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Annual Cycle (annual + cycle)
Selected AbstractsAnnual Cycle of Planktothrix agardhii(Gom.) Anagn.INTERNATIONAL REVIEW OF HYDROBIOLOGY, Issue 3 2004& Kom. Abstract Changes in abundance, biovolume and morphology of Planktothrix agardhii in a natural population were followed over one year period in shallow fishpond Bílá Lhota (Central Moravia, Czech Republic). The selected environmental parameters (pH, oxygen, temperature, conductivity, nutrients, light) were measured at the surface and at the bottom of the fishpond, together with the Planktothrix abundances and filament morphology , filament length, width, shape, aerotopes (gas vacuoles) formation. The annual cycle of P. agardhii in this hypertrophic fishpond starts in March with the germination of hormogonia and the growth of overwintered filaments. The filament length quickly increases to a maximum length in April. The following summer period can be characterized by filament shortening and by changes in the aerotopes shape. On the other hand the abundance and biomass of P. agardhii is increasing until the maximum in August. Further shortening of filaments, loss of aerotopes and hormogoniae formation is typical for the autumn (October) with the average temperature of 9.4 °C. The population overwinters near the pond bottom in the form of hormogonia (60%) and filaments (40%). (© 2004 WILEY-VCH Verlag GmbH & Co. KGaA, Weinheim) [source] Motility of Daphnia spinulata as Affected by Solar Radiation Throughout an Annual Cycle in Mid-latitudes of Patagonia,PHOTOCHEMISTRY & PHOTOBIOLOGY, Issue 4 2007Rodrigo J. Gonçalves During an annual cycle, Daphnia spinulata collected from a plankton community of Patagonia was exposed outdoors to assess the impact of recently received solar radiation on motility (i.e. swimming speed and gravitaxis). Individual values of these parameters were obtained by video recordings and image analysis at different time intervals during the day. Initial swimming speed varied throughout the year, and changes in speed during exposure were not significantly affected by any waveband used in our experimental design (i.e. PAB, 280,700 nm; PA, 320,700 nm; and P, 400,700 nm). Overall, most of the individuals swam downwards, regardless of the radiation treatment imposed to the samples. We found that multifactor interactions (i.e. not a single parameter explained more than 40% of the observed variability) explained most of our observations on motility parameters. These factors include not only solar radiation, but other physical (underwater radiation field and wind intensity) and biological parameters (food availability, presence of predators and congeners). Our findings indicate that the plankton dynamics in the study site is likely to be governed by a sum of factors which must be taken into account when considering solar radiation effects on aquatic ecosystems. [source] Annual cycle of clupeiform larvae around Gran Canaria Island, Canary IslandsFISHERIES OCEANOGRAPHY, Issue 4 2006P. BÉCOGNÉE Abstract The distribution and abundance of fish larvae was studied along the eastern and southern shelf of Gran Canaria Island (Canary Islands) from July 2000 to June 2001. Oblique bongo hauls were carried out fortnightly during the daytime, coinciding with days of full and new moon. During February, the area was sampled every 2,5 days. About 17.3% of the ichthyoplanktonic community was composed of clupeiform larvae: 92.9% of these larvae were Sardinella aurita, whereas 4.7% and 2.4% were respectively Engraulis encrasicolus and Sardina pilchardus. Sardinella aurita larvae appeared during the whole year with two periods of maximum abundance: June to September and December to February. During the full moon their abundance was on average 38.5% (±6.8%) of their numbers during the new moon, showing a clear lunar periodicity. Engraulis encrasicolus larvae appeared from November to March, also coinciding with the new moon. Sardina pilchardus larvae only appeared during two short periods, both coinciding with filaments shed from the African coastal upwelling which reached the island. This fact confirms the transport of fish larvae from the upwelling area off northwest Africa to the Canary Islands, promoting a genetic flow among both sites. [source] Annual cycle and inter-annual variability of gross primary production and ecosystem respiration in a floodprone river during a 15-year periodFRESHWATER BIOLOGY, Issue 5 2006URS UEHLINGER Summary 1. Temporal variation in ecosystem metabolism over a 15-year period (1986,2000) was evaluated in a seventh order channelised gravel bed river (mean annual discharge 48.7 m3 s,1) of the Swiss Plateau. The river is subject to frequent disturbance by bed-moving spates. Daily integrals of gross primary production (GPP) and ecosystem respiration (ER) were calculated based on single-station diel oxygen curves. 2. Seasonal decomposition of the time series of monthly metabolism rates showed that approximately 50% of the variation of GPP and ER can be attributed to season. Annual GPP averaged 5.0 ± 0.6 g O2 m,2 day,1 and showed no long-term trend. 3. Ecosystem respiration, averaging 6.2 ± 1.4 g O2 m,2 day,1, declined from 8.8 to 4.1 g O2 m,2 day,1 during the 15-year period. This significant trend paralleled a decline in nitrate and soluble reactive phosphorus concentrations, and the biochemical oxygen demand discharged by sewage treatment facilities upstream of the study reach. The ratio of GPP to ER (P/R) increased from 0.53 to about 1 as consequence of ER reduction. 4. Bed moving spates reduced GPP by 49% and ER by 19%. Postspate recovery of GPP was rapid between spring and autumn and slow during winter. Recovery of ER lacked any seasonal pattern. Annual patterns of daily GPP and to a minor extent of daily ER can be described as a sequence of recovery periods frequently truncated by spates. 5. The study showed that disturbance by frequent bed-moving spates resulted in major stochastic variation in GPP and ER but annual patterns were still characterised by a distinct seasonal cycle. It also became evident that stream metabolism is a suitable method to assess effects of gradual changes in water quality. [source] Annual cycles in the interstellar scintillation time-scales of PKS B1519,273 and PKS B1622,253MONTHLY NOTICES OF THE ROYAL ASTRONOMICAL SOCIETY, Issue 3 2009Steven J. B. Carter ABSTRACT We have used the University of Tasmania's 30-m radio telescope at Ceduna in South Australia to regularly monitor the flux density of a number of southern blazars. We report the detection of an annual cycle in the variability time-scale of the centimetre radio emission of PKS B1622,253. Observations of PKS B1519,273 over a period of nearly 2 yr confirm the presence of an annual cycle in the variability time-scale in that source. These observations prove that interstellar scintillation is the principal cause of inter-day variability at radio wavelengths in these sources. The best-fitting annual cycle model for