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Glacial Cycles (glacial + cycle)
Kinds of Glacial Cycles Selected AbstractsGeological constraints on Antarctic palaeo-ice-stream retreatEARTH SURFACE PROCESSES AND LANDFORMS, Issue 4 2008Colm Ó Cofaigh Abstract Submarine landforms preserved in bathymetric troughs on the Antarctic continental shelf show that the style of ice stream retreat across the shelf following the last glacial maximum varied between different troughs. Three styles of retreat are inferred from the geological evidence: rapid, episodic and slow. Rapid retreat by ice stream floatation and calving is recorded by the preservation of a landform assemblage of unmodified streamlined subglacial bedforms including mega-scale glacial lineations (MSGLs) that record streaming flow along these troughs. These elongate bedforms are not overprinted by recessional glacial landforms formed transverse to ice flow such as moraines or grounding-zone wedges, and overlying deglacial sediments are thin. A second type of landform assemblage consists of MSGLs overprinted or interrupted by transverse grounding-zone wedges. This assemblage implies episodic retreat between successive grounding-zone positions. The third type of landform assemblage is that of numerous, closely spaced, recessional moraines and intermittent grounding-zone wedges that overlie and interrupt MSGLs. This assemblage records the slow retreat of grounded ice across the shelf. Variation in the style of ice stream retreat between the different bathymetric troughs indicates that Antarctic palaeo-ice-streams did not respond uniformly to external forcing at the end of the last glacial cycle. Rather, their diachronous retreat reflects the dominance of local controls in the form of bathymetry and drainage basin size. More broadly, these data show that retreat of marine-based ice sheets in areas of reverse bed slope is not necessarily catastrophic, and they provide important constraints for numerical models that attempt to predict the dynamics of large polar ice sheets. Copyright © 2008 John Wiley & Sons, Ltd. [source] SPATIAL ASSOCIATIONS BETWEEN LONGEST-LASTING WINTER SNOW COVER AND COLD REGION LANDFORMS IN THE HIGH DRAKENSBERG, SOUTHERN AFRICAGEOGRAFISKA ANNALER SERIES A: PHYSICAL GEOGRAPHY, Issue 2 2009STEFAN W. GRAB ABSTRACT. Although snow is known to influence landform genesis and distribution, the spatial associations between snow and landforms within particular cold regions has received limited research attention. We present a case study from the high Drakensberg of southern Africa, comparing the contemporary spatial pattern of longest-lasting cold-season snow patches with the distribution patterns of active and relic cold region landforms. Two 30 m resolution sets of TM images dated 3 and 19 August 1990 and a DEM were used to demonstrate the geographic trends of snow patch depletion during late winter. Geomorphological phenomena with known coordinates were then incorporated into the GIS. The spatial distribution of several periglacial land-forms (earth hummocks, stone-/turf-banked lobes, block deposits, large sorted patterned ground) coincides with topographic positions that limit snow accumulation. However, the strong spatial association between longest-lasting snow patches and palaeo-moraines implies substantial snow accumulation at some high altitude south-facing sites during the last glacial cycle. [source] LATE-GLACIAL GLACIER EVENTS IN SOUTHERNMOST SOUTH AMERICA: A BLEND OF ,NORTHERN' AND 'SOUTHERN' HEMISPHERIC CLIMATIC SIGNALS?GEOGRAFISKA ANNALER SERIES A: PHYSICAL GEOGRAPHY, Issue 2 2005D.E. SUGDEN ABSTRACT. This paper examines new geomorphological, chronological and modelling data on glacier fluctuations in southernmost South America in latitudes 46,55°S during the last glacial,interglacial transition. Establishing leads and lags between the northern and southern hemispheres and between southern mid-latitudes and Antarctica is key to an appreciation of the mechanisms and resilience of global climate. This is particularly important in the southern hemisphere where there is a paucity of empirical data. The overall structure of the last glacial cycle in Patagonia has a northern hemisphere signal. Glaciers reached or approached their Last Glacial Maxima on two or more occasions at 25,23 ka (calendar) and there was a third less extensive advance at 17.5 ka. Deglaciation occurred in two steps at 17.5 ka and at 11.4 ka. This structure is the same as that recognized in the northern hemisphere and taking place in spite of glacier advances occurring at a time of high southern hemisphere summer insolation and deglaciation at a time of decreasing summer insolation. The implication is that at orbital time scales the,northern' signal dominates any southern hemisphere signal. During deglaciation, at a millennial scale, the glacier fluctuations mirror an antiphase 'southern' climatic signal as revealed in Antarctic ice cores. There is a glacier advance coincident with the Antarctic Cold Reversal at 15.3,12.2 ka. Furthermore, deglaciation begins in the middle of the Younger Dryas. The implication is that, during the last glacial,interglacial transition, southernmost South America was under the influence of sea surface temperatures, sea ice and southern westerlies responding to conditions in the 'southern' Antarctic domain. Such asynchrony may reflect a situation whereby, during deglaciation, the world is more sensitized to fluctuations in the oceanic thermohaline circulation, perhaps related to the bipolar seesaw, than at orbital timescales. [source] Patterns and determinants of shorebird species richness in the circumpolar ArcticJOURNAL OF BIOGEOGRAPHY, Issue 3 2005Sara S. Henningsson Abstract Aim, The intention with this study was first to investigate and describe the broad-scale geographical patterns of species richness of breeding shorebirds (Charadriiformes; families: Charadriidae, Scolopacidae and Haematopodidae) throughout the arctic tundra biome. Secondly, after compensating for the positive relationship between net primary productivity (NPP) and species richness, the relative importance of additional ecological and historical variables for species richness was investigated. The main variables considered are NPP, length of snow- and ice-free season, accessibility of regions depending on migratory flyway systems, tundra area at Pleistocene (120 and 20,18 ka bp) and Holocene (8 ka bp) times, and tundra area at present. Methods, Information on shorebird species breeding distributions was compiled from distribution atlases and species accounts. The breeding distributions of shorebirds with ranges partly or completely in the Arctic (a total of 50 species) were mapped in ArcView 3.2 to create a raster grid layer of shorebird species richness at a 1° latitude × longitude resolution. The total and mean species richness value was calculated per each 10° of longitude sector of the Arctic. The relationships between species richness and the different climatic and environmental variables were analysed on the basis of this sector-wise division of the arctic tundra. The influence of each variable on species richness was investigated using univariate and multivariate analyses (multivariate linear regression and general linear model). Results, We