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Shade
Kinds of Shade Terms modified by Shade Selected AbstractsThe effect of overlying absorbing aerosol layers on remote sensing retrievals of cloud effective radius and cloud optical depthTHE QUARTERLY JOURNAL OF THE ROYAL METEOROLOGICAL SOCIETY, Issue 598 2004Jim M. Haywood Abstract Two types of partially absorbing aerosol are included in calculations that are based on intensive aircraft observations: biomass burning aerosol characterized during the Southern AFricAn Regional science Initiative (SAFARI 2000) and mineral dust aerosol characterized during the SaHAran Dust Experiment (SHADE). Measurements during SAFARI 2000 reveal that the biomass burning aerosol layer is advected over the South Atlantic ocean at elevated altitudes above the marine boundary layer which is capped by semi-permanent stratocumulus cloud sheets. Similarly, the mineral dust is measured at elevated altitudes during SHADE resulting in transport above cloud for distances of several thousands of kilometres. We perform theoretical calculations of the effect of these partially absorbing aerosol layers on satellite retrievals of cloud effective radius and cloud optical depth, and show that, in these cases, retrievals of cloud optical depth or liquid water path are likely to be subject to systematic low biases. The theoretical calculations suggest that the cloud effective radius may be subject to a significant low bias for Moderate resolution Imaging Spectrometer (MODIS) retrievals that rely on the 0.86 and 1.63 µm radiance pair for an overlying aerosol layer of either biomass burning aerosol or mineral dust. Conversely, the cloud effective radius may be subject to a significant high bias for Advanced Very High Resolution Radiometer or MODIS retrievals that rely on the 0.63 and 3.7 µm radiance pair for an overlying aerosol layer of mineral dust. Analysis of 1 km resolution MODIS data for the SAFARI 2000 period suggests that the effective radius derived from the 0.86 and 1.63 µm radiance pair is, indeed, subject to a low bias in the presence of overlying biomass burning aerosol. These results show the difficulties associated with remote sensing retrievals, which must be kept in mind when attempting to assess any potential indirect effect. © Crown copyright 2004. [source] The SCUBA Half-Degree Extragalactic Survey (SHADES) , VIII.MONTHLY NOTICES OF THE ROYAL ASTRONOMICAL SOCIETY, Issue 1 2008SWIRE, SXDF surveys, The nature of faint submillimetre galaxies in SHADES ABSTRACT We present the optical-to-submillimetre spectral energy distributions (SEDs) for 33 radio and mid-infrared (mid-IR) identified submillimetre galaxies discovered via the SHADES 850-,m SCUBA imaging in the Subaru- XMM Deep Field (SXDF). Optical data for the sources come from the SXDF and mid- and far-IR fluxes from SWIRE. We obtain photometric redshift estimates for our sources using optical and IRAC 3.6- and 4.5-,m fluxes. We then fit SED templates to the longer wavelength data to determine the nature of the far-IR emission that dominates the bolometric luminosity of these sources. The IR template fits are also used to resolve ambiguous identifications and cases of redshift aliasing. The redshift distribution obtained broadly matches previous results for submillimetre sources and on the SHADES SXDF field. Our template fitting finds that active galactic nuclei, while present in about 10 per cent of our sources, do not contribute significantly to their bolometric luminosity. Dust heating by starbursts, with either Arp220 or M82 type SEDs, appears to be responsible for the luminosity in most sources (23/33 are fitted by Arp220 templates, 2/33 by the warmer M82 templates). 8/33 sources, in contrast, are fitted by a cooler cirrus dust template, suggesting that cold dust has a role in some of these highly luminous objects. Three of our sources appear to have multiple identifications or components at the same redshift, but we find no statistical evidence that close associations are common among our SHADES sources. Examination of rest-frame K -band luminosity suggests that ,downsizing' is underway in the submillimetre galaxy population, with lower redshift systems lying in lower mass host galaxies. Of our 33 identifications six are found to be of lower reliability but their exclusion would not significantly alter our conclusions. [source] Shade facilitates an invasive stem succulent in a chenopod shrubland in South AustraliaAUSTRAL ECOLOGY, Issue 5 2003TANJA I. LENZ Abstract The invasive stem succulent Orbea variegata (L) Haw. (Asclepiadaceae) tends to be more abundant underneath shrubs than in open areas in chenopod shrublands near Whyalla, South Australia. To assess the role of facilitation in the life cycle of O. variegata, we investigated the effect of chenopod shrubs on different life stages of the species by experimentally manipulating temperature, light, soil moisture and nutrient levels. Experimental results suggest that the reduction in light and temperature under shrubs, but not increased nutrient levels, are the main facilitative mechanisms for O. variegata. Temperatures above 30°C, which are more likely to occur on the soil surface of open areas than under shrubs, inhibited seed germination. Seedling survival at low watering frequency and the growth of established ramets were increased by 75,80% shade cloth. Ramets growing in full light contained a high concentration of anthocyanin pigments. One of the functions of these pigments is to absorb excess radiation, suggesting that O. variegata experiences radiation stress in full light. In the field O. variegata performed considerably better under Atriplex vesicaria Heward ex Benth. (Chenopodiaceae) or under 75,80% shade cloth than in full light. Monthly irrigation of 20 mm did not reduce this positive effect of the A. vesicaria canopy on O. variegata, suggesting that O. variegata is inhibited by high light intensities or temperatures, independent of water availability. In conclusion, whereas shrub canopies do not seem to be required for the establishment or survival of O. variegata, shrubs improve adult growth and can improve establishment. The possibility of exotic plants being facilitated by other plants has to be taken into account when assessing