both sources implies a high degree of anisotropy in the scattering screen and that it has a large velocity offset with respect to the local standard of rest. This is consistent with a greater screen distance for these ,slow' intra-day variability (IDV) sources than for rapid scintillators such as PKS B0405,385 or J1819+3845. [source] Optimal conservation planning for migratory animals: integrating demographic information across seasonsCONSERVATION LETTERS, Issue 3 2010Justin Sheehy Abstract Conservation strategies for migratory animals are typically based on ad-hoc or simple ranking methods and focus on a single period of the annual cycle. We use a density-dependent population model to examine one-time land purchase strategies for a migratory population with a breeding and wintering grounds. Under equal rates of habitat loss, we show that it is optimal to invest more, but never solely, in the habitat with the higher density dependence to habitat cost ratio. When there are two habitats that vary in quality within a season, the best strategy is to invest only in one habitat. Whether to purchase high- or low-quality habitat depends on the general life history of the species and the ratio of habitat quality to habitat cost. When carry-over effects are incorporated, it is almost always optimal to invest in high-quality habitat during the season that produces the carry-over effect. We apply this model to a threatened warbler population and show the optimal strategy is to purchase more breeding than wintering habitat despite the fact that breeding habitat is over ten times more expensive. Our model provides a framework for developing year-round conservation strategies for migratory animals and has important implications for long-term planning and management. [source] Impact of freshwater input and wind on landings of anchovy (Engraulis encrasicolus) and sardine (Sardina pilchardus) in shelf waters surrounding the Ebre (Ebro) River delta (north-western Mediterranean)FISHERIES OCEANOGRAPHY, Issue 2 2004J. Lloret Abstract Time series analyses (Box,Jenkins models) were used to study the influence of river runoff and wind mixing index on the productivity of the two most abundant species of small pelagic fish exploited in waters surrounding the Ebre (Ebro) River continental shelf (north-western Mediterranean): anchovy (Engraulis encrasicolus) and sardine (Sardina pilchardus). River flow and wind were selected because they are known to enhance fertilization and local planktonic production, thus being crucial for the survival of fish larvae. Time series of the two environmental variables and landings of the two species were analysed to extract the trend and seasonality. All series displayed important seasonal and interannual fluctuations. In the long term, landings of anchovy declined while those of sardine increased. At the seasonal scale, landings of anchovy peaked during spring/summer while those of sardine peaked during spring and autumn. Seasonality in landings of anchovy was stronger than in sardine. Concerning the environmental series, monthly average Ebre runoff showed a progressive decline from 1960 until the late 1980s, and the wind mixing index was highest during 1994,96. Within the annual cycle, the minimum river flow occurs from July to October and the wind mixing peaks in winter (December,April, excluding January). The results of the analyses showed a significant correlation between monthly landings of anchovy and freshwater input of the Ebre River during the spawning season of this species (April,August), with a time lag of 12 months. In contrast, monthly landings of sardine were significantly positively correlated with the wind mixing index during the spawning season of this species (November,March), with a lag of 18 months. The results provide evidence of the influence of riverine inputs and wind mixing on the productivity of small pelagic fish in the north-western Mediterranean. The time lags obtained in the relationships stress the importance of river runoff and wind mixing for the early stages of anchovy and sardine, respectively, and their impact on recruitment. [source] Near,surface ground temperature regime variability in selected microenvironments, Kärkevagge, Swedish LaplandGEOGRAFISKA ANNALER SERIES A: PHYSICAL GEOGRAPHY, Issue 3-4 2002Colin E. Thorn The importance of topographic microvariability in influencing shallow (10,50 cm depths) soil temperature regimes in arctic,alpine Kärkevagge, northern Sweden, from August 1999 to July 2000 is demonstrated using six sites. The ground microclimate on the tops of very large boulders forming an extensive boulder field in the central valley bottom is more comparable to that at an alpine ridge,crest site 300 m higher than it is to the microclimate at the base of one of the boulders. The boulder crests also differ substantially from the more generalized valley,bottom conditions outside the boulder field. Assuming that chemical processes may be active at temperatures at or above 0°C, sites in the valley experience favorable conditions from 159 to 324 days of the year. Aside from the annual cycle, freeze,thaw cycles are infrequent within Kärkevagge. [source] Turf exfoliation in the high Drakensberg, Southern AfricaGEOGRAFISKA ANNALER SERIES A: PHYSICAL GEOGRAPHY, Issue 1 2002Stefan W. Grab Limited research attention has focussed on turf exfoliation as a denudation process in mountain environments. This paper examines some characteristics of turf exfoliation forms identified within particular valley zones in the Drakensberg alpine belt. Morphological and sedimentological data are presented for turf exfoliated sites investigated in the Mashai Valley of eastern Lesotho. It is found that a variety of processes, including needle ice action, biological activity, fluvial processes and deflation, operating synergistically, are responsible for contemporary turf exfoliation in the high Drakensberg. It is apparent that the strong seasonality from mild, wet summers to cold, dry winters has helped induce the annual cycle of dominating processes. [source] Climatic effects on the phenology of lake processesGLOBAL CHANGE BIOLOGY, Issue 11 2004Monika Winder Abstract Populations living in seasonal environments are exposed to systematic changes in physical conditions that restrict the growth and reproduction of many species to only a short time window of the annual cycle. Several studies have shown that climate changes over the latter part of the 20th century affected the phenology and population dynamics of single species. However, the key limitation to forecasting the effects of changing climate on ecosystems lies in understanding how it will affect interactions among species. We investigated the effects of climatic and biotic drivers on physical and biological lake processes, using a historical dataset of 40 years from Lake Washington, USA, and dynamic time-series models to explain changes in the phenological patterns among physical and biological components of pelagic ecosystems. Long-term