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Competitors
Kinds of Competitors Terms modified by Competitors Selected AbstractsWHOLESALE PRICING WHEN BUYERS ARE ASYMMETRIC COURNOT COMPETITORS,THE MANCHESTER SCHOOL, Issue 2 2006GIUSEPPE COLANGELO This paper focuses on the pricing policy of a well-informed profit- maximizing producer selling to asymmetric retailers who compete à la Cournot. An optimal upstream two-part tariff implies the exit of the inefficient retailer, thus causing downstream monopolization. When this would bring about a significant increase in the efficient retailer's bargaining power, as is plausible, the producer will try to avoid this and consequently choose a pricing scheme that does not cause downstream monopolization. When this is the case, two alternatives emerge: a two-part tariff (ensuring no downstream monopolization) or third-degree price discrimination. The more asymmetric in cost retailers are (consistent with no downstream monopolization), the more likely it is to see third-degree price discrimination as the equilibrium wholesale pricing. When third-degree price discrimination is implemented, a welfare loss is easily produced. [source] OPTIMUM-WELFARE AND MAXIMUM-REVENUE TARIFFS IN MIXED OLIGOPOLY WITH FOREIGN COMPETITORSAUSTRALIAN ECONOMIC PAPERS, Issue 1 2010LEONARD F.S. WANG This paper re-examines the important tariff ranking issue under a linear mixed oligopoly model with foreign competitors and asymmetric costs. We demonstrate that under Cournot competition, when the size of domestic private and foreign private firms become more unequally distributed, optimum-welfare tariff will exceed maximum-revenue tariff. We also show that under Stackelberg competition, when the domestic government protects its domestic sector, it will levy higher optimum-welfare tariffs versus maximum-revenue tariffs; however, when it decides to open its doors more for foreign competitors, it will need to levy higher maximum-revenue tariffs versus optimum-welfare tariffs. The above results remain valid whether the domestic public firm acts as a leader or a follower. [source] Innate and Learned Components of Defence by Flickers Against a Novel Nest Competitor, the European StarlingETHOLOGY, Issue 10 2004Karen L. Wiebe Defence against predators is an important component of fitness in wild birds but the first step of defence, predator recognition, is not well understood. Anti-predator behaviour may innate, in which case the individual responds without prior contact with that predator, and/or there may be a learned component that develops only after direct experience. In the wild, the development of anti-predator behaviour is studied by exposing naive individuals to novel predators. I studied responses of 71 naive and experienced northern flickers Colaptes auratus, to a novel nest predator and competitor, the European starling Sturnus vulgaris that was introduced to North America. Naive individuals responded more intensely to the model starling than to the control model suggesting an innate component to recognition. However, there was also a learned component to defence because flickers nesting near to starlings reacted more aggressively than naive individuals far from starlings. Consistent with theory on life histories and optimal defence levels, no significant differences in aggression were found between the sexes or between age classes. Selection should favour more intense, and possibly innate, defence against the introduced starling. Variation in responses of naive individuals suggests that there may already be some alleles in the population associated with higher defence, but that these may not be uniform within the population. [source] Stackelberg Mixed Duopoly with a Foreign CompetitorBULLETIN OF ECONOMIC RESEARCH, Issue 3 2003Toshihiro Matsumura H42; L13 We investigate Stackelberg mixed duopoly models where a state-owned public firm and a foreign private firm compete. We examine a desirable role (either leader or follower) of the public firm. We also consider endogenous roles by adopting the observable delay game of Hamilton and Slutsky (1990). We find that, in contrast to Pal (1998) discussing a case of domestic competitors, the public firm should be the leader and that it becomes the leader in the endogenous role game. We also find that in contrast to Ono (1990) eliminating a foreign firm does not improve domestic welfare in mixed oligopolies. [source] The Smell of New Competitors: The Response of American Mink, Mustela vison, to the Odours of Otter, Lutra lutra and Polecat, M. putoriusETHOLOGY, Issue 5 2009Lauren A. Harrington We tested the response of wild American mink (an established alien species in the UK), to the odours of unfamiliar mink, European polecat and Eurasian otter. Polecats are similar in size and habits to mink, otters are larger than mink and a dominant competitor; both are native to the UK and both were absent during the original colonization by mink but are now undergoing natural population recoveries. The response of mink to experimental odours was assessed by counting the numbers of tracks (footprints) on rafts treated with anal gland secretions, and compared with response to a control raft, on two rivers in the Upper Thames valley, UK. Remote video showed that the number of tracks was positively correlated with the time that mink spent investigating an odour. We found that mink were attracted to the odours of both unfamiliar mink and polecats. There was little evidence that mink avoided the odour of otters. We suggest that, during an encounter with a polecat, mink may behave much as they would to a conspecific. We infer from the response of mink to the odour of otters, that, if mink do avoid otters, the mechanism of avoidance is likely to be complex, situation-dependent and perhaps affected by prior experience. [source] Does Host Value Influence