the probability and rate of invasion. [source] UK Assessment , Shades of the late 1980s?ECONOMIC OUTLOOK, Issue 4 2001Article first published online: 16 DEC 200 First page of article [source] Shades of Orientalism: Paradoxes and Problems in Indian HistoriographyHISTORY AND THEORY, Issue 2 2003Peter Heehs In Orientalism, Edward Said attempts to show that all European discourse about the Orient is the same, and all European scholars of the Orient complicit in the aims of European imperialism. There may be "manifest" differences in discourse, but the underlying "latent" orientalism is "more or less constant." This does not do justice to the marked differences in approach, attitude, presentation, and conclusions found in the works of various orientalists. I distinguish six different styles of colonial and postcolonial discourse about India (heuristic categories, not essential types), and note the existence of numerous precolonial discourses. I then examine the multiple ways exponents of these styles interact with one another by focusing on the early-twentieth-century nationalist orientalist, Sri Aurobindo. Aurobindo's thought took form in a colonial framework and has been used in various ways by postcolonial writers. An anti-British nationalist, he was by no means complicit in British imperialism. Neither can it be said, as some Saidians do, that the nationalist style of orientalism was just an imitative indigenous reversal of European discourse, using terms like "Hinduism" that had been invented by Europeans. Five problems that Aurobindo dealt with are still of interest to historians: the significance of the Vedas, the date of the vedic texts, the Aryan invasion theory, the Aryan-Dravidian distinction, and the idea that spirituality is the essence of India. His views on these topics have been criticized by Leftist and Saidian orientalists, and appropriated by reactionary "Hindutva" writers. Such critics concentrate on that portion of Aurobindo's work which stands in opposition to or supports their own views. A more balanced approach to the nationalist orientalism of Aurobindo and others would take account of their religious and political assumptions, but view their project as an attempt to create an alternative language of discourse. Although in need of criticism in the light of modern scholarship, their work offers a way to recognize cultural particularity while keeping the channels of intercultural dialogue open. [source] No Shades of Gray: The Binary Discourse of George W. Bush and an Echoing PressJOURNAL OF COMMUNICATION, Issue 2 2004Kevin Coe Binary communications represent the world as a place of polar opposites. Such conceptions of reality, although not uncommon in Western thought, take on a heightened importance when political leaders employ them in a concerted, strategic discourse in a mass media environment. With this in mind, this research offers a conception of binary discourse and uses this as a foundation to examine (a) the use of binaries by U.S. President George W. Bush in 15 national addresses, from his inauguration in January 2001 to commencement of the Iraq War in March 2003, and (b) the responses of editorials in 20 leading U.S. newspapers to the president's communications. [source] Books: Shades of Green: Business, Regulation, and EnvironmentJOURNAL OF INDUSTRIAL ECOLOGY, Issue 4 2004René Kemp [source] Shades of Melancholy in Gabriele Reuter's Aus guter Familie,THE GERMAN QUARTERLY, Issue 4 2006Lisabeth Hock First page of article [source] Religious Echoes of the Errant Text: Darker Shades of Derrida's Pathless WayTHE HEYTHROP JOURNAL, Issue 3 2003Ian Almond I employ these words, I admit, with a glance towards the operations of childbearing,but also with a glance towards those who, in a society from which I do not exclude myself, turn their eyes away when faced by the as yet unnameable which is proclaiming itself and which can do so ,only under the species of the nonspecies, in the formless, mute, infant and terrifying form of monstrosity. The question of writing could be opened only if the book was closed. The joyous wandering of the graphein then became wandering without return. [source] Soluble and Columnar Liquid Crystalline Peropyrenequinones by Coupling of Phenalenones in Caesium HydroxideCHEMISTRY - A EUROPEAN JOURNAL, Issue 19 2010Noémie Buffet Dr. Shades of violet: The nitrogen-free homologues of perylene tetracarboxylic diimide dyes have been prepared. CsOH succeeds where KOH fails to couple sterically demanding hydroxyphenalenones into the esterifiable precursors of ,-stacking long-wavelength absorbing disks (see figure). [source] Shade Effects on Phaseolus vulgaris L. Intercropped with Zea mays L. under Well-Watered ConditionsJOURNAL OF AGRONOMY AND CROP SCIENCE, Issue 3 2004M. Tsubo Abstract Field experiments were carried out under unstressed conditions of soil water during two summer crop growing seasons (1998,99 and 1999,2000 seasons) in a South African semi-arid region (Bloemfontein, Free State, South Africa). The aim of this study was to investigate shade effects on beans intercropped with maize in terms of plant mass and radiation use. The experimental treatments were two cropping systems (no shading/sole cropping and shading/intercropping) and two row orientations (north,south and east,west). At the top of bean canopies shaded by maize, incident radiation was reduced by up to 90 %. Shading reduced total dry matter of beans by 67 % at the end of the growing season, resulting in yield losses. The dry matter partitioning into leaf and stem (the ratios of leaf and stem to total biomass) was about 50 % higher in intercropping than sole cropping. In contrast, intercropped beans had 40 % lower dry matter partitioning into pod (the ratio of pod to total biomass). Fraction of radiation intercepted by sole-cropped beans steeply increased until canopy closure (0.9) and then slowly decreased, while fraction of radiation intercepted by intercropped beans remained constant between 0.0 and 0.2 throughout the growing seasons. However, intercropped beans had 77 % higher radiation use efficiency (RUE) than sole-cropped beans. In contrast, for maize, no effect of intercropping (shading) was found on growth, partitioning, yield, radiation interception or RUE. Consequently, lower