found that patterns of breeding shorebird species richness in the Arctic tundra biome are to a large degree determined by the NPP, the length of the snow- or ice-free season, the diversity of migratory flyways, as well as the historical extent of tundra habitat area during the maximum cooling of the last glacial period. Essentially, two main regions are distinguishable in the circumpolar Arctic regarding shorebird community richness. These are a species-rich Beringia-centred region and a species-poor Atlantic-centred region. Main conclusions, The underlying explanations to these major trends may primarily be attributed to factors that operate at present through accessibility of areas from contemporary migration flyways, as well as processes that operated in the past during and after the last glacial cycle. The most prominent influence on the shorebird diversity was found for NPP in combination with the diversity of migratory flyways. These flyways provide the links between breeding and wintering resources, often separated by huge distances, and the geographical and ecological conditions associated with the shorebirds' migration seem to be of particular importance for their breeding diversity in different sectors of circumpolar tundra. [source] Recognition and palaeoclimatic implications of late Quaternary niche glaciation in eastern Lesotho,JOURNAL OF QUATERNARY SCIENCE, Issue 7 2009Stephanie C. Mills Abstract Geomorphic evidence of former glaciation in the high Drakensberg of southern Africa has proven controversial, with conflicting glacial and non-glacial interpretations suggested for many landforms. This paper presents new geomorphological, sedimentological and micromorphological data, and glacier mass-balance modelling for a site in the Leqooa Valley, eastern Lesotho, preserving what are considered to be moraines of a former niche glacier that existed during the Last Glacial Maximum (LGM). The geomorphology and macro-sedimentology of the deposits display characteristics of both active and passive transport by glacial processes. However, micromorphological analyses indicate a more complex history of glacial deposition and subsequent reworking by mass movement processes. The application of a glacier reconstruction technique to determine whether this site could have supported a glacier indicates a reconstructed glacier equilibrium line altitude (ELA) of 3136,m a.s.l. and palaeoglacier mass balance characteristics comparable with modern analogues, reflecting viable, if marginal glaciation. Radiocarbon dates obtained from organic sediment within the moraines indicate that these are of LGM age. The reconstructed palaeoclimatic conditions during the LGM suggest that snow accumulation in the Drakensberg was significantly higher than considered by other studies, and has substantial relevance for tuning regional climate models for southern Africa during the last glacial cycle. Copyright © 2009 John Wiley & Sons, Ltd. [source] The Last Glacial Maximum in the North Sea Basin: micromorphological evidence of extensive glaciation,JOURNAL OF QUATERNARY SCIENCE, Issue 2 2006S. J. Carr Abstract Despite a long history of investigation, critical issues regarding the last glacial cycle in northwest Europe remain unresolved. One of these refers to the extent, timing and dynamics of Late Devensian/Weichselian glaciation of the North Sea Basin, and whether the British and Scandinavian ice sheets were confluent at any time during this period. This has been the result of the lack of the detailed sedimentological data required to reconstruct processes and environment of sediments recovered through coring. This study presents the results of seismic, sedimentological and micromorphological evidence used to reconstruct the depositional processes of regionally extensive seismic units across the North Sea Basin. Thin section micromorphology is used here to provide an effective means of discriminating between subglacial and glacimarine sediments from cored samples and deriving process-based interpretations from sediment cores. On the basis of micromorphology, critical formations from the basin have been reinterpreted, with consequent stratigraphic implications. Within the current stratigraphic understanding of the North Sea Basin, a complex reconstruction is suggested, with a minimum of three major glacial episodes inferred. On at least two occasions during the Weichselian/Devensian, the British and Scandinavian ice sheets were confluent in the central North Sea. Whilst micromorphology can provide much greater confidence in the interpretation of Late Quaternary offshore stratigraphic sequences, it is noted that a much better geochronology is required to resolve key stratigraphic issues between the onshore and offshore stratigraphic records. Copyright © 2006 John Wiley & Sons, Ltd. [source] Oxygen isotope and palaeotemperature records from six Greenland ice-core stations: Camp Century, Dye-3, GRIP, GISP2, Renland and NorthGRIPJOURNAL OF QUATERNARY SCIENCE, Issue 4 2001Sigfus J. Johnsen Abstract Oxygen isotope variations spanning the last glacial cycle and the Holocene derived from ice-core records for six sites in Greenland (Camp Century, Dye-3, GRIP, GISP2, Renland and NorthGRIP) show strong similarities. This suggests that the dominant influence on oxygen isotope variations reflected in the ice-sheet records was regional climatic change. Differences in detail between the records probably reflect the effects of basal deformation in the ice as well as geographical gradients in atmospheric isotope ratios. Palaeotemperature estimates have been obtained from the records using three approaches: (i) inferences based on the measured relationship between mean annual ,18O of snow and of mean annual surface temperature over Greenland; (ii) modelled inversion of the borehole temperature profile constrained either by the dated isotopic profile, or (iii) by using Monte Carlo simulation techniques. The third of these approaches was adopted to reconstruct Holocene temperature variations for the Dye 3 and GRIP temperature profiles, which yields remarkably compatible results. A new record of Holocene isotope variations obtained from the NorthGRIP ice-core matches the GRIP short-term isotope record, and also shows similar long-term trends to the Dye-3 and GRIP inverted temperature data. The NorthGRIP isotope record reflects: (i) a generally stronger isotopic signal than is found in the GRIP record; (ii) several short-lived temperature fluctuations during the first 1500 yr of the Holocene; (iii) a marked cold event at ca. 8.2 ka (the ,8.2 ka event'); (iv) optimum temperatures for the Holocene between ca. 8.6 and 4.3 ka, a signal that is 0.6, stronger than for the GRIP profile; (v) a clear signal for the Little Ice Age; and (vi) a clear signal of climate warming during the last century. These data suggest that the NorthGRIP stable isotope record responded in a sensitive manner to temperature fluctuations during the Holocene. Copyright © 2001 John Wiley & Sons, Ltd. [source] Intriguing climatic shifts in a 90 kyr old lake record from northern RussiaBOREAS, Issue 1 2008MONA HENRIKSEN A 22 m long sediment core from Lake Yamozero on the Timan Ridge in northern Russia has provided evidence of intriguing climatic shifts during the last glacial