climate warming and variability because of large-scale climatic patterns like Pacific decadal oscillation (PDO) and El Niño,southern oscillation (ENSO) extended the duration of the stratification period by 25 days over the last 40 years. This change was due mainly to earlier spring stratification (16 days) and less to later stratification termination in fall (9 days). The phytoplankton spring bloom advanced roughly in parallel to stratification onset and in 2002 it occurred about 19 days earlier than it did in 1962, indicating the tight connection of spring phytoplankton growth to turbulent conditions. In contrast, the timing of the clear-water phase showed high variability and was mainly driven by biotic factors. Among the zooplankton species, the timing of spring peaks in the rotifer Keratella advanced strongly, whereas Leptodiaptomus and Daphnia showed slight or no changes. These changes have generated a growing time lag between the spring phytoplankton peak and zooplankton peak, which can be especially critical for the cladoceran Daphnia. Water temperature, PDO, and food availability affected the timing of the spring peak in zooplankton. Overall, the impact of PDO on the phenological processes were stronger compared with ENSO. Our results highlight that climate affects physical and biological processes differently, which can interrupt energy flow among trophic levels, making ecosystem responses to climate change difficult to forecast. [source] Spatial and temporal variability of the phenological seasons in Germany from 1951 to 1996GLOBAL CHANGE BIOLOGY, Issue 6 2001Annette Menzel Abstract Various indications for shifts in plant and animal phenology resulting from climate change have been observed in Europe. This analysis of phenological seasons in Germany of more than four decades (1951,96) has several major advantages: (i) a wide and dense geographical coverage of data from the phenological network of the German Weather Service, (ii) the 16 phenophases analysed cover the whole annual cycle and, moreover, give a direct estimate of the length of the growing season for four deciduous tree species. After intensive data quality checks, two different methods ,,linear trend analyses and comparison of averages of subintervals , were applied in order to determine shifts in phenological seasons in the last 46 years. Results from both methods were similar and reveal a strong seasonal variation. There are clear advances in the key indicators of earliest and early spring (,0.18 to ,0.23 d y,1) and notable advances in the succeeding spring phenophases such as leaf unfolding of deciduous trees (,0.16 to ,0.08 d y,1). However, phenological changes are less strong during autumn (delayed by +,0.03 to +,0.10 d y,1 on average). In general, the growing season has been lengthened by up to ,0.2 d y,1 (mean linear trends) and the mean 1974,96 growing season was up to 5 days longer than in the 1951,73 period. The spatial variability of trends was analysed by statistical means and shown in maps, but these did not reveal any substantial regional differences. Although there is a high spatial variability, trends of phenological phases at single locations are mirrored by subsequent phases, but they are not necessarily identical. Results for changes in the biosphere with such a high resolution with respect to time and space can rarely be obtained by other methods such as analyses of satellite data. [source] The surface radiation budget over North America: gridded data assessment and evaluation of regional climate modelsINTERNATIONAL JOURNAL OF CLIMATOLOGY, Issue 15 2009Marko Markovic Abstract While surface station observations of downwelling radiation offer accuracy at high temporal resolution, they do not easily allow an evaluation of model surface radiation budgets (SRB) over a wide geographical area. We evaluate three gridded SRB data sets against detailed observations from six surface radiation sites from the US surface radiation (SURFRAD) network. We subsequently use the most accurate surrogate observational data set for evaluation of model-simulated SRB. The data sets assessed are: ERA40,reanalysis of European Centre for Medium-Range Weather Forecasts (ECMWF), North American Regional Reanalysis (NARR),regional reanalysis of National Centres for Environmental Prediction (NCEP) and the surface radiative budget (SRB) from the International Satellite Cloud Climatology Project (ISCCP). Due to varying constraints with respect to temporal coverage of each data set, the evaluation period used in this study is 1996,2001, inclusive. The ERA40 downwelling longwave radiation (DLR) appears the most accurate surrogate observation, while both ERA40 and ISCCP show accurate results when the incoming shortwave radiation (ISR) is considered across the annual cycle. Winter DLR is less accurate in ISCCP with a positive bias and lack of very low (<200 Wm,2) flux values. The NARR SRB shows a large positive bias in the ISR throughout the annual cycle, linked to a significant underestimate of cloud cover. The ERA40 data are subsequently used to evaluate the simulated SRB in three regional climate models across North America. With respect to solar radiation, cloud cover biases are seen to be crucial, while for longwave fluxes both cloud fraction and in-cloud water content are important to simulate correctly. Inclusion of trace gases beyond H2O, CO2 and O3 appears necessary for an accurate calculation of clear-sky longwave radiation. Error compensation frequently occurs between the various components contributing to a model total-sky SRB. This is important to consider when trying to identify the underlying causes of errors in the simulated total SRB. Copyright © 2009 Royal Meteorological Society [source] A regional climate study of Central America using the MM5 modeling system: results and comparison to observationsINTERNATIONAL JOURNAL OF CLIMATOLOGY, Issue 15 2006Jose L. Hernandez Abstract The Mesoscale Modeling system, version 3.6 (MM5) regional modeling system has been applied to Central America and has been evaluated against National Oceanic and Atmospheric Administration/National Climatic Data Center (NOAA/NCDC) daily observations and the Global Precipitation Climatology Project (GPCP) precipitation data. We compare model results and observations for 1997 and evaluate various climate parameters (temperature, wind speed, precipitation and water vapor mixing ratio), emphasizing the differences within the context of the station dependent geographical features and the land use (LU) categories. At 9 of the 16 analyzed stations the modeled temperature, wind speed and vapor mixing ratio are in agreement with observations with average model-observation differences consistently lower than 25%. MM5 has better performance at stations strongly impacted by monsoon systems, regions typified by low topography in coastal areas and areas characterized by evergreen, broad-leaf and shrub land vegetation types. At four stations the model precipitation is about a factor