Female Aggressiveness, Contest Outcome and Fitness Gain in Parasitoids?ETHOLOGY, Issue 4 2007Marlene Goubault Intraspecific competition for resources is common in animals and may lead to physical contests. Contest outcomes and aggressiveness can be influenced by the resource holding potential of contestants but also by their perception of the resource value (RV). Competitors may assess resource quality directly (real RV) but may also estimate it according to their physiological status and their experience of the habitat quality (subjective RV). In this article, we studied contests between females of the solitary parasitoid Pachycrepoideus vindemmiae Rondani (Hymenoptera: Pteromalidae) when exploiting simultaneously a host, a Delia radicum L. (Diptera: Anthomyiidae) pupa. We tested the effect of factors modifying host value on the occurrence of agonistic behaviours, contest outcomes and host exploitation. The factors tested were: the quality of the previous habitat experienced by females, female egg load, host parasitism status and the stage reached by the owner female in her behavioural oviposition sequence. Females successfully protected their host against intruders during its exploitation, but not after oviposition, and their aggressiveness did not seem to be influenced by their perception of the RV. The fact that the host is subsequently parasitized by the opponent females appears to mainly depend on the host selectiveness of females. [source] Parasitized Salamanders are Inferior Competitors for Territories and Food ResourcesETHOLOGY, Issue 4 2000Daria S. Maksimowich Parasites have been shown to impair the behaviour of their hosts, compromising the host's ability to exploit and compete for resources. We conducted two experiments to determine whether infestation with an ectoparasitic mite (Hannemania eltoni) was associated with changes in aggressive and foraging behaviour in the Ozark zigzag salamander, Plethodon angusticlavius. In a first experiment, male salamanders with high parasite loads were less aggressive overall than males with low parasite loads during territorial disputes. In addition, males with high parasite loads were more aggressive toward opponents with high parasite loads (symmetric contests) than toward opponents with low parasite loads (asymmetric contests). In contrast, males with low parasite loads did not adjust their level of aggression according to the parasite load of the opponent. In a second experiment, foraging behaviour of females was tested in response to ,familiar' (Drosophila) prey and ,novel' (termite) prey. Latency to first capture was significantly longer for parasitized than non-parasitized females when tested with ,familiar' prey, but not for ,novel' prey. Our results suggest that parasite-mediated effects may have profound influences on individual fitness in nature. [source] Discriminatory Dealing with Downstream Competitors: Evidence from the Cellular IndustryTHE JOURNAL OF INDUSTRIAL ECONOMICS, Issue 3 2000David Reiffen One concern about regulated monopolies entering unregulated vertically-related markets is that they will discriminate against competitors of their unregulated affiliates. However, prohibiting regulated monopolies from offering related goods may preclude production by the most efficient provider. We take advantage of variation across geographic cellular phone markets in the US to examine the effect of integration on output, quality and prices. We find some evidence consistent with efficiencies (greater concentration of lines to users is associated with greater output and higher quality) and some consistent with discrimination (greater interconnection facility ownership concentration is associated with lower output and quality). [source] Strategic Bidding By Potential Competitors: Will Monopoly Persist?THE JOURNAL OF INDUSTRIAL ECONOMICS, Issue 2 2000Yongmin Chen Who will win the bidding to become the sole producer of a new product: the monopolist of a related product or a new entrant? When there exists potential entry to the monopolist's existing business, the standard result that monopoly persists (Gilbert and Newbery, ,Preemptive Patenting and the Persistence of Monopoly', American Economic Review, 72, pp. 514,526, 1982) may or may not hold, depending crucially on how the new product relates to the existing product of the monopolist. The monopolist tends to win the bidding and to dominate both products if the two products are strategic complements; and the entrant tends to win the bidding if the two products are strategic substitutes. [source] Launch decisions and competitive reactions: an exploratory market signaling studyTHE JOURNAL OF PRODUCT INNOVATION MANAGEMENT, Issue 3 2002Erik Jan Hultink When firms launch a new product into the marketplace they often aim to find a balance between building scale and provoking extensive and quick competitive reactions. Competitors react to new products when they perceive the product introduction as hostile, committed or when they feel that the product entry will have a large impact on their profitability. The present study develops a framework that shows how strong and fast incumbents react to perceived market signals resulting from a new product's launch decisions (broad targeting, penetration pricing, advertising intensity and product advantage). The strength of the relationships between the launch decisions and the perceived market signals was expected to depend on one industry characteristic (i.e., market growth) and on one entrant characteristic (i.e., aggressive reputation). We distinguished three market signals in our framework: hostility, commitment and consequences. Signal hostility refers to the extent to which the approach used by an acting firm to introduce the new product is perceived hostile whereas the commitment signal refers to the extent to which incumbents perceive the entrant