both 0+ and 1+ salmon were found in relatively deep water by day under low flow, whilst 1+ salmon preferred relatively shallow water at night. First year trout increased their use of gravel and reduced their use of mud and submerged tree roots under low flow, using relatively deep and fast flowing water by day. Older trout increased their use of gravel and occupied relatively deep, slow flowing water by day and relatively fast and shallow water at night. Grayling showed little change in their preferred substratum, but occupied relatively shallow water following the introduction of low flow. 3.,The range of movement of juvenile salmon increased at night under low flow, and was greater by day under normal flow. There was also an initial increase in the range of movement of 0+ trout following the introduction of low flow. Older trout initially moved less under low flow. With reduced flow, the range of movement by grayling increased significantly during the day. 4.,There was no net downstream displacement of any species with reduced flow, but the mortality rate in 0+ salmon, trout and grayling increased. This may be related to their small size and increased vulnerability to predation under low flow, and due to the reduction in depth and loss of the stream margins that are normally the preferred habitat of 0+ trout and grayling. 5.,The findings of this field study have implications for the management of braided, and highly regulated, chalk stream systems. In particular, they underline the importance of the stream margins as juvenile salmonid habitat, and suggest that a flow management strategy is required to mitigate for drought conditions. Such a strategy might include pre-emptive controls on abstraction and the maintenance of river flow via a prioritised route, predetermined using fish or habitat surveys, to minimise the effects of drought conditions on the more vulnerable or valued fish groups. [source] Micro-scale sulphur isotope evidence for sulphur cycling in the late Archean shallow oceanGEOBIOLOGY, Issue 1 2007B. S. KAMBER ABSTRACT We report in situ secondary ion mass spectrometer sulphur isotope data for sedimentary pyrite from the 2.52 Ga Upper Campbellrand Subgroup, Transvaal, South Africa. The analysed sedimentary rocks represent a transition in depositional environment from very shallow to deeper water, with strong sedimentological, facies distribution and geochemical evidence for the presence of a shallow redox chemocline. Data were obtained directly in thin section in order to preserve petrographic context. They reveal a very large extent of isotopic fractionation both in mass-independent (MIF) and in mass-dependent fractionation (MDF) on unprecedentedly small scale. In the shallow-water microbical carbonates, three types of pyrite were identified. The texturally oldest pyrite is found as small, isotopically little fractionated grains in the microbial mats. Large (several mm) spheroidal pyrite concretions, which postdate the mat pyrite, record strong evidence for an origin by bacterial sulphate reduction. Rare pyrite surrounding late fenestral calcite is inferred to have formed from recycled bacterial pyrite on account of the slope of its correlated MIF and MDF array. This latter type of pyrite was also found in an interbedded black shale and a carbonate laminite. In a deeper water chert, pyrite with very heavy sulphur indicates partial to almost complete sulphate reduction across a chemocline whose existence has been inferred independently. The combined picture from all the studied samples is that of a sulphate availability-limited environment, in which sulphur was cycled between reservoirs according to changing redox conditions established across the chemocline. Cycling apparently reduced the extent of recorded sulphur isotope fractionation relative to what is expected from projection in the correlated MIF and MDF arrays. This is consistent with regionally relatively high free oxygen concentrations in the shallow water, permitting locally strong MDF. Our new observations add to the growing evidence for a complex, fluctuating evolution of free atmospheric oxygen between c. 2.7 Ga and 2.3 Ga. [source] Darriwilian (Middle Ordovician) graptolite faunas of the Sandia Region, southern PeruGEOLOGICAL JOURNAL, Issue 4 2010Jörg Maletz Abstract Ordovician graptolite faunas of Peru are restricted to a short interval in the Middle to basal Upper Ordovician, found in three regions of the country. All Peruvian graptolite faunas are strongly dominated by shallow water elements of the Atlantic Faunal Realm, represented largely by Didymograptus s. str. and Aulograptus, but a number of faunal elements of the pandemic isograptid biofacies have recently been discovered in the Sandia Region of SE Peru. Peruvian graptolite faunas are reviewed and the new records from the Sandia Region are discussed in detail. The faunas from the Purumpata and Iparo members of the San José Formation range in age approximately from the Undulograptus austrodentatus Biozone to the Holmograptus lentus Biozone (early to middle Darriwilian). The faunas provide a better understanding of faunal composition and diversity in this region and help to correlate shallow water and deeper water graptolite faunas from this time interval. Biserial graptolites are rare in most samples and usually indeterminable, but a single identifiable specimen of Undulograptus austrodentatus was found, indicating a level close to the base of the Darriwilian. A number of specimens of the genera Isograptus and Arienigraptus from the Sandia Region represent pandemic graptolite faunas of the isograptid biofacies, described for the first time from this region. Copyright © 2009 John Wiley & Sons, Ltd. [source] Sequence stratigraphy of the upper Millstone Grit (Yeadonian, Namurian), North WalesGEOLOGICAL JOURNAL, Issue 5 2007Rhodri M. Jerrett Abstract The upper Millstone Grit strata (Yeadonian, Namurian) of North Wales have been studied using sedimentological facies analysis and sequence stratigraphy. These strata comprise two cyclothems, each containing prodelta shales (Holywell Shale) that pass gradationally upwards into delta-front and delta-plain deposits (Gwespyr Sandstone Formation). The deltas formed in shallow water (<100,m), were fluvial-dominated, had elongate and/or sheet geometries and are assigned to highstand systems tracts. Two delta complexes with distinctive sandstone petrographies are identified: (1) a southerly derived, quartzose delta complex sourced locally from the Wales-Brabant Massif, and (2) a feldspathic delta complex fed by a regional source(s) to the north and/or west. The feldspathic delta complex extended further south in the younger cyclothem. A multistorey braided-fluvial complex (Aqueduct Grit, c. 25,m thick) is assigned to a lowstand systems tract, and occupies an incised valley that was eroded into the highstand feldspathic delta complex in the younger cyclothem. A candidate incised valley cut into the highstand feldspathic delta complex in the older cyclothem is also tentatively identified. Transgressive systems tracts are thin (<5,m) and contain condensed fossiliferous shales (marine bands). The high-resolution sequence stratigraphic framework interpreted for North Wales can be readily traced northwards into the Central Province Basin (,Pennine Basin'), supporting the notion that high-frequency, high-magnitude sea-level changes were the dominant control on stratigraphic architecture. Copyright © 2007 John Wiley & Sons, Ltd. [source] Bird species numbers in an archipelago of reeds at Lake Velence, HungaryGLOBAL ECOLOGY, Issue 6 2000András Báldi Abstract 1,Bird species numbers were studied on 109 reed islands at Lake Velence, Hungary, in the 1993 and 1994 breeding seasons. The aim was to describe and account for the abundance and distribution patterns of the bird species. 2,It was expected that an exponential model would fit the calculated species,area curves. However, for the 1993 data, both the power function (LogS ~ LogArea) and the exponential (S ~ LogArea) models did so, while the power function, exponential and linear (S ~ A) models fitted the curves for the 1994 data. 3,The results showed that the pattern was not random: a collection of small islands held more species than a few large islands with the same total area. 4,The relative species richness of small islands is a result of the preference of most common passerine bird species for the edges of reed islands. Most individuals were found in the first 5 m of the reedbed, and no edge avoidance was detected on a local spatial scale. Large, rarer species (e.g. Great White Egret), however, were found to be dependent on large reed islands. 5,Comparison of results with two other studies on bird communities of reed islands revealed that the type of landscape matrix (e.g. deep water, shallow water or agricultural lands) among reed patches significantly influences bird communities. Deep water was dominated by grebes and coot, shallow water by reed-nesting passerines, and farmed areas by reed- and bush-nesting passerines. [source] The natural volume method (NVM): Presentation and application to shallow water inviscid flowsINTERNATIONAL JOURNAL FOR NUMERICAL METHODS IN FLUIDS, Issue 1 2009R. Ata Abstract In this paper a fully Lagrangian formulation is used to simulate 2D shallow water inviscid flows. The natural element method (NEM), which has been used successfully with several solid and fluid mechanics applications, is used to approximate the fluxes over Voronoi cells. This particle-based method has shown huge potential in terms of handling problems involving large deformations. Its main advantage lies in the interpolant character of its shape function and consequently the ease it allows with respect to the imposition of Dirichlet boundary conditions. In this paper, we use the NEM collocationally, and in a Lagrangian kinematic description, in order to simulate shallow water flows that are boundary moving problems. This formulation is ultimately shown to constitute a finite-volume methodology requiring a flux computation on Voronoi cells rather than the standard elements, in a triangular or quadrilateral mesh. St Venant equations are used as the mathematical model. These equations have discontinuous solutions that physically represent the existence of shock waves, meaning that stabilization issues have thus been considered. An artificial viscosity deduced from an analogy with Riemann solvers is introduced to upwind the scheme and therefore stabilize the method. Some inviscid bidimensional flows were used as preliminary benchmark tests, which produced decent results, leading to well-founded hopes for the future of this method in real applications. Copyright © 2008 John Wiley & Sons, Ltd. [source] A mesh patching method for finite volume modelling of shallow water flowINTERNATIONAL JOURNAL FOR NUMERICAL METHODS IN FLUIDS, Issue 12 2006Keming Hu Abstract A new mesh-patching model is presented for shallow water flow described by the 2D non-linear shallow water (NLSW) equations. The mesh-patching model is based on AMAZON, a high-resolution NLSW engine with an improved HLLC approximate Riemann solver. A new patching algorithm has been developed, which not only provides improved spatial resolution of flow features in particular parts of the mesh, but also simplifies and speeds up the (structured) grid