cycle. An overall shallowing of the lake is reflected in the lower part of the cores, where pollen indicates a transition from glacial steppe vegetation to interstadial shrub-tundra. These beds are capped by a well-defined layer of compact clay deposited in relatively deep water, where pollen shows surrounding spruce forests and warmer-than-present summer temperatures. The most conservative interpretation is that this unit represents the last interglacial period. However, a series of Optical Stimulated Luminescence (OSL) dates suggests that it corresponds with the Early Weichselian Odderade interstadial (MIS 5a). This would imply that the Odderade interstadial was just as warm as a normal interglacial in this continental part of northern Europe. If correct, then pollen analysis, as a correlation tool, is less straightforward and the definition of an interglacial is more complex than previously thought. We discuss the validity and possible systematic errors of the OSL dates on which this age model is based, but conclude they really indicate a MIS 5a age for the warm period. Above the clay is an unconformity, most likely reflecting a period of subaerial exposure implying dry conditions. Deposition of silt under fluctuating cold climates in the Middle Weichselian continued until a second gap in the record at c. 40 kyr BP. The lake basin started to fill up again around 18 kyr BP. [source] Taxonomic diversity gradients through geological timeDIVERSITY AND DISTRIBUTIONS, Issue 4 2001J. Alistair Crame Abstract., There is evidence from the fossil record to suggest that latitudinal gradients in taxonomic diversity may be time-invariant features, although almost certainly not on the same scale as that seen at the present day. It is now apparent that both latitudinal and longitudinal gradients increased dramatically in strength through the Cenozoic era (i.e. the last 65 my) to become more pronounced today than at any time in the geological past. Present-day taxonomic diversity gradients, in both the marine and terrestrial realms, are underpinned by the tropical radiations of a comparatively small number of species-rich clades. Quite why these particular taxa proliferated through the Cenozoic is uncertain, but it could be that at least part of the explanation involves the phenomenon of evolutionary escalation. This is, in essence, a theory of biological diversification through evolutionary feedback mechanisms between predators and prey; first one develops an adaptive advantage, and then the other. However, there may also have been some form of extrinsic control on the process of tropical diversification, and this was most likely centred on the phenomenon of global climate change. This is especially so over the last 15 my Various Late Cenozoic (Neogene) vicariant events effectively partitioned the tropics into a series of high diversity centres, or foci. It has been suggested that, in the largest of these in the marine realm (the Indo-West Pacific or IWP centre), a critical patterns of islands acted as a template for rapid speciation during glacioeustatic sea level cycles. The same process occurred in the Atlantic, Caribbean and East Pacific (ACEP) centre, though on a lesser scale. Tropical terrestrial diversity may also have been promoted by rapid range expansions and contractions in concert with glacial cycles (a modified refugium hypothesis). We are beginning to appreciate that an integrated sequence of Neogene tectonic and climatic events greatly influenced the formation of contemporary taxonomic diversity patterns. [source] Geomorphology of the onset area of a paleo-ice stream, Marguerite Bay, Antarctic PeninsulaEARTH SURFACE PROCESSES AND LANDFORMS, Issue 4 2008John B. Anderson Abstract Geomorphic features indicate that both glacial and melt-water erosion characterize the onset area of the ancestral Marguerite Ice Stream. The large size of these features indicates that they formed over repeated glacial cycles, most recently during the Last Glacial Maximum. Ice drainage within the bay and on the inner continental shelf was strongly influenced by tectonic fabric. Deep, isolated basins surrounded by rugged bedrock bathymetry characterize the innermost part of the bay. Drumlins and other streamlined features occur in the floors of these basins at depths of up to 900 m. The outer bay has three large interconnected basins. Drumlins and megaflutings within these basins indicate ice was grounded at water depths up to 1000 m. The orientations of these features show convergence of drainage from the northeast, east and south into the Marguerite paleo-ice stream. On the inner continental shelf, the ice converged into a single, wide trough dominated by mega-scale glacial lineations. This transition in geomorphic features from drumlins and megaflutings to mega-scale glacial lineations occurs at the location on the continental shelf where sedimentary strata blanket bedrock, and marks a zone of acceleration of the ice stream. The glacially sculptured geomorphic features within Marguerite Bay co-exist with anastomosing, radial and relatively straight channels, which become increasingly focused in a seaward direction. This implies that a well organized subglacial drainage system existed within the bay at some point in the past. Copyright © 2008 John Wiley & Sons, Ltd. [source] A model computation of the temporal changes of surface gravity and geoidal signal induced by the evolving Greenland ice sheetGEOPHYSICAL JOURNAL INTERNATIONAL, Issue 3 2001Emmanuel Le Meur Summary This paper deals with present-day gravity changes in response to the evolving Greenland ice sheet. We present a detailed computation from a 3-D thermomechanical ice sheet model that is interactively coupled with a self-gravitating spherical viscoelastic bedrock model. The coupled model is run over the last two glacial cycles to yield the loading evolution over time. Based on both the ice sheet's long-term history and its modern evolution averaged over the last 200 years, results are presented of the absolute gravity trend that would arise from a ground survey and of the corresponding geoid rate of change a satellite would see from space. The main results yield ground absolute gravity trends of the order of ±1 µgal yr,1 over the ice-free areas and total geoid changes in the range between ,0.1 and +0.3 mm yr,1. These estimates could help to design future measurement campaigns by revealing areas of strong signal and/or specific patterns, although there are uncertainties associated with the parameters adopted for the Earth's rheology and aspects of the ice sheet model. Given the instrumental accuracy of a particular surveying method, these theoretical trends could also be useful to assess the required duration of a measurement campaign. According to our results, the present-day gravitational signal is dominated by the response to past loading changes rather than current mass changes of the Greenland ice sheet. We finally discuss the potential of inferring the present-day evolution of the Greenland ice sheet from the geoid rate of change measured by the future geodetic GRACE mission. We find that despite the anticipated high-quality data from satellites, such a method is compromised by the uncertainties in the earth model, the dominance of isostatic recovery on