bean yield losses can be attained in association with late shading rather than early shading. This can be controlled by growing crops with different temporal and spatial treatments. As regards row treatment, no effect of row direction was found on growth, partitioning, yield, radiation interception or RUE. [source] Stylized lighting for cartoon shaderCOMPUTER ANIMATION AND VIRTUAL WORLDS (PREV: JNL OF VISUALISATION & COMPUTER ANIMATION), Issue 2-3 2009Hideki Todo Abstract In the context of non-photorealistic imaging, such as digital cel animation, lighting is symbolic and stylized to depict the scene's mood and the geometric or physical features of the objects in the scene. Stylized light and shade should therefore be intentionally animated rather than rigorously simulated. However, it is difficult to achieve smooth animation of light and shade that are stylized with a user's intention, because such stylization cannot be achieved using just conventional 3D lighting. To address this problem, we propose a 3D stylized lighting method, focusing on several stylized effects including straight lighting, edge lighting, and detail lighting which are important features in hand-drawn cartoon animation. Our method is an extension of the conventional cartoon shader and introduces a light coordinate system for light shape control with smooth animations of light and shade. We also extend a toon mapping process for detailed feature lighting. Having these algorithms in a real-time cartoon shader, our prototype system allows the interactive creation of stylized lighting animations. We show several animation results obtained by our method to illustrate usefulness and effectiveness of our method. Copyright © 2009 John Wiley & Sons, Ltd. [source] Grammar-based Encoding of FacadesCOMPUTER GRAPHICS FORUM, Issue 4 2010Simon Haegler Abstract In this paper we propose a real-time rendering approach for procedural cities. Our first contribution is a new lightweight grammar representation that compactly encodes facade structures and allows fast per-pixel access. We call this grammar F -shade. Our second contribution is a prototype rendering system that renders an urban model from the compact representation directly on the GPU. Our suggested approach explores an interesting connection from procedural modeling to real-time rendering. Evaluating procedural descriptions at render time uses less memory than the generation of intermediate geometry. This enables us to render large urban models directly from GPU memory. [source] Use of Premontane Moist Forest and Shade Coffee Agroecosystems by Army Ants in Western PanamaCONSERVATION BIOLOGY, Issue 1 2000Dina L. Roberts Behavioral and distributional studies of these two species have been confined largely to humid lowland forest. We conducted intensive systematic area searches at elevations between 1200 and 1800 m in western Panama to assess the distribution of both species in intact premontane moist forest, shade coffee plantations, and sun coffee plantations. Both species were repeatedly observed in forest, shade coffee plantations close to forest, and shade coffee plantations distant from forest. Neither species was observed in sun coffee plantations. We believe that retention of certain forest-like characteristics in the traditional shade coffee farm contributes to the persistence of these forest organisms in modified landscapes. Large canopy trees not only provide shade that buffers temperature extremes but also supply the ground layer with regular inputs of leaf litter and coarse woody debris from fallen trunks. Both E. burchelli and L. praedator hunt in leaf litter, and E. burchelli uses coarse woody debris as nesting sites ( bivouacs). There were significantly fewer potential bivouacs available in sun coffee plantations than in forest and shade coffee habitats. Also, litter depth was less in sun coffee than in forest and shade coffee. Our results provide the first evidence that shade coffee plantations can provide additional habitat for E. burchelli and L. praedator, top predators of the leaf litter arthropod community. E. burchelli and L. praedator act as critical links between swarm-attendant bird species and leaf-litter arthropods, providing an easily exploited food resource that would otherwise be unavailable for many birds. Continued conversion of shade coffee plantations to sun coffee plantations could have negative effects on army ants and associated biodiversity. Resumen: Las hormigas arrierras Neotropicales, Eciton burchelli y Labidus praedator ( Hymenoptera: Formicidae: Ecitoninae) son especies que requieren de extensas áreas de hábitat para cazar. Los estudios conductuales y de la distribución de estas especies se han realizado principalmente en bosques húmedos en tierras bajas. Desarrollamos búsquedas sistemáticas intensivas en elevaciones entre 1200 y 1800 msnm en Panama occidental para determinar la distribución de ambas especies en bosque húmedo premontano intacto, en plantaciones de café con y sin sombra. Las dos especies fueron observadas recurrentemente en bosque y en plantaciones de café de sombra cercanos y lejanos al bosque. Consideramos que la retención de ciertas características del bosque en las plantaciones de café de sombra contribuye a la persistencia de estos organismos de bosque en ambientes modificados. Los árboles no solo proporcionan sombra que amortigua la temperatura, sino que proporcionan hojarasca y restos leñosos de troncos caídos. Tanto E. burchelli como L. praedator cazan en la hojarasca, E. burchelli utiliza restos leñosos para anidar (vivaques). Encontramos significativamente menos vivaques en plantaciones de café sin sombra al compararlos con bosque y plantaciones de café con sombra. Asimismo, la profundidad de la capa de hojarasca fue menor en plantaciones de café sin sombra en comparación con bosque y plantaciones de café con sombra. Nuestros resultados proporcionan la primera evidencia de que las plantaciones con sombra proporcionan hábitat adicional para E. burchelli y L. praedator, depredadores de la comunidad de artrópodos en la hojarasca. E. burchelli y L. praedator actúan como eslabones críticos entre especies de aves que se alimentan de hormigas y los artrópodos de la hojarasca, proporcionando un recurso