generation process for an area with complicated geometry. The new patching technique is also compatible with increasingly popular parallel computing and adaptive grid techniques. The patching algorithm has been tested with moving bores, and results of test problems are presented and compared to previous work. Copyright © 2005 John Wiley & Sons, Ltd. [source] Extension of an explicit finite volume method to large time steps (CFL>1): application to shallow water flowsINTERNATIONAL JOURNAL FOR NUMERICAL METHODS IN FLUIDS, Issue 1 2006J. Murillo Abstract In this work, the explicit first order upwind scheme is presented under a formalism that enables the extension of the methodology to large time steps. The number of cells in the stencil of the numerical scheme is related to the allowable size of the CFL number for numerical stability. It is shown how to increase both at the same time. The basic idea is proposed for a 1D scalar equation and extended to 1D and 2D non-linear systems with source terms. The importance of the kind of grid used is highlighted and the method is outlined for irregular grids. The good quality of the results is illustrated by means of several examples including shallow water flow test cases. The bed slope source terms are involved in the method through an upwind discretization. Copyright © 2005 John Wiley & Sons, Ltd. [source] Hybrid finite-volume finite-difference scheme for the solution of Boussinesq equationsINTERNATIONAL JOURNAL FOR NUMERICAL METHODS IN FLUIDS, Issue 11 2005K. S. Erduran Abstract A hybrid scheme composed of finite-volume and finite-difference methods is introduced for the solution of the Boussinesq equations. While the finite-volume method with a Riemann solver is applied to the conservative part of the equations, the higher-order Boussinesq terms are discretized using the finite-difference scheme. Fourth-order accuracy in space for the finite-volume solution is achieved using the MUSCL-TVD scheme. Within this, four limiters have been tested, of which van-Leer limiter is found to be the most suitable. The Adams,Basforth third-order predictor and Adams,Moulton fourth-order corrector methods are used to obtain fourth-order accuracy in time. A recently introduced surface gradient technique is employed for the treatment of the bottom slope. A new model ,HYWAVE', based on this hybrid solution, has been applied to a number of wave propagation examples, most of which are taken from previous studies. Examples include sinusoidal waves and bi-chromatic wave propagation in deep water, sinusoidal wave propagation in shallow water and sinusoidal wave propagation from deep to shallow water demonstrating the linear shoaling properties of the model. Finally, sinusoidal wave propagation over a bar is simulated. The results are in good agreement with the theoretical expectations and published experimental results. Copyright © 2005 John Wiley & Sons, Ltd. [source] Coupling between shallow water and solute flow equations: analysis and management of source terms in 2DINTERNATIONAL JOURNAL FOR NUMERICAL METHODS IN FLUIDS, Issue 3 2005J. Murillo Abstract A two-dimensional model for the simulation of solute transport by convection and diffusion into shallow water flow over variable bottom is presented. It is based on a finite volume method over triangular unstructured grids. A first order upwind technique is applied to solve the flux terms in both the flow and solute equations and the bed slope source terms and a centred discretization is applied to the diffusion and friction terms. The convenience of considering the fully coupled system of equations is indicated and the methodology is well explained. Three options are suggested and compared in order to deal with the diffusion terms. Some comparisons are carried out in order to show the performance in terms of accuracy and computational effort of the different options. Copyright © 2005 John Wiley & Sons, Ltd. [source] Modelling of wetting and drying of shallow water using artificial porosityINTERNATIONAL JOURNAL FOR NUMERICAL METHODS IN FLUIDS, Issue 11 2005B. van't Hof Abstract A new method for wetting and drying in two-dimensional shallow water flow models is proposed. The method is closely related to the artificial porosity method used by different authors in Boussinesq-type models, but is further extended for use in a semi-implicit (ADI-type) time integration scheme. The method is implemented in the simulation model WAQUA using general boundary fitted coordinates and is applied to realistic schematization for a portion of the river Meuse in the Netherlands. A large advantage of the artificial porosity method over traditionally used methods on the basis of ,screens' is a strongly reduced sensitivity of model results. Instead of blocking all water transport in grid points where the water level becomes small, as in screen-based methods, the flow is gradually closed off. Small changes in parameters such as the initial conditions or bottom topography therefore no longer lead to large changes in the model results. Copyright © 2005 John Wiley & Sons, Ltd. [source] A stabilized SPH method for inviscid shallow water flows,INTERNATIONAL JOURNAL FOR NUMERICAL METHODS IN FLUIDS, Issue 2 2005Riadh Ata Abstract In this paper, the smoothed particle hydrodynamics (SPH) method is applied to the solution of shallow water equations. A brief review of the method in its standard form is first described then a variational formulation using SPH interpolation is discussed. A new technique based on the Riemann solver is introduced to improve the stability of the method. This technique leads to better results. The treatment of solid boundary conditions is