the current bedrock signal, and other inaccuracies inherent to the method itself. [source] Genetic divergences pre-date Pleistocene glacial cycles in the New Zealand speckled skink, Oligosoma infrapunctatumJOURNAL OF BIOGEOGRAPHY, Issue 5 2008Stephanie N. J. Greaves Abstract Aim, To examine the hypothesis raised by Graham S. Hardy that Pleistocene glacial cycles suffice to explain divergence among lineages within the endemic New Zealand speckled skink, Oligosoma infrapunctatum Boulenger. Location, Populations were sampled from across the entire range of the species, on the North and South Islands of New Zealand. Methods, We sequenced the mitochondrial genes ND2 (550 bp), ND4 + tRNAs (773 bp) and cytochrome b (610 bp) of 45 individuals from 21 locations. Maximum likelihood, maximum parsimony and Bayesian methods were used for phylogenetic reconstruction. The Shimodaira,Hasegawa test was used to examine hypotheses about the taxonomic status of morphologically distinctive populations. Results, Our analysis revealed four strongly supported clades within O. infrapunctatum. Clades were largely allopatric, except on the west coast of the South Island, where representatives from all four clades were found. Divergences among lineages within the species were extremely deep, reaching over 5%. Two contrasting phylogeographical patterns are evident within O. infrapunctatum. Main conclusions, The deep genetic divisions we found suggest that O. infrapunctatum is a complex of cryptic species which diverged in the Pliocene, contrary to the existing Pleistocene-based hypothesis. Although Pleistocene glacial cycles do not underlie major divergences within this species, they may be responsible for the shallower phylogeographical patterns that are found within O. infrapunctatum, which include a radiation of haplotypes in the Nelson and Westland regions. [source] Phylogeographic structuring and volant mammals: the case of the pallid bat (Antrozous pallidus)JOURNAL OF BIOGEOGRAPHY, Issue 7 2007Sarah E. Weyandt Abstract Aim, To examine the phylogeographic pattern of a volant mammal at the continental scale. The pallid bat (Antrozous pallidus) was chosen because it ranges across a zone of well-studied biotic assemblages, namely the warm deserts of North America. Location, The western half of North America, with sites in Mexico, the United States, and Canada. Methods, PCR amplification and sequencing of the mitochondrial control region was performed on 194 pallid bats from 36 localities. Additional sequences at the cytochrome- b locus were generated for representatives of each control-region haplotype. modeltest was used to determine the best set of parameters to describe each data set, which were incorporated into analyses using paup*. Statistical parsimony and measurements of population differentiation (amova, FST) were also used to examine patterns of genetic diversity in pallid bats. Results, We detected three major lineages in the mitochondrial DNA of pallid bats collected across the species range. These three major clades have completely non-overlapping geographic ranges. Only 6 of 80 control-region haplotypes were found at more than a single locality, and sequences at the more conserved cytochrome- b locus revealed 37 haplotypes. Statistical parsimony generated three unlinked networks that correspond exactly to clades defined by the distance-based analysis. On average there was c. 2% divergence for the combined mitochondrial sequences within each of the three major clades and c. 7% divergence between each pair of clades. Molecular clocks date divergence between the major clades at more than one million years, on average, using the faster rates, and at more than three million years using more conservative rates of evolution. Main conclusions, Divergent haplotypic lineages with allopatric distributions suggest that the pallid bat has responded to evolutionary pressures in a manner consistent with other taxa of the American southwest. These results extend the conclusions of earlier studies that found the genetic structuring of populations of some bat species to show that a widespread volant species may comprise a set of geographically replacing monophyletic lineages. Haplotypes were usually restricted to single localities, and the clade showing geographic affinities to the Sonoran Desert contained greater diversity than did clades to the east and west. While faster molecular clocks would allow for glacial cycles of the Pleistocene as plausible agents of diversification of pallid bats, evidence from co-distributed taxa suggests support for older events being responsible for the initial divergence among clades. [source] Deep genealogies and the mid-peninsular seaway of Baja CaliforniaJOURNAL OF BIOGEOGRAPHY, Issue 8 2006Johan Lindell Abstract Geological forces and long-term climate changes can have profound effects on species. Such effects may be manifested in the pattern and magnitude of genealogical diversity, as revealed by mitochondrial DNA (mtDNA) lineages. The relative importance of the different forces on a regional biota must be evaluated along with a good understanding of geological and climatological history. The peninsula of Baja California of north-western Mexico is one area where both geology and climate have affected the historical biogeography of the regional biota. Molecular studies based on the genealogical relationships among mtDNA lineages have contributed greatly towards elucidating the historical biogeography of Baja California. Perhaps most noticeably, numerous concordant breaks in mtDNA genealogies half-way along the peninsula suggest a vicariant history in which the mid-peninsula was temporarily submerged. This vicariant explanation has recently been criticized, as no conclusive geological evidence exists for a continuous submergence of the mid-peninsula. As an alternative, a scenario based on climatological factors has been suggested. Here we discuss the validity of the hypothesized mid-peninsular vicariance event and the climate-based alternative in explaining the concordant genealogical breaks. We argue that, despite the significant changes in climate brought about by the glacial cycles throughout the Quaternary, a vicariant history involving a mid-peninsular seaway remains the most parsimonious explanation of the observed patterns in mtDNA genealogies. [source] Breeding bird species richness in Taiwan: distribution on gradients of elevation, primary productivity and urbanizationJOURNAL OF BIOGEOGRAPHY, Issue 2 2004Pei-Fen Lee Abstract Aim, To examine the richness of breeding bird species in relation to elevation, primary productivity and urbanization. Location, The island of Taiwan (120°,122° E, 22°,25° N). Methods, We arranged bird species richness (BSR) data from 288 bird censuses undertaken in Taiwan into a 2 × 2 km quadrat system and calculated average values of elevation, primary productivity [surrogated by normalized difference vegetation index (NDVI)], and urbanization (surrogated by road density and percentage of built area) for each 2 × 2 km quadrat. Results, Bird species richness showed a hump-shaped relationship with elevation. It increased with elevation from sea level (10,64 species per 2 × 2 km quadrat), peaked around 2000 m (43,76 species), and then decreased with elevation towards