alimenticio fácilmente explotado que de otra manera no estaría disponible para muchas aves. La continua transformación de plantaciones de café con sombra a plantaciones sin sombra pudiera tener efectos negativos sobre las hormigas arrieras y la biodiversidad asociada. [source] Topical and systemic photoprotectionDERMATOLOGIC THERAPY, Issue 1 2003Cheryl F. Rosen ABSTRACT: Sunscreens are a valuable method of sun protection. Several new compounds are now available. It is important to remember, however, that photoprotection includes more than the use of sunscreens. There are a number of sun-protective behaviors that people can use to decrease their exposure to ultraviolet (UV) radiation. Dermatologists and other health professionals can work toward changing public policy, greatly increasing the ability of people to access shade. In addition, there is growing evidence about the effectiveness of other sun-protective agents. The only systemic medication for sun protection is ,-carotene, which is effective in erythropoietic protoporphyria (EPP). [source] Age-related change in canopy traits shifts conspecific facilitation to interference in a semi-arid shrublandECOGRAPHY, Issue 4 2007Orna Reisman-Berman Shifts between facilitation and interference and their importance in shaping plant population and community dynamics have received wide recognition. Nevertheless, the causes and spatio-temporal scales of these shifts are poorly understood, yet strongly debated. This study tested the hypothesis that age-related changes in canopy structure shift the effect of a nurse shrub on their protégé from facilitation to interference, using as a model system the interaction between the dwarf shrub Sarcopoterium spinosum and conspecific new recruits, in the shrubland of the transition area between the Mediterranean and the semi-arid climatic zones of Israel. Foliation level (i.e. the percentage of canopy surface area covered with leaves), a measure of shrub canopy structure, increased with age. Shading level was significantly and positively related to foliation level. Densities of new recruits in the shrubland showed a unimodal response to canopy structure and cover: the highest densities were associated with canopies presenting low and medium foliation levels (providing 71 and 82% shade, respectively), while high foliation levels (93% shade) and open spaces among canopies were characterized by very low densities. A related field experiment using shading nets revealed that seedling survival rates followed a similar unimodal pattern, with the highest survival (ca 60%) detected in moderate shade (70%), twice as much as in full sun, and the lowest survival (ca 10%) observed in extreme shade (90%). These results support the study hypothesis on age-dependent interactions. Thus, in a semi-arid shrubland ecosystem, the transition of the "nurse shrub" from "young" to "old" stage can shift facilitation to interference. Hence, the age structure of established shrub populations determines a) the availability of suitable sites for seedling recruitment and b) the balance between facilitation versus interference effects on seedling establishment. [source] Woody plants modulate the temporal dynamics of soil moisture in a semi-arid mesquite savanna,ECOHYDROLOGY, Issue 1 2010Daniel L. Potts Abstract Climate variability and human activities interact to increase the abundance of woody plants in arid and semi-arid ecosystems worldwide. How woody plants interact with rainfall to influence patterns of soil moisture through time, at different depths in the soil profile and between neighboring landscape patches is poorly known. In a semi-arid mesquite savanna, we deployed a paired array of sensors beneath a mesquite canopy and in an adjacent open area to measure volumetric soil water content (,) every 30 min at several depths between 2004 and 2007. In addition, to quantify temporally dynamic variation in soil moisture between the two microsites and across soil depths we analysed , time-series using fast Fourier transforms (FFT). FFT analyses were consistent with the prediction that by reducing evaporative losses through shade and reducing rainfall inputs through canopy interception of small rainfall events, the mesquite canopy was associated with a decline in high-frequency (hour-to-hour and day-to-day) variation in shallow ,. Finally, we found that, in both microsites, high-frequency , variation declined with increasing soil depth as the influence of evaporative losses and inputs associated with smaller rainfall events declined. In this case, we argue that the buffering of shallow soil moisture against high-frequency variations can enhance nutrient cycling and alter the carbon cycle in dryland ecosystems. Copyright © 2009 John Wiley & Sons, Ltd. [source] Best host-plant attribute for species,area relationship, and effects of shade, conspecific distance and plant phenophase in an arthropod community within the grass Muhlenbergia robustaENTOMOLOGICAL SCIENCE, Issue 2 2010Víctor LÓPEZ-GÓMEZ Abstract Increased understanding of the species,area relationship (SAR) can improve its usefulness as a tool for prediction of species loss for biodiversity conservation targets. This study was conducted: (i) to determine the best plant attribute for the SAR in the community of arthropods living within the grass Muhlenbergia robusta; (ii) to determine the contribution of phenophases of plant foliage (dry and fresh), shade and conspecific distance to the variation in arthropod richness within the plant; (iii) to determine the best functional model of changes in the abundance, diversity and biomass in communities of arthropods in response to increases in plant size; (iv) to determine the best host-plant attribute for prediction of these community attributes; and (v) to determine the effect of the plant phenophase, shade and M. robusta isolation on the abundance, diversity and biomass of the arthropod community. The above-ground dry weight of grass was found to be the best host-plant attribute for the SAR, while the light environment explained the arthropod richness within the grass, with higher richness observed in shaded environments. This study also showed that the best functional mathematical models for estimation of changes in the abundance, dry weight