discussed but remains an open problem for general geometries. The dam-break problem with a flat bed is used as a benchmark test. Copyright © 2004 John Wiley & Sons, Ltd. [source] A staggered conservative scheme for every Froude number in rapidly varied shallow water flowsINTERNATIONAL JOURNAL FOR NUMERICAL METHODS IN FLUIDS, Issue 12 2003G. S. Stelling Professor Abstract This paper proposes a numerical technique that in essence is based upon the classical staggered grids and implicit numerical integration schemes, but that can be applied to problems that include rapidly varied flows as well. Rapidly varied flows occur, for instance, in hydraulic jumps and bores. Inundation of dry land implies sudden flow transitions due to obstacles such as road banks. Near such transitions the grid resolution is often low compared to the gradients of the bathymetry. In combination with the local invalidity of the hydrostatic pressure assumption, conservation properties become crucial. The scheme described here, combines the efficiency of staggered grids with conservation properties so as to ensure accurate results for rapidly varied flows, as well as in expansions as in contractions. In flow expansions, a numerical approximation is applied that is consistent with the momentum principle. In flow contractions, a numerical approximation is applied that is consistent with the Bernoulli equation. Both approximations are consistent with the shallow water equations, so under sufficiently smooth conditions they converge to the same solution. The resulting method is very efficient for the simulation of large-scale inundations. Copyright © 2003 John Wiley & Sons, Ltd. [source] Relaxation schemes for the shallow water equationsINTERNATIONAL JOURNAL FOR NUMERICAL METHODS IN FLUIDS, Issue 7 2003A. I. Delis Abstract We present a class of first and second order in space and time relaxation schemes for the shallow water (SW) equations. A new approach of incorporating the geometrical source term in the relaxation model is also presented. The schemes are based on classical relaxation models combined with Runge,Kutta time stepping mechanisms. Numerical results are presented for several benchmark test problems with or without the source term present. Copyright © 2003 John Wiley & Sons, Ltd. [source] Numerical simulation of the vertical structure of discontinuous flowsINTERNATIONAL JOURNAL FOR NUMERICAL METHODS IN FLUIDS, Issue 1 2001Guus S. Stelling Abstract A numerical method to solve the Reynolds-averaged Navier,Stokes equations with the presence of discontinuities is outlined and discussed. The pressure is decomposed into the sum of a hydrostatic component and a hydrodynamic component. The numerical technique is based upon the classical staggered grids and semi-implicit finite difference methods applied for quasi- and non-hydrostatic flows. The advection terms in the momentum equations are approximated in order to conserve mass and momentum following the principles recently developed for the numerical simulation of shallow water flows with large gradients. Conservation of these properties is the most important aspect to represent near local discontinuities in the solution, following from sharp bottom gradients or hydraulic jumps. The model is applied to reproduce the flow over a step where a hydraulic jump forms downstream. The hydrostatic pressure assumption fails to represent this type of flow mainly because of the pressure deviation from the hydrostatic values downstream the step. Fairly accurate results are obtained from the numerical model compared with experimental data. Deviation from the data is found to be inherent to the standard k,, model implemented. Copyright © 2001 John Wiley & Sons, Ltd. [source] Output feedback control design for station keeping of AUVs under shallow water wave disturbancesINTERNATIONAL JOURNAL OF ROBUST AND NONLINEAR CONTROL, Issue 13 2009Shuyong Liu Abstract In this paper, we consider the problem of autonomous underwater vehicle (AUV) station keeping (SK) in shallow water area. During SK, an AUV is required to maintain position and orientation with respect to a fixed reference point at the sea floor. When AUV operates in shallow water, high-frequency disturbances due to waves will significantly affect the motion of the AUV. In order to derive wave disturbance information for control purposes, a nonlinear observer is first designed to estimate the shallow water wave velocities and AUV relative velocities by using position and attitude measurement. Using the observer estimates, a nonlinear output feedback controller is subsequently synthesized by applying observer backstepping technique. Global exponential stability (GES) of the proposed nonlinear observer,controller design is proved through Lyapunov stability theory. Simulation studies on a model based on an actual AUV were performed to verify the performance of the proposed nonlinear observer and output feedback controller. Copyright © 2008 John Wiley & Sons, Ltd. [source] Variations in the retinal designs of pulmonate snails (Mollusca, Gastropoda): squaring phylogenetic background and ecophysiological needs (I)INVERTEBRATE BIOLOGY, Issue 2 2004Marina V. Bobkova Abstract. The eyes of aquatic pulmonates differ from those of terrestrial pulmonates; the latter, in species such as Cepaea nemoralis and Trichia hispida, possess conventional, cup-shaped retinas, but the aquatic species Lymnaea stagnalis, Radix peregra, Physa fontinalis, and Planorbarius corneus have retinas that are partitioned into dorsal and ventral depressions ("pits"). The pits are separated by an internal ridge, called the "crest", and on account of their pigmentation can be seen in