its minimum at the highest elevation. Road density and percentage of built area decreased with elevation, and NDVI showed a hump-shaped relationship with elevation and inverse relationships with road density and percentage of built area. BSR increased with NDVI and decreased with road density and percentage of built area. Linear and cubic terms of elevation together explained 31.3% of the variance in BSR, and road density explained additional 3.4%. The explanatory power of NDVI on BSR was insignificant after the effects of elevation and road density had been justified. Main conclusions, We argue that urbanization plays an important role in the BSR of Taiwan. Urbanization might indirectly decrease BSR through decreasing primary productivity and therefore change the hypothetical inverse relationship between BSR and elevation into a hump-shaped relationship. We also propose a time hypothesis that the biotic communities in the mid-elevation zone of Taiwan had relatively longer periods of existence during the Pleistocene glacial cycles, which might be one underlying process of the observed hump-shaped relationship between species diversity and elevation. [source] Phylogeography of the bigeye chub Hybopsis amblops (Teleostei: Cypriniformes): early Pleistocene diversification and post-glacial range expansionJOURNAL OF FISH BIOLOGY, Issue 8 2008P. B. Berendzen The bigeye chub, Hybopsis amblops, is a member of the Central Highlands ichthyofauna of eastern North America. Phylogenetic analyses of the H. amblops species group based on a 1059 bp fragment of the mitochondrial DNA cytochrome b gene did not recover a monophyletic group. The inclusion of Hybopsis hypsinotus in the species complex is questionable. Within H. amblops, five strongly supported clades were identified; two clades containing haplotypes from the Ozark Highlands and three clades containing haplotypes from the Eastern Highlands and previously glaciated regions of the Ohio and Wabash River drainages. Estimates of the timing of divergence indicated that prior to the onset of glaciation, vicariant events separated populations east and west of the Mississippi River. East of the Mississippi River glacial cycles associated with the blocking and rerouting of the Teays River system caused populations to be pushed southward into refugia of the upper Ohio River. Following the most recent Wisconsinan glaciation, populations expanded northward into previously glaciated regions and southward into the Cumberland River drainage. In the Ozarks, west of the Mississippi River, isolation of clades appears to be maintained by the lack of stream capture events between the upper Arkansas and the White River systems and a barrier formed by the Arkansas River. [source] Response of mid-latitude North Pacific surface temperatures to orbital forcing and linkage to the East Asian summer monsoon and tropical ocean,atmosphere interactions,JOURNAL OF QUATERNARY SCIENCE, Issue 8 2009Masanobu Yamamoto Abstract We present a palaeoceanographic perspective of the North Pacific during the last two glacial cycles based on U -derived palaeotemperature records of IMAGES Core MD01-2421 off the coast of central Japan and cores from the Ocean Drilling Program (ODP) Sites 1014 and 1016 off the coast of California. The sea surface temperature (SST) differences between ODP Sites 1014 and 1016 (,SSTnortheastern Pacific (NEP),=,SSTODP1014 , SSTODP1016) indicate the intensity of the California Current. Comparison of ,SSTNEP and the SST from Core MD01-2421 revealed anti-phase variation; high ,SSTNEP (indicating weakening of the California Current) corresponded to low SST at the Japan margin (indicating the southward displacement of the north-western Pacific subarctic boundary and weakening of the Kuroshio Extension), and vice versa. This finding suggests that the intensity of the North Pacific subtropical gyre circulation has varied in response to precessional forcing and that this response has been linked with changes in tropical ocean,atmosphere interactions. In the precessional cycle, the SST variation derived from Core MD01-2421 lags ca. 2.5,4,ka behind the variations shown by Hulu and Sanbao stalagmite ,18O records and by the pollen temperature index from Core MD01-2421, suggesting out-of-phase variations of the North Pacific subtropical gyre circulation and the East Asian summer monsoon. These findings indicate that the behaviour of interactions between tropical ocean,atmosphere dynamics and the East Asian summer monsoon may have varied in response to the precessional cycle. Copyright © 2009 John Wiley & Sons, Ltd. [source] Comparative phylogeography of five avian species: implications for Pleistocene evolutionary history in the Qinghai-Tibetan plateauMOLECULAR ECOLOGY, Issue 2 2010Y. QU Abstract Pleistocene climate fluctuations have shaped the patterns of genetic diversity observed in extant species. In contrast to Europe and North America where the effects of recent glacial cycles on genetic diversity have been well studied, the genetic legacy of the Pleistocene for the Qinghai-Tibetan (Tibetan) plateau, a region where glaciation was not synchronous with the North Hemisphere ice sheet maxima, remains poorly understood. Here, we compared the phylogeographical patterns of five avian species on the Qinghai-Tibetan plateau by three mitochondrial DNA fragments: the Tibetan snow finch (Montifringilla adamsi), the Blanford's snow finch (Pyrgilauda blanfordi), the horned lark (Eremophila alpestris), the twite (Carduelis flavirostris) and the black redstart (Phoenicurus ochruros). Our results revealed the three species mostly distributed on the platform region of the plateau that experienced population expansion following the retreat of the extensive glaciation period (0.5,0.175 Ma). These results are at odds with the results from avian species of Europe and North America, where population expansions occurred after Last Glacial Maximum (LGM, 0.023,0.018 Ma). A single refugium was identified in a restricted semi-continuous area around the eastern margin of the plateau, instead of multiple independent refugia for European and North American species. For the other two species distributed on the edges of the plateau (the twite and black redstart), populations were maintained at stable levels. Edge areas are located on the eastern margin, which might have had little or no ice cover during the glaciation period. Thus, milder climate may have mitigated demographic stresses for edge species relative to the extremes experienced by platform counterparts, the present-day ranges of which were heavily ice covered during the glaciation period. Finally, various behavioural and ecological characteristics, including dispersal capacities, habitat preference and altitude specificity along with evolutionary history might have helped to shape different phylogeographical structures appearing in these five species. [source] Sea-level changes and palaeo-ranges: reconstruction of ancient shorelines and river drainages and the phylogeography of the Australian land crayfish Engaeus sericatus Clark (Decapoda: Parastacidae)MOLECULAR ECOLOGY, Issue 24 2008MARK B. SCHULTZ Abstract Historical sea levels have been influential in shaping the phylogeography of freshwater-limited taxa via palaeodrainage and palaeoshoreline