and diversity of arthropods in response to increases in grass size (dry weight) are the power model, exponential model and logarithmic model, respectively. Furthermore, the host-plant foliage phenophase, shade and the isolation of M. robusta with other conspecifics had no effect on the abundance, biomass or diversity per basal area of the grass. [source] THE CONTRIBUTION OF SPONTANEOUS MUTATION TO VARIATION IN ENVIRONMENTAL RESPONSES OF ARABIDOPSIS THALIANA: RESPONSES TO LIGHTEVOLUTION, Issue 2 2005Christina M. Kavanaugh Abstract It has been hypothesized that new, spontaneous mutations tend to reduce fitness more severely in more stressful environments. To address this hypothesis, we grew plants representing 20 Arabidopsis thaliana mutationaccumulation (M-A) lines, advanced to generation 17, and their progenitor, in differing light conditions. The experiment was conducted in a greenhouse, and two treatments were used: full sun and shade, in which influx of red light was reduced relative to far-red. The shade treatment was considered the more stressful because mean absolute fitness was lower in that treatment, though not significantly so. Plants from generation 17 of M-A developed significantly faster than those from generation 0 in both treatments. A significant interaction between generation and treatment revealed that, counter to the hypothesis, M-A lines tended to have higher fitness on average relative to the progenitor in the shaded conditions, whereas, in full sun, the two generations were similar in fitness. A secondary objective of this experiment was to characterize the contribution of new mutations to genotype x environment interaction. We did not, however, detect a significant interaction between M-A line and treatment. Plots of the line-specific enviromental responses indicate no tendency of new mutations to contribute to fitness trade-offs between environments. They also do not support a model of conditionally deleterious mutation, in which a mutatn reduces fitness only in a particular environment. These results suggest that interactions between genotype and light environment previously documented for A. thaliana are not explicable primarily as a consequence of steady input of spontaneous mutations having environment-specific effects. [source] PLASTICITY TO LIGHT CUES AND RESOURCES IN ARABIDOPSIS THALIANA: TESTING FOR ADAPTIVE VALUE AND COSTSEVOLUTION, Issue 6 2000Lisa A. Dorn Abstract Plants shaded by neighbors or overhead foliage experience both a reduction in the ratio of red to far red light (R:FR), a specific cue perceived by phytochrome, and reduced photosynthetically active radiation (PAR), an essential resource. We tested the adaptive value of plasticity to crowding and to the cue and resource components of foliage shade in the annual plant Arabidopsis thaliana by exposing 36 inbred families from four natural populations to four experimental treatments: (1) high density, full sun; (2) low density, full sun; (3) low density, neutral shade; and (4) low density, low R:FR-simulated foliage shade. Genotypic selection analysis within each treatment revealed strong environmental differences in selection on plastic life-history traits. We used specific contrasts to measure plasticity to density and foliage shade, to partition responses to foliage shade into phytochrome-mediated responses to the R:FR cue and responses to PAR, and to test whether plasticity was adaptive (i.e., in the same direction as selection in each environment). Contrary to expectation, we found no evidence for adaptive plasticity to density. However, we observed both adaptive and maladaptive responses to foliage shade. In general, phytochrome-mediated plasticity to the R:FR cue of foliage shade was adaptive and counteracted maladaptive growth responses to reduced PAR. These results support the prediction that active developmental responses to environmental cues are more likely to be adaptive than are passive resource-mediated responses. Multiple regression analysis detected a few costs of adaptive plasticity and adaptive homeostasis, but such costs were infrequent and their expression depended on the environment. Thus, costs of plasticity may occasionally constrain the evolution of adaptive responses to foliage shade in Arabidopsis, but this constraint may differ among environments and is far from ubiquitous. [source] Waterlogging and canopy interact to control species recruitment in floodplainsFUNCTIONAL ECOLOGY, Issue 4 2010Wiktor Kotowski Summary 1.,The extent to which seedling recruitment contributes to local functional diversity depends on the environmental filters operating in a plant community. Classical community assembly models assume that habitat constraints and competition act like hierarchical filters with habitat filtering as the dominant one. Alternative models assume a synergic interaction since responses to environmental stress and competition may impose physiological trade-offs in plants. 2.,River floodplains are an ideal system to test the relationship between habitat and competition filtering in community (re)assembly, as flooding causes changes in both habitat stress (waterlogging, resulting in anoxia and toxicity) and competition (dieback of vegetation) on one hand and acts as an effective seed dispersal vector on the other hand. 3.,We conducted a mesocosm experiment on early community assembly from a pool of 34 floodplain species covering a wetness gradient. Seed mixtures were sown in a full factorial design with water level, canopy and mowing as controlling factors. We measured the biomass of all species after one growing season and determined germination and seedling growth traits, both outside (response to waterlogging/no waterlogging) and in a growth-chamber (response to light/darkness). 4.,Species recruitment was analysed in relation to the controlling factors and measured functional traits using co-inertia analysis. Furthermore we analysed the effects of the controlling factors on several aspects of functional diversity. 5.,There was no establishment in grass sward, unless mowing was applied. Species-rich communities only developed when germination and early establishment phases occurred on waterlogged bare soil. High water level did not suppress establishment but reduced the total biomass and lowered inter-specific competition. The effect of mowing on species richness depended upon the interplay between waterlogging and canopy. 