vivo. The dominant cellular components of the retinae of terrestrial as well as aquatic snails are pigmented cells and microvillar photoreceptors, the latter occurring in two morphologically distinct types (I and II). Aquatic snails with preferences for shallow water possess eyes with both type I and type II photoreceptive cells, but Pl. corneus, an inhabitant of deeper water, only has type-I receptors, supporting an earlier finding that type I cells represent dim- and type II cells bright-light receptors. On the basis of histological and optical comparisons, we conclude that the eyes of L. stagnalis and R. peregra, species that are known to escape and seek temporary refuge above the water surface, are well adapted to function in water as well as air, but that the eyes of P. fontinalis and Pl. corneus are less modified from those of their terrestrial ancestors. [source] Diving in shallow water: the foraging ecology of darters (Aves: Anhingidae)JOURNAL OF AVIAN BIOLOGY, Issue 4 2007Peter G. Ryan Diving birds have to overcome buoyancy, especially when diving in shallow water. Darters and anhingas (Anhingidae) are specialist shallow-water divers, with adaptations for reducing their buoyancy. Compared to closely-related cormorants (Phalacrocoracidae), darters have fully wettable plumage, smaller air sacs and denser bones. A previous study of darter diving behaviour reported no relationship between dive duration and water depth, contrary to optimal dive models. In this study I provide more extensive observations of African darters Anhinga melanogaster rufa diving in water<5 m deep at two sites. Dive duration increases with water depth at both sites, but the relationship is weak. Dives were longer than dives by cormorants in water of similar depth (max 108 s in water 2.5 m deep), with dives of up to 68 s observed in water<0.5 m deep. Initial dives in a bout were shorter than expected, possibly because their plumage was not fully saturated. Dive efficiency (dive:rest ratio) was 5,6, greater than cormorants (2.7±0.4 for 18 species) and other families of diving birds (average 0.2,4.3). Post-dive recovery periods increased with dive duration, but only slowly, resulting in a strong increase in efficiency with dive duration. All dives are likely to fall within the theoretical anaerobic dive limit. Foraging bouts were short (17.8±4.3 min) compared to cormorants, with birds spending 80±5% of time underwater. Darters take advantage of their low buoyancy to forage efficiently in shallow water, and their slow, stealthy dives are qualitatively different from those of other diving birds. However, they are forced to limit the duration of foraging bouts by increased thermoregulatory costs associated with wettable plumage. [source] The island rule and the evolution of body size in the deep seaJOURNAL OF BIOGEOGRAPHY, Issue 9 2006Craig R. McClain Abstract Aim, Our goal is to test the generality of the island rule , a graded trend from gigantism in small-bodied species to dwarfism in large-bodied species , in the deep sea, a non-insular but potentially analogous system. Location, Shallow-water and deep-sea benthic habitats in the western Atlantic Ocean from the North to South Poles. Methods, We conducted regression analyses of body size of deep-sea gastropods species relative to their shallow-water congeners using measurements from the Malacolog ver. 3.3.3 database. Results, Our results indicate that, consistent with the island rule, gastropod genera with small-bodied, shallow-water species have significantly larger deep-sea representatives, while the opposite is true for genera that are large-bodied in shallow water. Bathymetric body size clines within the deep sea are also consistent with predictions based on the island rule. Main conclusions, Like islands, the deep sea is characterized by low absolute food availability, leading us to hypothesize that the island rule is a result of selection on body size in a resource-constrained environment. The body size of deep-sea species tends to converge on an optimal size for their particular ecological strategy and habitat. [source] Marine nematode deep-sea biodiversity , hyperdiverse or hype?JOURNAL OF BIOGEOGRAPHY, Issue 4 2003P. John D. Lambshead Abstract Nematodes have been identified as a potentially hyperdiverse group and the deep sea as a potentially hyperdiverse environment (i.e. > 1 million species). A large-scale data set from the equatorial central Pacific is used to estimate regional diversity with results that challenge this view; regional diversity is higher in some coastal waters despite lower sample diversity in coastal waters than in the deep sea. The data suggests a paradigm where the deep sea has modest regional diversity, despite high local diversity through patch dynamics, because similar patches in a similar habitat are repeated for considerable distances. Disturbance in shallow water dominates over patch-dynamic mechanisms reducing local diversity but regional diversity is high because of the close packing of multiple habitats within a single region. The Pacific data are also used to demonstrate the pitfalls of extrapolating from local to global diversity. There is no reason to conclude that nematodes are less diverse than other benthic groups, indeed where direct comparison is possible the Nematoda appear to be as diverse as the Polychaeta, the most diverse macrofaunal taxon. This analysis is not consistent with the hypothesis that either marine nematodes or the deep-sea benthos are hyperdiverse raising the question whether any environment or metazoan taxon has more than a million species. [source] Hybridization dynamics of invasive cattail (Typhaceae) stands in the Western Great Lakes Region of North America: a molecular analysisJOURNAL OF ECOLOGY, Issue 1 2010Steven E. Travis Summary 1.,By increasing vigour and broadening ecological tolerances, hybridization between native and introduced species may serve as a primary driver of invasiveness. 