connections. In this study, we demonstrate an approach to phylogeographic analysis incorporating historical sea-level information in a nested clade phylogeographic analysis (NCPA) framework, using burrowing freshwater crayfish as the model organism. Our study area focuses on the Bass Strait region of southeastern Australia, which is marine region encompassing a shallow seabed that has emerged as a land bridge during glacial cycles connecting mainland Australia and Tasmania. Bathymetric data were analysed using Geographical Information Systems (GIS) to delineate a palaeodrainage model when the palaeocoastline was 150 m below present-day sea level. Such sea levels occurred at least twice in the past 500 000 years, perhaps more often or of larger magnitude within the last 10 million years, linking Victoria and Tasmania. Inter-locality distance measures confined to the palaeodrainage network were incorporated into an NCPA of crayfish (Engaeus sericatus Clark 1936) mitochondrial 16S rDNA haplotypes. The results were then compared to NCPAs using present-day river drainages and traditional great-circle distance measures. NCPA inferences were cross-examined using frequentist and Bayesian procedures in the context of geomorphological and historical sea-level data. We found distribution of present-day genetic variation in E. sericatus to be partly explained not only by connectivity through palaeodrainages but also via present-day drainages or overland (great circle) routes. We recommend that future studies consider all three of these distance measures, especially for studies of coastally distributed species. [source] Lineage diversification and historical demography of a sky island salamander, Plethodon ouachitae, from the Interior HighlandsMOLECULAR ECOLOGY, Issue 24 2008DONALD B. SHEPARD Abstract Sky islands provide ideal opportunities for understanding how climatic changes associated with Pleistocene glacial cycles influenced species distributions, genetic diversification, and demography. The salamander Plethodon ouachitae is largely restricted to high-elevation, mesic forest on six major mountains in the Ouachita Mountains. Because these mountains are separated by more xeric, low-elevation valleys, the salamanders appear to be isolated on sky islands where gene flow among populations on different mountains may be restricted. We used DNA sequence data along with ecological niche modelling and coalescent simulations to test several hypotheses related to diversifications in sky island habitats. Our results revealed that P. ouachitae is composed of seven well-supported lineages structured across six major mountains. The species originated during the Late Pliocene, and lineage diversification occurred during the Middle Pleistocene in a stepping stone fashion with a cyclical pattern of dispersal to a new mountain followed by isolation and divergence. Diversification occurred primarily on an east,west axis, which is likely related to the east,west orientation of the Ouachita Mountains and the more favourable cooler and wetter environmental conditions on north slopes compared to south-facing slopes and valleys. All non-genealogical coalescent methods failed to detect significant population expansion in any lineages. Bayesian skyline plots showed relatively stable population sizes over time, but indicated a slight to moderate amount of population growth in all lineages starting approximately 10 000,12 000 years ago. Our results provide new insight into sky island diversifications from a previously unstudied region, and further demonstrate that climatic changes during the Pleistocene had profound effects on lineage diversification and demography, especially in species from environmentally sensitive habitats in montane regions. [source] Across the southern Andes on fin: glacial refugia, drainage reversals and a secondary contact zone revealed by the phylogeographical signal of Galaxias platei in PatagoniaMOLECULAR ECOLOGY, Issue 23 2008TYLER S. ZEMLAK Abstract We employed DNA sequence variation at two mitochondrial (control region, COI) regions from 212 individuals of Galaxias platei (Pisces, Galaxiidae) collected throughout Patagonia (25 lakes/rivers) to examine how Andean orogeny and the climatic cycles throughout the Quaternary affected the genetic diversity and phylogeography of this species. Phylogenetic analyses revealed four deep genealogical lineages which likely represent the initial division of G. platei into eastern and western lineages by Andean uplift, followed by further subdivision of each lineage into separate glacial refugia by repeated Pleistocene glacial cycles. West of the Andes, refugia were likely restricted to the northern region of Patagonia with small relicts in the south, whereas eastern refugia appear to have been much larger and widespread, consisting of separate northern and southern regions that collectively spanned most of Argentinean Patagonia. The retreat of glacial ice following the last glacial maximum allowed re-colonization of central Chile from nonlocal refugia from the north and east, representing a region of secondary contact between all four glacial lineages. Northwestern glacial relicts likely followed pro-glacial lakes into central Chilean Patagonia, whereas catastrophic changes in drainage direction (Atlantic , Pacific) for several eastern palaeolakes were the likely avenues for invasions from the east. These mechanisms, combined with evidence for recent, rapid and widespread population growth could explain the extensive contemporary distribution of G. platei throughout Patagonia. [source] Living on the edge: demographic and phylogeographical patterns in the woodlouse-hunter spider Dysdera lancerotensis Simon, 1907 on the eastern volcanic ridge of the Canary IslandsMOLECULAR ECOLOGY, Issue 15 2007LETICIA BIDEGARAY-BATISTA Abstract The Eastern Canary Islands are the emerged tips of a continuous volcanic ridge running parallel to the northeastern African coast, originated by episodic volcanic eruptions that can be traced back to the Miocene and that, following a major period of quiescence and erosion, continued from the Pliocene to the present day. The islands have been periodically connected by eustatic sea-level changes resulting from Pleistocene glacial cycles. The ground-dwelling spider Dysdera lancerotensis Simon, 1907 occurs along the entire ridge, except on recent barren lavas and sand dunes, and is therefore an ideal model for studying the effect of episodic geological processes on terrestrial organisms. Nested clade and population genetic analyses using 39 haplotypes from 605 base pairs of mitochondrial DNA cytochrome c oxidase I sequence data, along with phylogenetic analyses including two additional mitochondrial genes, uncover complex phylogeographical and demographic patterns. Our results indicate that D. lancerotensis colonized the ridge from north to south, in contrast to what had been expected given the SSW-NNE trend of volcanism and to what had been reported for other terrestrial arthropods. The occurrence of several episodes of extinction, recolonization and expansion are hypothesized for this species, and areas that act as refugia during volcanic cycles are identified. Relaxed molecular clock methods reveal divergence times between main haplotype lineages