of 3,5 higher than the observations, while the simulated wind is roughly twice what is observed. These stations include two inland stations characterized by croplands close to water bodies; one coastal station in El Salvador adjacent to a mountain-based cropland area and one station at sea-level. This suggests that the model does not adequately represent the influence of topographic features and water bodies close to these stations. In general, the model agrees reasonably well with measurements and therefore provides an acceptable description of regional climate. The simulations in this study use only two seasonal maps of land cover. The main model discrepancies are likely attributable to the actual annual cycle of land,atmosphere vapor and energy exchange that has a temporal scale of days to weeks. These fluxes are impacted by surface moisture availability, albedo and thermal inertia parameters. Copyright © 2006 Royal Meteorological Society. [source] An event-based jet-stream climatology and typologyINTERNATIONAL JOURNAL OF CLIMATOLOGY, Issue 3 2006Patrick Koch Abstract A novel climatology is developed for upper-tropospheric jet streams, which is complementary to and an alternative for the traditional depictions of the time-mean jets. It entails identifying the occurrence of a jet event at a given location and then compiling the spatial frequency distribution of such events. The resulting climatology, derived using the ERA-15 reanalysis data set of the ECMWF for the period 1979,1993 indicates that (1) in both hemispheres the annual cycle of jet events takes the form of comparatively smooth transition from a quasi-annular structure in summer to a more spiral-like structure in winter with a temporally asymmetric return to the summer pattern; (2) the hemispheres differ primarily in the amplitude of the frequencies and the longitudinal overlap of the spiral portion of the pattern. In addition, the jet events are subdivided using a two-class typology comprising shallow and deep jets whose vertical shear (sic. baroclinicity) are/are not confined principally to the upper troposphere. This provides a conceptually simple and dynamically meaningful classification since deep jets are more likely to spawn tropospheric-spanning cyclones. The accompanying climatology displays important longitudinal variations and significant inter-hemispheric differences. A comparison is drawn between these new and conventional climatologies and typologies. Also, comments are proffered on the relationship between, on the one hand, the patterns of jet frequency including the differing distributions of the shallow and deep types and, on the other hand, the location of the time-mean jets and the downstream storm tracks. Copyright © 2006 Royal Meteorological Society. [source] The surface winds of Sweden during 1999,2000INTERNATIONAL JOURNAL OF CLIMATOLOGY, Issue 2 2006Christine Achberger Abstract This study aims at increasing our understanding of the regional wind climate in Sweden. Spatial and temporal patterns of the surface winds are presented for the years 1999,2000. Annual mean wind speeds range between 2 and 5 m/s with high values at exposed mountainous sites and on islands off the coast. Combining wind speed and direction into mean wind velocities shows that flow conditions are stronger and more coherent in space in southern Sweden than in central and northern Sweden. The spatial scale, defined as the distance between stations when the correlation for wind speed drops to ,0.37, was determined by pairwise correlations between all possible station pairs. Scales range from 38 to 530 km for wind speed and from 40 to 830 km for wind direction depending on the region. They tend to be smaller in central and northern Sweden, where the more pronounced relief has a larger influence on the local wind conditions. The strength and the timing of the annual and diurnal wind speed cycle have been estimated for each station. Amplitudes of the annual cycle are greater at exposed sites and correlate generally well with annual mean wind speeds. Monthly mean wind speeds peak in winter in southern Sweden, but peak in other seasons in the remaining regions. In winter, weaker pressure gradients over northern Sweden and surface-near temperature inversions contribute to weaker surface winds. Diurnal cycles vary in strength between summer and winter. Compared to the last normal climate period (1961,1990), 1999,2000 is characterized by the increased occurrence of westerly and southerly geostrophic flow. Copyright © 2005 Royal Meteorological Society. [source] Onset characteristics of the southwest monsoon over IndiaINTERNATIONAL JOURNAL OF CLIMATOLOGY, Issue 2 2005P. V. S. Raju Abstract Dynamic and thermodynamic characteristics of the Asian summer monsoon during the onset phase over the Indian Peninsula (Kerala coast) and its variability are examined with reanalysis data sets. For this study, daily averaged (0000 and 1200 UTC) reanalysis data sets of National Centre for Environmental Prediction,National Centre for Atmospheric Research for the period 1948,99 are used. Based on 52 years of onset dates of the Indian summer monsoon, we categorized pre-onset, onset and post-onset periods (each averaged 5 days) to investigate the mean circulation characteristics and the large-scale energetics of the Asian summer monsoon. It is found that the strength of the low-level Somali jet and upper tropospheric tropical easterly jet increase rapidly during the time of evolution of the summer monsoon over India. Over the Bay of Bengal and the Arabian Sea, predominant changes are noticed in the large-scale balances of kinetic energy, heat and moisture from the pre-onset to the post-onset periods. Prior to the onset of the summer monsoon over India, a zone of flux convergence of heat and moisture is noticed over the eastern sector of the Bay of Bengal and this intensifies in the onset and post-onset periods. During onset of the monsoon over India, the horizontal flux convergence of heat and moisture, as well as diabatic heating, are enhanced over the Arabian Sea. These subsequently increase with the evolution and advancement of the monsoon over India. Further, the dynamics of the evolution processes (15 days before and 30 days after the onset date of the monsoon over Kerala for each annual cycle) are studied over various sectors, such as the Arabian Sea, Bay of Bengal and Indian Peninsula region. The study reveals that the low-level kinetic energy, vertically integrated generation of kinetic energy and net tropospheric moisture over Arabian Sea can be used as potential predictors for the prediction of the possible onset date of the summer monsoon over the Indian Peninsula. Copyright © 2005 Royal Meteorological Society [source] Seasonal to interannual variations of soil moisture measured in OklahomaINTERNATIONAL JOURNAL OF CLIMATOLOGY, Issue 15 2004Bradley G. Illston Abstract Agriculture is a $2 billion component of the state economy in Oklahoma. As a result, meteorological, climatological, and