2.,Cattails (Typha, Typhaceae) are clonal wetland graminoids that are known to hybridize where anthropogenic influences have resulted in distributional overlap. 3.,In order to gauge the relative performance of hybrid vs. pure Typha, we characterized hybridization and clonal growth where native Typha latifolia and introduced Typha angustifolia occur together in the Western Great Lakes Region of North America. 4.,Based on microsatellite markers, we documented F1 hybrids as the most common class at five intensively sampled sites, constituting up to 90% of the genets and 99% of the ramets. Backcrosses to one or the other parent constituted 5,38% of the genets. Pure T. latifolia was rare and never constituted more than 12% of the genets. 5.,F1 hybrid genets achieved the highest mean ramet numbers at three sites, and were second in size only to T. angustifolia at two sites; however, these differences were not significant based on site-specific one-way anovas. 6.,F1 hybrids exhibited little height advantage over other Typha classes, although there was a general tendency for hybrids in relatively mixed stands to be among the tallest genets in shallow water, but among the shortest genets in deeper water. 7.,Native T. latifolia was found growing at the shallowest water depths at the only site where it was sufficiently abundant to be included in statistical comparisons. 8.,Synthesis. The role of hybridization in plant invasions can be difficult to confirm in the absence of molecular data, particularly for clonal species where the boundaries separating individuals are otherwise difficult to discern. Here, we used molecular markers to document the prevalence and performance of hybrid genets in five invasive Typha stands covering a broad area of the Western Great Lakes Region. We found an extremely high prevalence of F1 hybrids within mixed Typha stands. This, coupled with the typically larger sizes of hybrid genets, suggests that hybrids are capable of outperforming other Typha spp. and that hybridization has played an influential role in the North American cattail invasion. [source] Seasonal pattern of depth selection in smallmouth bassJOURNAL OF ZOOLOGY, Issue 2 2009C. D. Suski Abstract The current study used a stationary acoustic telemetry array to monitor the depth selection of adult smallmouth bass Micropterus dolomieu in a large, oligotrophic lake in Ontario, Canada. At an annual scale, smallmouth bass demonstrated regular, seasonal changes in inhabited depths: fish occupied shallow water during the summer (2,5 m depth) and descended to deeper water (12,15 m depth) during winter under ice. Smallmouth bass remained above the thermocline in the summer, seasonal depth patterns did not vary across fish size and movements to and from seasonal depths were closely linked to the development and degradation of the thermocline. At finer time scales, smallmouth bass exhibited diel vertical migrations in summer, with fish moving to <2 m at night, and then descending to 3,5 m during the day. This pattern remained constant during the summer period examined, and varied with size such that larger fish remained deeper than smaller fish. During winter, depth did not vary across the 3-month monitoring period (c. 14 m), but small (<2 m) changes in depth were observed periodically, suggesting limited movements were occurring. Results are further discussed in the context of climate change and reproductive success for this species. [source] Fight or flight: antipredator strategies of baleen whalesMAMMAL REVIEW, Issue 1 2008JOHN K. B. FORD ABSTRACT 1The significance of killer whale Orcinus orca predation on baleen whales (Mysticeti) has been a topic of considerable discussion and debate in recent years. Discourse has been constrained by poor understanding of predator-prey dynamics, including the relative vulnerability of different mysticete species and age classes to killer whales and how these prey animals avoid predation. Here we provide an overview and analysis of predatory interactions between killer whales and mysticetes, with an emphasis on patterns of antipredator responses. 2Responses of baleen whales to predatory advances and attacks by killer whales appear to fall into two distinct categories, which we term the fight and flight strategies. The fight strategy consists of active physical defence, including self-defence by single individuals, defence of calves by their mothers and coordinated defence by groups of whales. It is documented for five mysticetes: southern right whale Eubalaena australis, North Atlantic right whale Eubalaena glacialis, bowhead whale Balaena mysticetus, humpback whale Megaptera novaeangliae and grey whale Eschrichtius robustus. The flight strategy consists of rapid (20,40 km/h) directional swimming away from killer whales and, if overtaken and attacked, individuals do little to defend themselves. This strategy is documented for six species in the genus Balaenoptera. 