that suggest an older origin of the northern islets than anticipated based on geological evidence. This study supports the key role of volcanism in shaping the distribution of terrestrial organisms on oceanic islands and generates phylogeographical predictions that warrant further research into other terrestrial endemisms of this fascinating region. [source] Pleistocene refugia in an arid landscape: analysis of a widely distributed Australian passerineMOLECULAR ECOLOGY, Issue 12 2007ALICIA TOON Abstract While many studies have documented the effect that glacial cycles have had on northern hemisphere species, few have attempted to study the associated effect of aridification at low latitudes in the southern hemisphere. We investigated the past effects that cyclic aridification may have had on the population structure and history of a widespread endemic Australian bird species, the Australian magpie (Gymnorhina tibicen). One thousand one hundred and sixty-six samples from across its native range were analysed for mitochondrial control region sequence variation and variation at six microsatellite loci. Analysis of mitochondrial control region sequence data indicated monophyletic clades that were geographically congruent with an eastern and western region. The contemporary distribution of east and west clades is nonoverlapping but in close proximity. Populations were estimated to have diverged in the Pleistocene around 36 000 years ago. The putative Carpentarian and Nullarbor arid barriers appear to be associated with the divergence between east and west mainland populations. Nested clade analysis indicated a signature of range expansion in the eastern region suggesting movement possibly inland and northward subsequent to the last period of aridity. The island population of Tasmania was of very recent origin, possibly since sea levels rose 16 000 years ago. Given the east-west structure, there was no congruence between morphology and recent history of this species indicating a lack of support for morphological taxa. Overall mitochondrial DNA and microsatellite variation suggest that increasing aridity and Pleistocene refugia played a role in structuring populations of the Australian magpie; however, the dispersal ability and generalist habitat requirements may have facilitated the movement of magpies into an almost contiguous modern distribution across the continent. This study supports the idea that Pleistocene aridification played an important role in structuring intraspecific variation in low latitudinal southern hemisphere avian species. [source] Phylogeography of the mountain chickadee (Poecile gambeli): diversification, introgression, and expansion in response to Quaternary climate changeMOLECULAR ECOLOGY, Issue 5 2007GARTH M. SPELLMAN Abstract Since the late 1990s, molecular techniques have fuelled debate about the role of Pleistocene glacial cycles in structuring contemporary avian diversity in North America. The debate is still heated; however, there is widespread agreement that the Pleistocene glacial cycles forced the repeated contraction, fragmentation, and expansion of the North American biota. These demographic processes should leave genetic ,footprints' in modern descendants, suggesting that detailed population genetic studies of contemporary species provide the key to elucidating the impact of the late Quaternary (late Pleistocene,Holocene). We present an analysis of mitochondrial DNA (mtDNA) variation in the mountain chickadee (Poecile gambeli) in an attempt to examine the genetic evidence of the impact of the late Quaternary glacial cycles. Phylogenetic analyses reveal two strongly supported clades of P. gambeli: an Eastern Clade (Rocky Mountains and Great Basin) and a Western Clade (Sierra Nevada and Cascades). Post-glacial introgression is apparent between these two clades in the Mono Lake region of Central California. Within the Eastern Clade there is evidence of isolation-by-distance in the Rocky Mountain populations, and of limited gene flow into and around the Great Basin. Coalescent analysis of genetic variation in the Western Clade indicates that northern (Sierra Nevada/Cascades) and southern (Transverse/Peninsular Ranges) populations have been isolated and evolving independently for nearly 60 000 years. [source] Refugial isolation and secondary contact in the dyeing poison frog Dendrobates tinctoriusMOLECULAR ECOLOGY, Issue 14 2006BRICE P. NOONAN Abstract Recent palaeoclimactic research suggests that fluctuating environmental conditions throughout the Pleistocene of Amazonia occurred with previously unrecognized frequency. This has resulted in a theoretical shift from glacially influenced fluctuations to those driven by precessional rhythms. This theoretical revolution has a profound impact on expectations of biotic diversity within biogeographical regions that have long been based on the idea of large-scale landscape fragmentation associated with increased aridity and glacial cycles. Generally speaking, this shifts phylogeographical expectations from that of (i) large areas of sympatry of closely related (but not sister) species whose origins lie in separate refugia, and current distributions are the results of cyclic connectivity of those two refugia (refuge hypothesis), to that of (ii) fine scale genetic structure, often associated with elevation, and divergence well below expected speciation levels [disturbance,vicariance (DV) hypothesis]. To date there have been few tests of the expectations of the DV hypothesis based on empirical studies of Neotropical floral and faunal communities. Herein we examine phylogeographical structure of Dendrobates tinctorius, an amphibian species endemic to the uplands of the eastern Guiana Shield, based on sampling of 114 individuals from 24 localities. Phylogenetic, nested clade, and dispersal,vicariance (DIVA) analyses of cytochrome b sequence data reveal the presence of two mitochondrial lineages that are associated with previously identified western and eastern uplands of this area. The geographical distribution of mitochondrial haplotypes and the results of DIVA and coalescent analyses suggest that there has been extensive secondary contact between these lineages indicating a complex history of connectivity between these western and eastern highlands, supporting the predictions of the DV hypothesis. [source] Phylogeography of the montane caddisfly Drusus discolor: evidence for multiple refugia and periglacial survivalMOLECULAR ECOLOGY, Issue 8 2006STEFFEN U. PAULS Abstract We studied the genetic population structure and phylogeography of the montane caddisfly Drusus discolor across its entire range in central and southern Europe. The species is restricted to mountain regions and exhibits an insular distribution across the major mountain ranges. Mitochondrial sequence data (COI) of 254 individuals from the entire species range is analysed to reveal population genetic structure. The data show little molecular variation within populations and regions, but distinct genetic differentiation between mountain ranges. Most populations are significantly differentiated based on FST and exact tests of population differentiation and most haplotypes are unique to a single mountain range. Phylogenetic analyses reveal deep divergence between