6.,Establishment success under canopy required seedling strategies to tolerate shade. The elimination of typical wetland specialists from oxic mesocosms was clearly an effect of their poorer and/or slower germination and lower competitive abilities in comparison to non-wetland plants, leading to their disappearance in this low-stress environment. 7.,Our results indicate that single stress factors can enhance species richness and functional diversity through limiting competition but a synergic interaction of different stresses can lead to reduced richness. [source] Xylem root and shoot hydraulics is linked to life history type in chaparral seedlingsFUNCTIONAL ECOLOGY, Issue 1 2010Robert B. Pratt Summary 1.,Shrubs in fire prone chaparral communities have evolved different life history types in response to fire. A key to understanding the evolution of life history type differences is to understand how physiological traits are linked to differences in life history type. Vascular adaptations are important for delivering an efficient and stable water supply to evergreen chaparral shrub leaves. This study tested for a link between vascular physiology and life history type in chaparral shrubs. 2.,Chaparral shrub species along the south-western coast of North America survive wildfire by three different life histories. Non-sprouters are killed by fire and re-establish exclusively through germination of fire-stimulated seeds, facultative sprouters re-establish by a combination of vegetative sprouting and fire-stimulated seeds, and obligate sprouters re-establish exclusively by vegetative sprouting because their seeds do not survive fire. Non-sprouters and facultative sprouters establish seedlings in the open canopy post fire environment, whereas obligate sprouters establish seedlings in the shady understory of the mature chaparral canopy. 3.,Seedlings of nine species (Rhamnaceae) representing three each of the different life history types were grown in deep containers in a common garden under treatments of sun and shade. Hydraulic conductance was measured using a high-pressure flow meter for all organs, and a vacuum technique was used to measure conductance of fine and woody roots. We predicted that non-sprouters would exhibit greater hydraulic efficiency than the sprouting species, and that facultative sprouters would be more efficient than the shade tolerant obligate sprouters. 4.,Non-sprouters had the greatest hydraulic conductance per unit leaf and sapwood area at the whole seedling level, whereas facultative and obligate sprouters were not different. Comparing hydraulic conductance across major organs (from fine roots to leaves) showed that the hydraulic system was well coordinated. At the whole seedling level, the root system was more of a bottleneck than the shoot system. This pattern was consistent with high resistance extraxylary pathways in roots and differences in root architecture. 5.,The greater hydraulic efficiency of the non-sprouter life history type is attributed to its post-fire pioneering habit and may partially explain the relatively high speciation in the non-sprouters. Lower hydraulic efficiency is associated with a sprouting life history and greater shade tolerance. The seedling root systems represent a hydraulic bottleneck that may place roots under especially intense selection. [source] Patterns and consequences of differential vascular sectoriality in 18 temperate tree and shrub speciesFUNCTIONAL ECOLOGY, Issue 2 2006A. E. ZANNE Summary 1Resource delivery within plants depends on supply pathways. Some species have relatively constrained (sectored) vascular connections, while others have relatively unconstrained (integrated) vascular connections. 2In this study, patterns of vascular hydraulic sectoriality, anatomy and ecological tolerance were examined for 18 Northern Hemisphere temperate woody species growing at Arnold Arboretum, Jamaica Plain, MA, USA. A hydraulic technique was used to measure axial and tangential conductivity on branch segments. From a ratio of these values, a sectoriality index was calculated. 3Species that were more hydraulically sectored had greater vessel size, variation in vessel area and tangential nearest-neighbour distance, as well as lower vessel density, than did integrated species. 4Ecologically, higher tolerance to drought and wind was correlated with being sectored, while higher tolerance to flood and shade was correlated with being integrated. 5These results suggest that sectored species should be especially prominent in xeric environments where sectoriality may reduce embolism spread by minimizing vessel-to-vessel contact and pitting, and integrated species should be especially prominent when resources are spatially patchy or heterogeneous. [source] Light gains and physiological capacity of understorey woody plants during phenological avoidance of canopy shadeFUNCTIONAL ECOLOGY, Issue 4 2005C. K. AUGSPURGER Summary 1Carbon gain during phenological avoidance of canopy shade by an understorey plant depends on the extent of avoidance, the leaf stage during avoidance, and whether young and old leaves can exploit greater light availability in spring and autumn. 2For Asimina triloba (L.) Dunal., Aesculus glabra Willd., Acer saccharum Marsh., Lindera benzoin (L.) Blume and Carpinus caroliniana Walt. in a deciduous forest in Illinois, USA, spring avoidance with leaves at full size ranged from 0 days for Asimina to 24 days for Aesculus, and brought 36,98% of estimated total annual irradiance. Autumn avoidance was non-existent to minimal in all species. 3Total chlorophyll reached maximum concentrations at the middle of leaf life span, and declined well before senescence. Leaf nitrogen concentrations and net photosynthetic capacity both peaked in youngest leaves during spring avoidance, and were low in old leaves during autumn avoidance. 4Aesculus had especially high photosynthetic capacity during precanopy closure, while Asimina had relatively low capacity in its later developing leaves. 