agricultural communities should benefit from an improved understanding of soil moisture conditions and how those conditions vary spatially and temporally. The Oklahoma Mesonet is an automated observing network that provides real-time hydrometeorological observations at 115 stations across Oklahoma. In 1996, sensors were installed at 60 Mesonet sites to provide near-real-time observations of soil moisture. This study focuses on 6 years of soil moisture data collected between 1997 and 2002 to analyse the annual cycle and temporal characteristics of soil moisture across Oklahoma. The statewide analysis of the annual cycle of soil moisture revealed four distinct soil moisture phases. In addition, the four statewide phases were also observed in each of the nine climate divisions across Oklahoma, although the temporal characteristics of each phase were unique for each division. Further analysis demonstrated that, at shallow soil depths (5 and 25 cm), the spatial variability of soil moisture across Oklahoma was most homogeneous during the winter and spring periods and most heterogeneous during the summer and autumn periods. Conversely, at greater depths (60 and 75 cm), soil moisture was most heterogeneous during the winter period and the most homogeneous during the late spring. Copyright © 2004 Royal Meteorological Society [source] Intra-seasonal variability of wintertime temperature over East AsiaINTERNATIONAL JOURNAL OF CLIMATOLOGY, Issue 2 2004Dao-Yi Gong Abstract There has been a profound warming over East Asia during the winter months (November through to March) over the past few decades. The goal of this study is to address the question of whether the daily temperature has become more variable in conjunction with this warming by using observed temperature data obtained from 155 Chinese and Korean stations. Prior to the analysis, the annual cycle is removed to obtain daily temperature anomalies for each winter for each station. Results show that the intra-seasonal variance generally decreases, implying that the daily temperatures are becoming less variable. Considering all stations as a whole, the rate of change is ,0.49°C2 per decade (equivalent to ,3.59% per decade). The changes are more robust in the northeastern portion of China. In contrast, there are no dominant trends for the skewness coefficients, except for clear negatively skewed trends in northeastern China. These results are consistent with an increase in the number of extremely cold events. Over the region, the frequency of low-temperature extremes (as low as below minus two standard deviations) increases at a rate of change of 0.26 days per decade, significant at the 95% confidence level. Both the Siberian high and Arctic oscillation (AO) exert a notable influence on the temperature variance. Intra-seasonal variance of the Siberian high and AO are significantly correlated with the temperature variance, whereas the seasonal mean state of the AO affects the temperature variance by modulating the high-frequency components of the Siberian high. The intra-seasonal variance of the Siberian high tends to decline at a rate of change of ,10.7% per decade, significant at the 99% level; meanwhile, the mean wintertime AOs have strengthened in the last few decades. These two climate features together make a considerable contribution to the changes in intra-seasonal temperature variance in East Asia. Copyright © 2004 Royal Meteorological Society [source] Precipitation characteristics of the Eurasian Arctic drainage systemINTERNATIONAL JOURNAL OF CLIMATOLOGY, Issue 11 2003Mark C. Serreze Abstract This study examines characteristics of precipitation over the major watersheds of the Eurasian Arctic drainage system over the period 1960,92. In addition to the Ob, Yenisey and Lena (the three largest drainage systems), we examine the combined Kolyma,Indigirka in eastern Eurasia. Each basin exhibits approximately symmetric mean annual cycles of monthly total precipitation and daily event size, with winter minima and July maxima. These are strikingly similar to the annual cycles of total column water vapour (precipitable water), which fundamentally reflects the control on saturation vapour pressure by temperature. Effective precipitation mechanisms exist in all seasons. However, because of the long distance from strong moisture sources (continentality), precipitation tends to follow the seasonality in column water vapour. An effective contrast is presented for the Iceland sector. Here, the annual cycle of precipitation is tied not to the seasonality in column water vapour, but to the stronger precipitation-generating mechanisms in winter. Hence, the annual cycles of precipitation and column water vapour in this region oppose each other. Mean winter precipitation over the Eurasian watersheds is primarily driven by a modest convergence of water vapour. Whereas precipitation peaks in summer, the mean flux convergence exhibits a general minimum (negative in the Ob). Summer precipitation is hence primarily associated with surface evaporation. A strong role of convection is supported from consideration of static stability, the fairly weak spatial organization of precipitation totals and results from prior studies. On daily time scales, the largest basin-averaged precipitation events, for both summer and winter, are allied with synoptic-scale forcing. This is seen in relationships with cyclone frequency, and patterns of 500 hPa height, vertical motion and the 700 hPa vapour flux. The relative frequency of four 500 hPa synoptic types captures the basic time series structures of precipitation. Copyright © 2003 Royal Meteorological Society [source] The annual cycle and interannual variability of atmospheric pressure in the vicinity of the North PoleINTERNATIONAL JOURNAL OF CLIMATOLOGY, Issue 10 2003Richard I. Cullather Abstract A comparison of National Centers for Environmental Prediction,National Center for Atmospheric Research reanalysis six-hourly sea-level pressure data with former Soviet drifting station observations over the central Arctic Basin reveals high monthly correlations throughout the period 1950,91, but also a preferred winter season negative bias of about 1.4 hPa. Using the reanalysis, supplemented by Arctic Ocean Buoy Program fields and in situ observations, a generalized depiction of the annual cycle of pressure fields over the Arctic may be constructed. Above the Canada Basin,Laptev Sea side of the Arctic, the annual cycle of surface pressure is dominated by the first harmonic, which has an amplitude of about 5 hPa and maximum pressure occurring in March. Along the periphery of northern Greenland and extending to the North Pole, a weak semiannual cycle is found in surface pressure with maxima in May and November. The presence of the semiannual variation over time is highly variable. Dynamically, this progression of the annual cycle may be attributed to the transfer of atmospheric mass from Eurasia and into the Canadian Archipelago in spring and the reverse condition