3Many aspects of the life history, behaviour and morphology of mysticetes are consistent with their antipredator strategy, and we propose that evolution of these traits has been shaped by selection for reduced predation. Fight species tend to have robust body shapes and are slow but relatively manoeuvrable swimmers. They often calve or migrate in coastal areas where proximity to shallow water provides refuge and an advantage in defence. Most fight species have either callosities (rough and hardened patches of skin) or encrustations of barnacles on their bodies, which may serve (either primarily or secondarily) as weapons or armour for defence. Flight species have streamlined body shapes for high-speed swimming and they can sustain speeds necessary to outrun pursuing killer whales (>15,20 km/h). These species tend to favour pelagic habitats and calving grounds where prolonged escape sprints from killer whales are possible. 4The rarity of observed successful attacks by killer whales on baleen whales, especially adults, may be an indication of the effectiveness of these antipredator strategies. Baleen whales likely offer low profitability to killer whales, relative to some other marine mammal prey. High-speed pursuit of flight species has a high energetic cost and a low probability of success while attacks on fight species can involve prolonged handling times and a risk of serious injury. [source] Existence of periodic traveling wave solutions for the Ostrovsky equationMATHEMATICAL METHODS IN THE APPLIED SCIENCES, Issue 14 2008Naoyuki Ishimura Abstract We are concerned with the Ostrovsky equation, which is derived from the theory of weakly nonlinear long surface and internal waves in shallow water under the presence of rotation. On the basis of the variational method, we show the existence of periodic traveling wave solutions. Copyright © 2008 John Wiley & Sons, Ltd. [source] Light dependent shift in the anti-predator response of a pyralid mothOIKOS, Issue 2 2003A. Monica Svensson Male small china-mark moth Cataclysta lemnata (Pyralidae) swarming over shallow water show a flight activity that peaks during the afternoon but which sometimes is extended into the night. We exposed wild, naturally flying C. lemnata to simulated predator attacks consisting of a) bursts of ultrasound (26 kHz, simulating a bat) and b) a thrown stick (rapid movement, simulating a small bird), during day and night, respectively. We thus investigated the possibility that these moths are able to switch between defensive strategies as the predator regime shifts from insectivorous birds to bats in the evening. The defensive response differed qualitatively between day and night, as expected, but it was independent of the kind of stimulus. We thus demonstrate a previously unknown flexibility in the defensive strategy of moths. [source] Patterns of permafrost formation and degradation in relation to climate and ecosystemsPERMAFROST AND PERIGLACIAL PROCESSES, Issue 1 2007Y. L. Shur Abstract We develop a permafrost classification system to describe the complex interaction of climatic and ecological processes in permafrost formation and degradation that differentiates five patterns of formation: ,climate-driven'; ,climate-driven, ecosystem-modified'; ,climate-driven, ecosystem-protected'; ,ecosystem-driven'; and ,ecosystem-protected' permafrost. Climate-driven permafrost develops in the continuous permafrost zone, where permafrost forms immediately after the surface is exposed to the atmosphere and even under shallow water. Climate-driven, ecosystem-modified permafrost occurs in the continuous permafrost zone when vegetation succession and organic-matter accumulation lead to development of an ice-rich layer at the top of the permafrost. During warming climates, permafrost that has formed as climate-driven can occur in the discontinuous permafrost zone, where it can persist for a long time as ecosystem-protected. Climate-driven, ecosystem protected permafrost, and its associated ground ice, cannot re-establish in the discontinuous zone once degraded, although the near surface can recover as ecosystem-driven permafrost. Ecosystem-driven permafrost forms in the discontinuous permafrost zone in poorly drained, low-lying and north-facing landscape conditions, and under strong ecosystem influence. Finally, ecosystem-protected permafrost persists as sporadic patches under warmer climates, but cannot be re-established after disturbance. These distinctions are important because the various types react differently to climate change and surface disturbances. For example, climate-driven, ecosystem-modified permafrost can experience thermokarst even under cold conditions because of its ice-rich layer formed during ecosystem development, and ecosystem-driven permafrost is unlikely to recover after disturbance, such as fire, if there is sufficient climate warming. Copyright © 2007 John Wiley & Sons, Ltd. [source] Salamander Blue-sensitive Cones Lost During Metamorphosis,PHOTOCHEMISTRY & PHOTOBIOLOGY, Issue 4 2008Ying Chen The tiger salamander lives in shallow water with bright light in the aquatic phase, and in dim tunnels or caves in the terrestrial phase. In the aquatic phase, there are five types of photoreceptors,two types of rods and three types of cones. Our previous studies showed that the green rods and blue-sensitive cones contain the same visual pigment and have the same absorbance spectra; however, the green rods have a larger photon-catch area and thus have higher light sensitivity than the blue-sensitive cones. Here we show that after metamorphosis, the terrestrial salamander looses the blue-sensitive cones, while the density of the green rods increases. Moreover, the size of the green rod outer segments is increased in the terrestrial phase, compared to that in the aquatic phase. This switch from the blue-sensitive cones to the green rods may represent an adaptation to the dim light environment of the terrestrial phase. [source] Shallow-water habitats as sources of fallback foods for homininsAMERICAN JOURNAL OF PHYSICAL ANTHROPOLOGY, Issue 4 2009Richard Wrangham Abstract Underground storage organs (USOs) have