5Young leaves of species with phenological avoidance can enhance C gain, while old leaves in autumn do not. Thus phenological avoidance in spring may enhance the persistence of understorey woody individuals of some species. [source] Allocation of above-ground growth is related to light in temperate deciduous saplingsFUNCTIONAL ECOLOGY, Issue 4 2003D. A. King Summary 1Allocational shifts in response to light may be an important factor in allowing plants to survive in shade, while increasing their extension rates and competitive ability in sun. To investigate this response, the allocation of above-ground growth between leaves, branches and stems was studied in saplings of Acer pensylvanicum L. and Castenea dentata (Marsh.) Borkh. in the Appalachian mountains of western Virginia, USA. Measurements of current leaf biomass, current and past year leaf numbers and the growth ring widths of branches and stem were used to estimate biomass partitioning for saplings growing in locations ranging from forest understorey to large openings. 2Both species showed higher leaf area per unit leaf biomass (SLA) and higher allocation of above-ground growth to leaves in shade than in sun. 3There were no differences between species in the slopes of the relationships of allocation and SLA vs estimated irradiance, but SLA was significantly greater in A. pensylvanicum than in C. dentata at a given light level. Hence, somewhat lower production per unit leaf area is required to maintain the canopy in A. pensylvanicum, consistent with foresters' ratings of greater shade tolerance for this species. 4Greater foliar allocation in shade than sun has also been observed in broad-leaved evergreen saplings, but generally not in seedlings. This difference is probably related to differences in size and age between seedlings and saplings. Young seedlings typically show exponential growth with no immediate foliar losses, while shaded saplings lie closer to the steady state where new leaves replace old ones with little additional stem growth. 5Thus trees shift their allocation patterns in an acclimatory fashion, depending on their size and light environment, with the costs of replacing senesced leaves becoming of consequence as juveniles age. [source] Do shade-tolerant tropical tree seedlings depend longer on seed reserves?FUNCTIONAL ECOLOGY, Issue 4 2002Functional growth analysis of three Bignoniaceae species Summary 1.,A functional growth analysis was used to determine the duration of strict dependency on seed reserves for energy and nitrogen in three woody Bignoniaceae species (Tabebuia rosea DC., Challichlamys latifolia K. Schum. and Pithecoctenium crucigerum A. Gentry) which differed in cotyledon function (photosynthetic, semi-photosynthetic and storage) and shade tolerance (probability of seedling establishment and survival in the understorey). 2.,Seedlings were raised from seeds in sand culture under combinations of three nitrogen levels (daily supply of nutrient solution containing 100, 10 and 0% of 2·6 mm N) and two irradiances (27 and 1% full sun). Time course of biomass, non-cotyledonous biomass and leaf area for 40 days post-germination were compared to identify when the external availability of nitrogen or light began to affect seedling growth. 3.,Seedlings of all species became dependent on external energy supply earlier than they did on nitrogen supply. In all species seed nitrogen was sufficient to support positive seedling growth for 40 days in shade, but not in sun. 4.,Tabebuia rosea with photosynthetic cotyledons responded to light availability earlier than more shade-tolerant species with storage cotyledons. Challichlamys latifolia, the most shade-tolerant species, had the highest nitrogen concentration in seeds and was the last to respond to external nitrogen availability. Thus seedlings of the most shade-tolerant species depended on seed reserves for the longest period for both energy and nitrogen. 5.,Relative growth rate after seedlings initiated autotrophic growth was in a trade-off relationship with seedling survivorship in the understorey across the three species. Tabebuia rosea, the least shade-tolerant species, had the highest positive net carbon balance in sun and shade. 6.,Functional morphology of cotyledons and concentration of seed nitrogen deserve as much attention as seed size as correlates of contrasting seedling regeneration strategies. [source] Ecophysiological controls over the net ecosystem exchange of mountain spruce stand.GLOBAL CHANGE BIOLOGY, Issue 1 2007Comparison of the response in direct vs. diffuse solar radiation Abstract Cloud cover increases the proportion of diffuse radiation reaching the Earth's surface and affects many microclimatic factors such as temperature, vapour pressure deficit and precipitation. We compared the relative efficiencies of canopy photosynthesis to diffuse and direct photosynthetic photon flux density (PPFD) for a Norway spruce forest (25-year-old, leaf area index 11 m2 m,2) during two successive 7-day periods in August. The comparison was based on the response of net ecosystem exchange (NEE) of CO2 to PPFD. NEE and stomatal conductance at the canopy level (Gcanopy) was estimated from half-hourly eddy-covariance measurements of CO2 and H2O fluxes. In addition, daily courses of CO2 assimilation rate (AN) and stomatal conductance (Gs) at shoot level were measured using a gas-exchange technique applied to branches of trees. The extent of spectral changes in incident solar radiation was assessed using a spectroradiometer. We found significantly higher NEE (up to 150%) during the cloudy periods compared with the sunny periods at corresponding PPFDs. Prevailing diffuse radiation under the cloudy days resulted in a significantly lower compensation irradiance (by ca. 50% and 70%), while apparent quantum yield was slightly higher (by ca. 7%) at canopy level and significantly higher (by ca. 530%) in sun-acclimated shoots. The main reasons for these differences appear to be (1) more favourable microclimatic conditions during cloudy periods, (2) stimulation of photochemical reactions and stomatal opening via an increase of blue/red light ratio, and (3) increased penetration of light into the canopy and thus a more equitable distribution of light between leaves. Our analyses identified the most important reason of enhanced NEE under cloudy sky conditions to be the effective