in autumn. Over the central Arctic Basin, springtime pressure increases result from an enhanced poleward mass transport from Eurasia. An increase of equatorward transport over the Canadian Archipelago in May and June results in central Arctic pressure decreases into summer. A less distinct temporal separation between the poleward Canadian transport and the equatorward Eurasian transport results in the weaker second pressure maximum in autumn. On interannual time-scales, atmospheric mass over the central Arctic is exchanged with the storm track centres of action in the North Atlantic and North Pacific. In particular, the large decrease in central Arctic Basin sea-level pressure during the late 1980s is due to a large transfer of atmospheric mass into the North Pacific. Copyright © 2003 Royal Meteorological Society [source] A 10 year cloud climatology over Scandinavia derived from NOAA Advanced Very High Resolution Radiometer imageryINTERNATIONAL JOURNAL OF CLIMATOLOGY, Issue 9 2003Karl-Göran Karlsson Abstract Results from a satellite-based method to compile regional cloud climatologies covering the Scandinavian region are presented. Systematic processing of multispectral image data from the NOAA Advanced Very High Resolution Radiometer (AVHRR) instrument has been utilized to provide monthly cloud climatologies covering the period 1991,2000. Considerable local-scale variation of cloud amounts was found in the region. The inland Baltic Sea and adjacent land areas exhibited a large-amplitude annual cycle in cloudiness (high cloud amounts in winter, low cloud amounts in summer) whereas a weak-amplitude reversed annual cycle (high cloud amounts with a weak maximum in summer) was found for the Scandinavian mountain range. As a contrast, conditions over the Norwegian Sea showed high and almost unchanged cloud amounts during the course of the year. Some interesting exceptions to these patterns were also seen locally. The quality of the satellite-derived cloud climatology was examined through comparisons with climatologies derived from surface cloud observations, from the International Satellite Cloud Climatology Project (ISCCP) and from the European Centre for Medium-range Weather Forecasts ERA-40 data set. In general, cloud amount deviations from surface observations were smaller than 10% except for some individual winter months, when the separability between clouds and snow-covered cold land surfaces is often poor. The ISCCP data set showed a weaker annual cycle in cloudiness, generally caused by higher summer-time cloud amounts in the region. Very good agreement was found with the ERA-40 data set, especially for the summer season. However, ERA-40 showed higher cloud amounts than SCANDIA and ISCCP during the winter season. The derived cloud climatology is affected by errors due to temporal AVHRR sensor degradation, but they appear to be small for this particular study. The data set is proposed as a valuable data set for validation of cloud description in numerical weather prediction and regional climate simulation models. Copyright © 2003 Royal Meteorological Society [source] Spatial and temporal variabilities of rainfall in tropical South America as derived from Climate Prediction Center merged analysis of precipitationINTERNATIONAL JOURNAL OF CLIMATOLOGY, Issue 2 2002H. Matsuyama Abstract We investigated the spatial and temporal variabilities of Climate Prediction Center merged analysis of precipitation (CMAP) in tropical South America from 1979 to 1998. First, we validated CMAP using other hydrometeorological data. In comparison with the high-density precipitation data of the Global Historical Climatology Network (GHCN) Ver. 2, CMAP reproduces the spatial pattern well, although it underestimates (overestimates) heavy (light) precipitation. CMAP also reproduces the interannual variability well, compared with the discharge data of the River Amazon. Next, we applied the rotated empirical orthogonal function (REOF) to CMAP after subtracting the annual cycle. Simultaneous and lag correlations were calculated among the scores of REOFs 1 to 4, the southern oscillation index, and the dipole index of the Atlantic. REOF 1 (15%) represents the north,south pattern that exhibits the maximum precipitation in the summer hemisphere. REOF 2 (12%) indicates the gradual decrease of precipitation in the northern part of tropical South America, reflecting the effect of the Atlantic. REOF 3 (11%) exhibits an east,west pattern related to El Niño. In REOF 4 (7%), the centre of the factor loading is located in Colombia, and the score jumps abruptly around 1985,86. The Lepage test detected the abrupt increase of CMAP in 1985,86 around Colombia. Since such a jump is not found in GHCN Ver. 2, the discontinuous changes of CMAP and REOF 4 around 1985,86 are artificial and peculiar to CMAP. In this region, CMAP should be applied with caution when evaluating recent trends and the interannual variability. The importance of the abrupt increase of precipitation around Colombia is also addressed. Copyright © 2002 Royal Meteorological Society. [source] Climate of the seasonal cycle in the North Pacific and the North Atlantic oceansINTERNATIONAL JOURNAL OF CLIMATOLOGY, Issue 4 2001Igor M. Yashayaev Abstract Time series of monthly sea-surface temperature (SST), air temperature (AT) and sea level pressure (SLP) were constructed from merged releases of the Comprehensive Ocean-Atmosphere Data Set (COADS). The time series were decomposed into seasonal and non-seasonal (short and long-term) components. The contribution of the seasonal cycle to the total variance of SST and AT exceeds 80% in the mid and in some high latitude locations and reaches its peak (>95%) in the centres of subtropical gyres. In most cases, a combination of annual and semiannual harmonics accounts for more than 95% of the seasonal variability. Amplitudes of SST and AT annual cycles are highest near the western boundaries of the oceans; annual phases of SST and AT increase toward the eastern tropical oceans, revealing a southeastern propagation of the annual cycle over the Northern Hemisphere oceans. The annual cycle of AT leads that of SST by 1,3 weeks. The largest phase differences are observed in the regions of western boundary currents in the North Pacific and the North Atlantic oceans. This is consistent with spatial patterns of integral air,sea heat fluxes. Annual phases of SST increase along the Gulf Stream and the Kuroshio Current. This points to the importance of signal transport by the major ocean currents. The lowest annual amplitudes of SLP are observed along the equator (0°,10°N) in both oceans. There are three distinct areas of high annual amplitudes of SLP in the North Pacific Ocean: Asian, Aleutian and Californian. Unlike the North Pacific, only one such area exists in the North Atlantic centred to the west of Iceland. A remarkable feature in the climate of the North Pacific is a maximum of semiannual SLP amplitudes, centred near 40°N and 170°W. It is also an absolute maximum