geographically isolated lineages. Combined, these results suggest that past fragmentation is the prominent process structuring the populations across Europe. We use tests of selective neutrality and mismatch distributions, to study the demographic population history of regions with haplotype overlap. The high level of genetic differentiation between mountain ranges and estimates of demographic history provide evidence for the existence of multiple glacial refugia, including several in central Europe. The study shows that these aquatic organisms reacted differently to Pleistocene cooling than many terrestrial species. They persisted in numerous refugia over multiple glacial cycles, allowing many local endemic clades to form. [source] Substantial genetic substructuring in southeastern and alpine Australia revealed by molecular phylogeography of the Egernia whitii (Lacertilia: Scincidae) species groupMOLECULAR ECOLOGY, Issue 5 2005DAVID G. CHAPPLE Abstract Palaeoclimatic events and biogeographical processes since the mid-Tertiary are believed to have strongly influenced the evolution and distribution of the terrestrial vertebrate fauna of southeastern Australia. We examined the phylogeography of the temperate-adapted members of the Egernia whitii species group, a group of skinks that comprise both widespread low- to mid-elevation (E. whitii) and montane-restricted species (Egernia guthega, Egernia montana), in order to obtain important insights into the influence of past biogeographical processes on the herpetofauna of southeastern Australia. Sequence data were obtained from all six temperate-adapted species within the E. whitii species group, and specifically from across the distributional ranges of E. whitii, E. guthega and E. montana. We targeted a fragment of the ND4 mitochondrial gene (696 bp) and analysed the data using maximum likelihood and Bayesian methods. Our data reveal a deep phylogeographical break in the east Gippsland region of Victoria between ,northern' (Queensland, New South Wales, Australian Capital Territory) and ,southern' (Victoria, Tasmania, South Australia) populations of E. whitii. This divergence appears to have occurred during the late Miocene,Pliocene, with the Gippsland basin possibly forming a geographical barrier to dispersal. Substantial structuring within both the ,northern' and the ,southern' clades is consistent with the effects of Plio,Pleistocene glacial-interglacial cycles. Pleistocene glacial cycles also appear to have shaped the phylogeographical patterns observed in the alpine species, E. guthega and E. montana. We used our results to examine the biogeographical process that led to the origin and subsequent diversification of the lowland and alpine herpetofauna of southeastern Australia. [source] Mitochondrial phylogeography of the moor frog, Rana arvalisMOLECULAR ECOLOGY, Issue 6 2004W. Babik Abstract The moor frog Rana arvalis is a lowland species with a broad Eurasiatic distribution, from arctic tundra through forest to the steppe zone. Its present-day range suggests that glacial refugia of this frog were located outside southern European peninsulas. We studied the species-wide phylogeographical pattern using sequence variation in a 682 base pairs fragment of mtDNA cytochrome b gene; 223 individuals from 73 localities were analysed. Two main clades, A and B, differing by c. 3.6% sequence divergence were detected. The A clade is further subdivided into two subclades, AI and AII differing by 1.0%. All three lineages are present in the Carpathian Basin (CB), whereas the rest of the species range, including huge expanses of Eurasian lowlands, are inhabited solely by the AI lineage. We infer that AII and B lineages survived several glacial cycles in the CB but did not expand, at least in the present interglacial, to the north. The geographical distribution and genealogical relationships between haplotypes from the AI lineage indicate that this group had two glacial refugia, one located in the eastern part of the CB and the other probably in southern Russia. Populations from both refugia contributed to the colonization of the western part of the range, whereas the eastern part was colonized from the eastern refugium only. The effective population size as evidenced by ,ML is an order of magnitude higher in the AI lineage than in the AII and B lineages. Demographic expansion was detected in all three lineages. [source] Deep, hierarchical divergence of mitochondrial DNA in Amplirhagada land snails (Gastropoda: Camaenidae) from the Bonaparte Archipelago, Western AustraliaBIOLOGICAL JOURNAL OF THE LINNEAN SOCIETY, Issue 1 2010MICHAEL S. JOHNSON Continental islands have experienced cycles of isolation and connection. Although complex genetic patterns have been described for mainland species affected by glacial cycles of isolation, island biotas have received little attention. We examined mitochondrial DNA in Amplirhagada land snails from 16 islands and two adjacent mainland areas of the Bonaparte Archipelago, in the Kimberley region of northern Western Australia. Four major clades, with sequence divergence of 16,27% in the 16S ribosomal RNA gene, correspond to the major geographic groupings, separated by 10,160 km. Distinct lineages also characterize islands that are only a few kilometres apart. The large differences indicate that the lineages are much older than the islands themselves, and show no evidence of geologically recent connection. Three of the major clades match the morphological description of Amplirhagada alta. Either this named species comprises several morphologically cryptic species, or it is a single, genetically very diverse species, distributed over much of the Bonaparte Archipelago. © 2010 The Linnean Society of London, Biological Journal of the Linnean Society, 2010, 100, 141,153. [source] Varying rates of diversification in the genus Melitaea (Lepidoptera: Nymphalidae) during the past 20 million yearsBIOLOGICAL JOURNAL OF THE LINNEAN SOCIETY, Issue 2 2009JULIEN LENEVEU The influence of Quarternary glacial cycles on the extant diversity of Holarctic species has been intensively studied. It has been hypothesized that palaeoclimatic changes are responsible for divergence events in lineages. A constant improvement in DNA sequencing and modeling methods, as well as palaeoclimatic reconstruction, permit a deeper exploration of general causes of speciation in geological time. In the present study, we sampled, as exhaustively as possible, the butterflies belonging to the genus Melitaea (Lepidoptera: Nymphalidae), which are widely spread in the Palaearctic region. We conducted analyses to assess the phylogeny of the genus and estimated the timing of divergence and the most likely distribution of ancestral populations. The results obtained indicate that the systematics of the genus is in need of revision and that the diversity of the genus has been profoundly shaped by palaeoenvironmental changes during its evolutionary history. The present study also emphasizes that, when employed with caveats, major palaeoenvironmental events could represent very powerful tools for the calibration of the dating of divergences using molecular data. © 2009 The Linnean Society of London, Biological Journal of the Linnean Society, 2009, 97, 346,361. [source] | ||||||||||||||||