penetration of diffuse radiation to lower depths of the canopy. This subsequently led to the significantly higher solar equivalent leaf area compared with the direct radiation. Most of the leaves in such dense canopy are in deep shade, with marginal or negative carbon balances during sunny days. These findings show that the energy of diffuse, compared with direct, solar radiation is used more efficiently in assimilation processes at both leaf and canopy levels. [source] Evaluation of soil saturation, soil chemistry, and early spring soil and air temperatures as risk factors in yellow-cedar declineGLOBAL CHANGE BIOLOGY, Issue 3 2006D. V. D'AMORE Abstract Yellow-cedar (Callitropsis nootkatensis (D. Don) Oerst.) is a valuable tree species that is experiencing a widespread decline and mortality in southeast Alaska. This study evaluated the relative importance of several potential risk factors associated with yellow-cedar decline: soil saturation, soil aluminum (Al) toxicity or calcium (Ca) deficiency, and air and soil temperature. Data were collected from permanent vegetation plots established in two low-elevation coastal forests exhibiting broad ranges of cedar mortality. Measurements of each risk factor were contrasted among classified forest zones to indicate if there were strong links with decline. Hydrology alone is weakly associated with yellow-cedar decline, but could have a predisposing role in the decline by creating exposed conditions because of reduced forest productivity. Yellow-cedar decline is not strongly associated with soil pH and extractable Al and Ca, but there appears to be Ca enrichment of surface soils by feedback from dead yellow-cedar foliage. Air and soil temperature factors are strongly associated with decline. Based on these results, an hypothesis is presented to explain the mechanism of tree injury where exposure-driven tree mortality is initiated in gaps created by soil saturation and then expands in gaps created by the tree-mortality itself. The exposure allows soils to warm in early spring causing premature dehardening in yellow-cedar trees and subsequent freezing injury during cold events. Yellow-cedars growing in the protection of shade or snow are not preconditioned by this warming, and thus not as susceptible to cold injury. Yellow-cedar decline appears to be associated with regional climate changes, but whether the cause of these changes is related to natural or human-induced climate shifts remains uncertain. Management implications, the possible role of climate, and recommended research are discussed. [source] Abiotic constraints on the establishment of Quercus seedlings in grasslandGLOBAL CHANGE BIOLOGY, Issue 2 2003Brett T. Danner Abstract High evaporative demand and periodic drought characterize the growing season in midwestern grasslands relative to deciduous forests of the eastern US, and predicted climatic changes suggest that these climatic extremes may be exacerbated. Despite this less than optimal environment for tree seedling establishment, deciduous trees have expanded into adjacent tallgrass prairie within the last century leading to a dramatic land cover change. In order to determine the role of light and temperature on seedling establishment, we assessed carbon and water relations and aboveground growth of first-year Quercus macrocarpa seedlings exposed to one of three conditions: (1) intact tallgrass prairie communities (control), (2) aboveground herbaceous biomass removed (grass removal), and (3) shade plus biomass removal to reduce light (PFD) to levels typical of the grassland-forest ecotone (shade). In the 2000 growing season, precipitation was 35% below the long-term average, which had a significant negative effect on oak seedling carbon gain at midseason (photosynthesis declined to 10% of maximum rates). However, net photosynthesis and stomatal conductance in the shade treatment was ca. 2.5 and 1.5 times greater, respectively, than in control treatment seedlings during this drought. During this period, leaf and air temperatures in control seedlings were similar whereas leaf temperatures in the shade treatment remained below air temperature. A late-season recovery period, coincident with decreased air temperatures, resulted in increased net photosynthesis for all seedlings. Increased photosynthetic rates and water relations in shaded seedlings compared to seedlings in full sun suggest that, at least in dry years, high light and temperature may negatively impact oak seedling performance. However, high survival rates for all seedlings indicate that Q. macrocarpa seedlings are capable of tolerating both present-day and future climatic extremes. Unless historic fire regimes are restored, forest expansion and land cover change are likely to continue. [source] Halloween genes and nuclear receptors in ecdysteroid biosynthesis and signalling in the pea aphidINSECT MOLECULAR BIOLOGY, Issue 2010O. Christiaens Abstract The pea aphid (Acyrthosiphon pisum) is the first whole genome sequenced insect with a hemimetabolic development and an emerging model organism for studies in ecology, evolution and development. The insect steroid moulting hormone 20-hydroxyecdysone (20E) controls and coordinates development in insects, especially the moulting/metamorphosis process. We, therefore present here a comprehensive characterization of the Halloween genes phantom, disembodied, shadow, shade, spook and spookiest, coding for the P450 enzymes that control the biosynthesis of 20E. Regarding the presence of nuclear receptors in the pea aphid genome, we found 19 genes, representing all of the seven known subfamilies. The annotation and phylogenetic analysis revealed a strong conservation in the class of Insecta. But compared with other sequenced insect genomes, three orthologues are missing in the Acyrthosiphon genome, namely HR96, PNR-like and Knirps. We also cloned the EcR, Usp, E75 and HR3. Finally, 3D-modelling of the ligand-binding domain of Ap-EcR exhibited the typical canonical structural scaffold with 12 ,-helices associated with a short hairpin of two antiparallel ,-strands. Upon docking, 20E was located in the hormone-binding groove, supporting the hypothesis that EcR has a role in 20E signalling. [source] | |||||||||||||||||||||||||||||||||||||||||||||||||