in the entire Northern Hemisphere. Analysis of phases of harmonics of SLP seasonal cycle has revealed the trajectories of propagation of the annual and semiannual cycles. Analysis of semiannual to annual amplitudes ratio has revealed the regions of semiannual cycle dominance. Copyright © 2001 Royal Meteorological Society [source] ,It Takes Two Hands to Clap': How Gaddi Shepherds in the Indian Himalayas Negotiate Access to GrazingJOURNAL OF AGRARIAN CHANGE, Issue 1 2007RICHARD AXELBY This article examines the effects of state intervention on the workings of informal institutions that coordinate the communal use and management of natural resources. Specifically it focuses on the case of the nomadic Gaddi shepherds and official attempts to regulate their access to grazing pastures in the Indian Himalayas. It is often predicted that the increased presence of the modern state critically undermines locally appropriate and community-based resource management arrangements. Drawing on the work of Pauline Peters and Francis Cleaver, I identify key instances of socially embedded ,common' management institutions and explain the evolution of these arrangements through dynamic interactions between individuals, communities and the agents of the state. Through describing the ,living space' of Gaddi shepherds across the annual cycle of nomadic migration with their flocks I explore the ways in which they have been able to creatively reinterpret external interventions, and suggest how contemporary arrangements for accessing pasture at different moments of the annual cycle involve complex combinations of the formal and the informal, the ,traditional' and the ,modern'. [source] Cell proliferation and death in the brain of active and hibernating frogsJOURNAL OF ANATOMY, Issue 2 2009Silvia Cerri Abstract ,Binomial' cell proliferation and cell death have been studied in only a few non-mammalian vertebrates, such as fish. We thought it of interest to map cell proliferation/apoptosis in the brain of the frog (Rana esculenta L.) as this animal species undergoes, during the annual cycle, physiological events that could be associated with central nervous system damage. Therefore, we compared the active period and the deep underground hibernation of the frog. Using western blot analysis for proliferating cell nuclear antigen (PCNA), we revealed a positive 36 kDa band in all samples and found higher optical density values in the hibernating frogs than in active frogs. In both active and hibernating frogs, we found regional differences in PCNA-immunoreactive cells and terminal transferase dUTP nick-end labelling apoptotic cells in the ventricular zones and parenchyma areas of the main encephalon subdivisions. During the active period of the frogs, the highest concentration of PCNA-immunoreactive cells was found in the ventricle dorsal zone of the cerebral hemispheres but only some of the cells were apoptotic. By contrast, the tectal and cerebellar ventricular zones had a small or medium amount of PCNA-immunoreactive cells, respectively, and a higher number of apoptotic cells. During hibernation, an increased PCNA-immunoreactive cell number was observed in both the brain ventricles and parenchyma compared with active frogs. This increase was primarily evident in the lateral ventricles, a region known to be a proliferation ,hot spot'. Although differences existed among the brain areas, a general increase of apoptotic cell death was found in hibernating frogs, with the highest number of apoptotic cells being detected in the parenchyma of the cerebral hemispheres and optic tectum. In particular, the increased number of apoptotic cells in the hibernating frogs compared with active frogs in the parenchyma of these brain areas occurred when cell proliferation was higher in the corresponding ventricular zones. We suggest that the high number of dying cells found in the parenchymal regions of hibernating frogs might provide the stimulus for the ventricular zones to proliferate. Hibernating frogs could utilize an increased cell proliferation in the brain areas as a neuroprotective strategy to face cell death and the onset of neurological damages. Therefore, the hibernator promises to be a valuable model for studying the mechanisms naturally carried out by the central nervous system in order to adapt itself or survive adverse conditions. [source] Carry-over effects in a Pacific seabird: stable isotope evidence that pre-breeding diet quality influences reproductive successJOURNAL OF ANIMAL ECOLOGY, Issue 2 2009Marjorie C. Sorensen Summary 1Understanding the interactions between different periods of the annual cycle in migratory animals has been constrained by our inability to track individuals across seasons. In seabirds, virtually nothing is known about how diet quality during the non-breeding period, away from the breeding grounds, might influence subsequent reproductive success. 2We used stable nitrogen (,15N) and carbon (,13C) isotopes to evaluate the effects of non-breeding diet quality on the timing of breeding and egg size in a population of Cassin's auklets (Ptychoramphus aleuticus) breeding on Triangle Island, British Columbia. Adult feathers are grown during two different periods of the annual cycle, which allowed us to estimate diet quality from the previous fall (October,November) and pre-breeding (February,March) period. 3We found that the estimated proportion of energetically superior copepods (Neocalanus spp.) in the pre-breeding diet tended to be higher in females that bred earlier and laid larger eggs, whereas energetically poor juvenile rockfish (Sebastes spp.) were dominant in the pre-breeding diets of females that bred later and laid smaller eggs. We detected no effect of fall diet quality on breeding date or egg size, and no effect of pre-breeding diet quality on breeding date in males. 4Pre-breeding diet quality was not related to body condition measured 1,2 days after laying, which suggests that females may need to attain a threshold condition before they initiate breeding and successfully rear young. 