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Constant Proportion (constant + proportion)
Selected AbstractsA comparative study of the dispersal of 10 species of stream invertebratesFRESHWATER BIOLOGY, Issue 9 2003J. M. Elliott Summary 1. Apart from downstream dispersal through invertebrate drift, few quantitative data are available to model the dispersal of stream invertebrates, i.e. the outward spreading of animals from their point of origin or release. The present study provides comparative data for 10 species, using two independent methods: unmarked animals in six stream channels built over a stony stream and marked animals in the natural stream. Experiments were performed in April and June 1973 and 1974, with initial numbers of each species varying from 20 to 80 in the stream channels and 20 to 60 for marked animals. 2. Results were the same for marked invertebrates and those in the channels. Dispersal was not density-dependent; the number of dispersing animals was a constant proportion of the initial number for each species. The relationship between upstream or downstream dispersal distance and the number of animals travelling that distance was well described by an inverse power function for all species (exponential and log models were poorer fits). Results varied between species but were similar within species for the 4 months, and therefore were unaffected by variations in mean water velocity (range 0.04,0.35 m s,1) or water temperature (range 6.7,8.9 °C in April, 12.1,14.8 °C in June). 3. Species were arranged in order, according to their dispersal abilities. Three carnivores (Perlodes, Rhyacophila, Isoperla) dispersed most rapidly (70,91% in 24 h, maximum distances 9.5,13.5 m per day), followed by two species (Protonemura, Rhithrogena) in which about half their initial numbers dispersed (50,51% in 24 h, 7.5,8 m per day), and four species (Ecdyonurus, Hydropsyche, Gammarus, Baetis) in which less than half dispersed (33,40% in 24 h, 5.5,7 m per day). Dispersal was predominantly upstream for all nine species. Few larvae (20%) of Potamophylax dispersed, with similar maximum upstream and downstream distances of 3.5 m per day. The mean time spent drifting downstream was known for seven species from previous studies, and correlated positively with their dispersal distances. Therefore, the species formed a continuum from rapid to very slow dispersers. These interspecific differences should be considered when evaluating the role of dispersal in the maintenance of genetic diversity in stream invertebrates, and in their ability to colonise or re-colonise habitats. [source] Temperature and host species affect nuptial gift size in a seed-feeding beetleFUNCTIONAL ECOLOGY, Issue 6 2006C. W. FOX Summary 1In many insects species, males contribute large nutritional gifts to females during mating, generally as seminal fluids (ejaculates) or spermatophores. These nuptial gifts can affect both male and female fitness, and can mediate selection on male body size. However, it is unclear how environmental variables, such as temperature and diet, affect gift size and the consequences of gift size for male and female fitness. 2We examine how temperature and rearing host affect male nuptial gift size (both total ejaculate size and the proportion of a male's mass allocated to his seminal fluids), and the relationship between gift size and female reproduction, in two populations of the seed-feeding beetle Callosobruchus maculatus. 3Males reared at lower temperature (20 °C) produced substantially larger ejaculates than males reared at higher temperatures (25, 30 and 35 °C). However, males allocated a smaller proportion of their body mass to their ejaculate at the lowest temperature compared with other temperatures. This effect of temperature on male allocation to their ejaculates mirrored the effect of temperature on female body size , male ejaculate size remained a relatively constant proportion of their mate's body mass across temperatures. 4Rearing host also affected male ejaculate size but the magnitude and direction of the host effect differed between populations. 5Rearing temperature affected the relationship between male body mass and ejaculate size. Temperature also affected the relationship between female body mass and fecundity. The relationship between male body mass and ejaculate size was significantly lower when beetles were reared on cowpea than when beetles were reared on azuki or mung. 6We found no evidence that male body size or nuptial gift size affected female fecundity in either population of C. maculatus. We thus propose that the effect of nuptial gift size on male fitness is through a reduction in female mating frequency and thus increased paternity for males producing larger nuptial gifts. [source] The evolutionary species pool hypothesis and patterns of freshwater diatom diversity along a pH gradientJOURNAL OF BIOGEOGRAPHY, Issue 3 2005Jason Pither Abstract Aim, To interpret the unimodal relationship between diatom species richness and lake pH within the context of the evolutionary species pool hypothesis (SPH). We test the following primary prediction arising from the SPH: the size of the potential species pool (PSP) will increase along a gradient representing the historical commonness of different pH environments (pH commonness). To do this we assume that the present-day spatial dominance of near-neutral pH conditions compared with acidic and alkaline conditions reliably mimics the relative spatial availabilities of historical pH conditions among freshwater lakes. We also determine whether local richness represents a constant proportion of PSP size along the pH commonness gradient. Location, Two hundred and thirty-four lakes distributed over a 405,000 km2 region of the north-eastern United States of America. Methods, Sediment diatom morphospecies lists and pH data were acquired from the US Environmental Protection Agency's Environmental Monitoring and Assessment Program (EMAP) website. Using 248 morphospecies that occurred in at least 10 of the 234 lakes, four different measures of PSPs were calculated along the pH gradient. Local species richness was equated with the number of species occurring within the lake. Alpha diversity was equated with the average species richness of lakes with similar pH values. A combination of statistical methods were employed, including correlations, quadratic regression and piecewise regression. Results, PSP size increased significantly with pH commonness for all four measures of PSP size, thus supporting the primary prediction of the evolutionary SPH. Local richness comprised a larger proportion of the PSP within acidic lakes than within circumneutral lakes. Alpha diversity and lake species richness both increased significantly with pH commonness, but the former did so in a two-step fashion. We test and reject several alternative contemporary time-scale explanations for our findings. Main Conclusions, Our findings are consistent with the hypothesis that diatom taxonomic richness is presently lower within acidic and highly alkaline lakes than in circumneutral lakes owing to the limited opportunity in space and/or time for the evolution of suitably adapted species. Whereas ecological processes can explain why certain species are excluded from particular habitats, e.g. acidic lakes, they cannot account for why so few species are adapted to those habitats in the first place. [source] The importance of growth and mortality costs in the evolution of the optimal life historyJOURNAL OF EVOLUTIONARY BIOLOGY, Issue 6 2006D. A. ROFF Abstract A central assumption of life history theory is that the evolution of the component traits is determined in part by trade-offs between these traits. Whereas the existence of such trade-offs has been well demonstrated, the relative importance of these remains unclear. In this paper we use optimality theory to test the hypothesis that the trade-off between present and future fecundity induced by the costs of continued growth is a sufficient explanation for the optimal age at first reproduction, ,, and the optimal allocation to reproduction, G, in 38 populations of perch and Arctic char. This hypothesis is rejected for both traits and we conclude that this trade-off, by itself, is an insufficient explanation for the observed values of , and G. Similarly, a fitness function that assumes a mortality cost to reproduction but no growth cost cannot account for the observed values of ,. In contrast, under the assumption that fitness is maximized, the observed life histories can be accounted for by the joint action of trade-offs between growth and reproductive allocation and between mortality and reproductive allocation (Individual Juvenile Mortality model). Although the ability of the growth/mortality model to fit the data does not prove that this is the mechanism driving the evolution of the optimal age at first reproduction and allocation to reproduction, the fit does demonstrate that the hypothesis is consistent with the data and hence cannot at this time be rejected. We also examine two simpler versions of this model, one in which adult mortality is a constant proportion of juvenile mortality [Proportional Juvenile Mortality (PJM) model] and one in which the proportionality is constant within but not necessarily between species [Specific Juvenile Mortality (SSJM) model]. We find that the PJM model is unacceptable but that the SSJM model produces fits suggesting that, within the two species studied, juvenile mortality is proportional to adult mortality but the value differs between the two species. [source] TAXES, GROWTH AND THE CURRENT ACCOUNT TICK-CURVE EFFECTAUSTRALIAN ECONOMIC PAPERS, Issue 1 2010CREINA DAY This paper examines the dynamic and long run effects of a shift from income taxes to consumption taxes in a growing small open economy. We introduce a government sector that maintains a balanced budget and expenditure at a constant proportion of domestic income to a small open economy Swan-Solow model. Our framework provides a previously unidentified dynamic effect that is robust to endogenising the savings rate. Lowering the income tax rate promotes economic growth and has a tick-curve effect on the current account balance, characterised by instantaneous deterioration, a period of recovery and gradual convergence to an improved position in the long run. [source] | ||||||||||