been proposed as critical fallback foods for early hominins in savanna, but there has been little discussion as to which habitats would have been important sources of USOs. USOs consumed by hominins could have included both underwater and underground storage organs, i.e., from both aquatic and terrestrial habitats. Shallow aquatic habitats tend to offer high plant growth rates, high USO densities, and relatively continuous USO availability throughout the year. Baboons in the Okavango delta use aquatic USOs as a fallback food, and aquatic or semiaquatic USOs support high-density human populations in various parts of the world. As expected given fossilization requisites, the African early- to mid-Pleistocene shows an association of Homo and Paranthropus fossils with shallow-water and flooded habitats where high densities of plant-bearing USOs are likely to have occurred. Given that early hominins in the tropics lived in relatively dry habitats, while others occupied temperate latitudes, ripe, fleshy fruits of the type preferred by African apes would not normally have been available year round. We therefore suggest that water-associated USOs were likely to have been key fallback foods, and that dry-season access to aquatic habitats would have been an important predictor of hominin home range quality. This study differs from traditional savanna chimpanzee models of hominin origins by proposing that access to aquatic habitats was a necessary condition for adaptation to savanna habitats. It also raises the possibility that harvesting efficiency in shallow water promoted adaptations for habitual bipedality in early hominins. Am J Phys Anthropol 140:630,642, 2009. © 2009 Wiley-Liss, Inc. [source] On the representation of gravity waves in numerical models of the shallow-water equationsTHE QUARTERLY JOURNAL OF THE ROYAL METEOROLOGICAL SOCIETY, Issue 563 2000A. R. Mohebalhojeh Abstract Gravity waves, or imbalanced motions, that develop during the evolution of vortical flows in numerical models of the shallow water (SW) equations are examined in detail. The focus here is on nearly-balanced flows, with small but non-zero gravity-wave activity. For properly initialized flows, it is reasonable to expect small GW activity when Froude numbers Fr < 1 and Rossby numbers Ro , 1. The guiding principle in the present study is that an accurate representation of potential vorticity (PV) is the pre-requisite to a fair assessment of the generation of gravity waves. The contour-advective semi-Lagrangian (CASL) algorithm for the SW equations is applied, as it shows a remarkable improvement in the simulation of PV. However, it is shown that the standard CASL algorithm for SW leads to a noticeable numerical generation of gravity waves. The false generation of gravity waves can equivalently be thought of as the false, or numerical, breakdown of balance. In order to understand the maintenance of balance in the SW equations, a hierarchy of CASL algorithms is introduced. The main idea behind the new hierarchy is to implement PV inversion partially, balancing algorithms directly within the SW algorithm, while still permitting imbalanced motions. The results of the first three members of the hierarchy, CA0 (standard CASL), CA1, and CA2, are described and are compared with the results of two other SW algorithms, a pseudo-spectral and a semi-Lagrangian one. The main body of results is obtained for a highly ageostrophic regime of flow, with|Ro|max , 1 and Frmax , 0.5, where sub-index 'max' denotes maximum over the domain. Other flow regimes in the relevant parts of the Ro-Fr parameter space are also explored. It is found that, for a given resolution and Froude number, there is an optimal CASL algorithm, i.e. one which gives rise to the least numerical generation of gravity waves. [source] Seasonal changes in distribution of Hector's dolphin at Banks Peninsula, New Zealand: implications for protected area designAQUATIC CONSERVATION: MARINE AND FRESHWATER ECOSYSTEMS, Issue 1 2010William Rayment Abstract 1.The efficacy of protected areas for wildlife management is largely dependent on appropriate design. It is therefore crucial that the distribution of target species is well understood. 2.The Banks Peninsula Marine Mammal Sanctuary (BPMMS) was designed to protect Hector's dolphins from bycatch in gillnets. However, the Banks Peninsula dolphin population is likely to be still declining, partly due to continued bycatch outside the boundaries of the BPMMS. 3.A three year series of aerial line-transect surveys around Banks Peninsula was carried out to investigate seasonal changes in distribution of Hector's dolphins out to 20 nautical miles (37,km) from the coast. 4.Dolphin sightings were concentrated close to shore in shallow water in summer, but were more evenly distributed throughout the study area in winter. A greater proportion of dolphins were sighted outside the 4 nautical miles (7.4,km) offshore boundary of the BPMMS in winter (mean=56%) than in summer (mean=19%) (G=88.25, df=1, P<0.001). 5.Partial Mantel tests revealed the effects of distance offshore and depth on dolphin occurrence while controlling for spatial autocorrelation and multicollinearity within the data. Distance offshore had the strongest and most consistent effect on dolphin presence, while depth had a strong effect in summer only. 6.It is proposed that restrictions on gillnetting around Banks Peninsula must be extended in order to reduce bycatch of Hector's dolphins to a sustainable level, and that a new offshore boundary of the BPMMS would be best defined by distance from the coast. Copyright © 2009 John Wiley & Sons, Ltd. [source] | |||||||||||||||||||||||||||||||||||||