5Our results suggest that changes in climatic conditions during the pre-breeding period may have severe consequences for reproductive success by influencing breeding date and egg size. Our work emphasizes the importance of determining how events are linked throughout the annual cycle for understanding the fitness and population dynamics of migratory animals. [source] Site fidelity and the demographic implications of winter movements by a migratory bird, the harlequin duck Histrionicus histrionicusJOURNAL OF AVIAN BIOLOGY, Issue 3 2006Samuel A. Iverson Understanding the degree of demographic connectivity among population segments is increasingly recognized as central to the fields of population ecology and conservation biology. However, delineating discrete population units has proven challenging, particularly for migratory birds as they move through their annual cycle. In this study, radio telemetry was used to assess movement rates among habitats by harlequin ducks Histrionicus histrionicus during the non-breeding season in Prince William Sound, Alaska. A total of 434 females were outfitted with radio transmitters over six years of data collection, and their signals tracked by aircraft. Using a spatially nested design, it was determined that 75% of radioed females remained in the bay or coastline area where they were originally trapped, 94% remained on the same island or mainland region of Prince William Sound where they began the winter period, and 98% remained within the 4500 km2 study area as a whole. Home range analyses corroborated these findings, indicating that the scale of individual movements was small, with 95% kernel home range estimates averaging only 11.5±2.2 km2. A simple demographic model, which incorporated estimates for population size, survival, and movement rates, was used to infer the degree of independence among population segments. Immigrant females were found to contribute little to population numbers in most areas, accounting for only 4% of the adult female population at a scale of approximately 100 km2. These results have important implications for the scale of conservation action for the species and demonstrate that winter movements can have a strong influence local population dynamics. [source] Micro- and macroscopic characteristics to stage gonadal maturation of female Baltic codJOURNAL OF FISH BIOLOGY, Issue 2 2003J. Tomkiewicz A set of histological characteristics to judge ovarian development was established and used to elaborate morphological criteria of 10 maturity stages of Baltic cod Gadus morhua sampled throughout the annual cycle to represent different macroscopic maturity stages. The applied characteristics confirmed most stages of the macroscopic scale, but the separation of late immature and resting mature females remained imprecise. Atretic vitellogenic oocytes or encapsulated residual eggs identified the resting condition morphologically, but not all ovaries with visible signs of previous spawning showed such features. One ovarian stage that was previously classified as ,ripening' was changed to ,spawning', owing to the prevalence of hydrated eggs and empty follicles. Ovaries with malfunctions were defined by a separate stage. Macroscopic criteria were revised by comparing the gross anatomy of ovaries with their histology. Female length and gonado-somatic index supported stage definitions, but substantial variation in Fulton's condition factor and the hepato-somatic index rendered these of little use for this purpose. The time of sampling influenced staging accuracy. A female spawner probability function based on the proportion of ripening and ripe specimens in early spring seems to be the most appropriate method to estimate spawner biomass and reproductive potential. [source] Energy density patterns of nectar resources permit coexistence within a guild of Neotropical flower-visiting batsJOURNAL OF ZOOLOGY, Issue 1 2004Marco Tschapka Abstract Neotropical rainforests support guilds of nectar feeding bats (Phyllostomidae: Glossophaginae) with up to six coexisting species. To analyse guild structure and mechanisms of coexistence in a Costa Rican tropical lowland rainforest, the resource use and morphology of bats were compared to the energetic characteristics of preferred nectar resources and their spatio-temporal distribution. The relative abundance of nectar-feeding bats was determined from mistnet captures over 26 months. Food items were identified by analysis of pollen loads and faecal samples. Phenology, flower density and nectar sugar content of resource plants permitted quantitative estimations of resource availability expressed as energy density (kJ ha,1 day,1) throughout the annual cycle. Four glossophagine bat species co-occurred at La Selva: two permanent residents (Glossophaga commissarisi, Hylonycteris underwoodi) and two seasonal species (Lichonycteris obscura, Lonchophylla robusta) that were found in small numbers during a period of high nectar availability. The two resident species differed in their abundance and in their temporal feeding strategies. After the main flowering peak, the common G. commissarisi shifted to a more frugivorous diet, while the rarer H. underwoodi fed on the few remaining bat-flowers. Resource plant species differed in their energy density by up to two orders of magnitude. Hylonycteris underwoodi visited more often plant species with a low energy yield than G. commissarisi. Because of its smaller body size and a wing morphology that promotes fast flight, H. underwoodi appears to be better adapted to low and scattered nectar resource levels. The two seasonal species differed greatly in body mass, which suggests different strategies for high-quality resource tracking. Large body mass in Lonchophylla robusta provides an energy buffer that permits daily commuting flights between a permanent roost and profitable foraging areas, while the small Lichonycteris obscura seems to track resources nomadically. It is proposed that energy density may be a major niche dimension that restricts access of species to certain habitats and that may profoundly influence the structure of nectar-feeding bat guilds. [source] HAUL-OUT ACTIVITY OF RINGED SEALS (PHOCA HISPIDA) DETERMINED FROM SATELLITE TELEMETRYMARINE MAMMAL SCIENCE, Issue 1 2002E. W. Born Abstract The haul-out activity of 15 ringed seals (Phoca hispida) equipped with satellite-linked radio transmitters was studied in NW Greenland (ca. 73°-78°N). Between 19 June 1997 and 30 June 1999, telemetry data on haulout activity were obtained by the "Land-Sea-Reporter" (LSR), "Time-at-Depth" (TAD), and "Timelines" (TIM) systems housed within the satellite transmitters. The haul-out activity (% of total time hauled out) reported by the TIM system, which is specifically designed for collecting haul-out data, was about 1.4 times higher than that inferred from the LSR, but only about 0.7 of that inferred from TAD data. The TIM were used to describe haul-out activity. A total of 1,011 d with TIM were obtained (64.5% of a total of 1,568 "seal-days" monitored) representing data from nearly an entire annual cycle. No differences were found in percentage of time hauled out per month among various age categories. At all seasons the haul-out time showed considerable individual variation. There were no trends in percentage of time hauled out per month during late summer, fall, and winter (August-February). During the High Arctic winter darkness (November-January) the percentage of haul-out per month ranged between 3.9% in an adult (SD = 2.44, range: 1.1%-5.7%, n= 3 mo) and 15.7% in a subadult (SD = 1.95, range: 13.7%-17.6%, n= 3 mo). From late March there was a significant increase in haul-out time. Between 1 and 30 June, when aerial surveys of basking ringed seals usually are conducted, the haul-out time (% per day) increased from about 25% to about 57%. No tendencies in diel haul-out activity were revealed. [source] | ||||||||||||||||||||||||||||||||||