firm to be committed to the new product introduction. The consequence signal is defined as the incumbents' perception of the impact of a new product entry on their profitability. We tested our framework using cross-sectional data provided by 73 managers in The Netherlands who recently reacted to a new product entry. The results clearly reveal which launch decisions create which market signals. For example, incumbents consider high advantage new products hostile and consequential. Penetration pricing and an intense advertising campaign are also considered hostile, especially in fast growing markets. Broad targeting is not perceived hostile, especially not when used by entrants with an aggressive reputation. In addition, this study explored the impact of three perceived market signals on the strength and speed of competitive reaction. The results reveal that perceived signals of hostility and commitment positively impact the strength of reaction, whereas the perceived consequence signal positively impacts the speed of reaction. The article concludes with the implications of our study for managers and academics. The relevance to managers was assessed from both the perspective of the incumbent firm that must defend, and that of the rival firm that is introducing the new product. [source] Dominance Effects from Local Competitors: Setting Institutional Parameters for Employment Relations in Multinational Subsidiaries; a Case from the Spanish Supermarket SectorBRITISH JOURNAL OF INDUSTRIAL RELATIONS, Issue 4 2009Tony Royle Dominance effects are normally associated with multinational corporations (MNCs). However, we argue that a strong local competitor can create ,dominance effects' setting the institutional parameters for employment relations in multinational subsidiaries. Moreover such an effect can be persistent. In this case the Spanish-owned El Corte Inglés (ECI) used its power and influence to establish an employer's federation and two ,yellow unions'. These yellow unions infiltrated the French-owned MNC Carrefour and most of the Spanish supermarket sector by the early 1980s and continue to dominate collective bargaining rounds and works council elections, marginalizing the main independent trade unions. This has resulted in poor pay and working conditions and a lack of effective employee representation across most of the Spanish supermarket sector. The fact that Carrefour established an international framework agreement to observe union rights in 2001 has as yet not changed this situation. [source] Competition and Cost Accounting: Adapting to Changing Markets,CONTEMPORARY ACCOUNTING RESEARCH, Issue 2 2002Ranjani Krishnan Abstract The relation of competition and cost accounting has been the subject of conflicting prescriptions, theories, and empirical evidence. Practitioner literature and textbooks argue that higher competition generally requires more accurate product costing. Theoretical economic analysis, in contrast, predicts that the optimal level of product-costing accuracy is sometimes higher at lower levels of competition. Results of survey research are inconsistent, suggesting a need for further identification of conditions under which higher competition leads to more accurate product costing. This study shows experimentally that individuals' choices of the level of product-costing accuracy depend not only on the current level of competition but also on the previous level of competition , that is, on an interaction between market structure (monopoly, duopoly, and four-firm competition) and market history (increasing versus decreasing competition). In the experiment, subjects decide on the quantity of data to collect at a pre-set price per datum to support more accurate product-cost estimates. Subjects collect the most cost data (i.e., choose the most accurate product costing) in monopoly, collect the least in duopoly, and an intermediate amount in the four-firm market, consistent with the pattern of optimal cost-data collection in Hansen's 1998 model. The process of convergence to the optimum differs significantly across market types and market histories, however. Subjects who begin in four-firm competition adapt more successfully to change than those who begin in monopoly. The lowest levels of decision performance occur when ex-monopolists face their first competitor: they overreact to this first encounter with competition and overspend on cost data. [source] Phorid fly parasitoids of invasive fire ants indirectly improve the competitive ability of a native antECOLOGICAL ENTOMOLOGY, Issue 5 2004Natasha J. Mehdiabadi Abstract., 1.,The red imported fire ant, Solenopsis invicta Buren (Hymenoptera: Formicidae), is an invasive species of south-eastern U.S.A. Since its introduction from South America approximately 70 years ago, this pest has devastated natural biodiversity. 2.,Due to such ecological costs, Pseudacteon phorid fly parasitoids (Diptera: Phoridae) from South America are being introduced into the U.S.A. as a potential biological control agent. Here, the indirect effects of these specialised parasitoids on an interspecific native ant competitor, Forelius mccooki (Hymenoptera: Formicidae), are evaluated. 3.,Over the course of a 50-day laboratory experiment, the results show that the native ant improved aspects of exploitative, but not interference, competition when S. invicta -attacking flies were present compared with when they were absent. 4.,Forelius mccooki colonies from the phorid treatment had approximately twice as many foragers at food baits relative to controls; however, there was no significant difference in interference aspects of competition or native ant colony growth between the two treatments. 5.,These results suggest that the S. invicta -specialised parasitoids help shift the competitive balance more in favour of F. mccooki than if these flies were not present; however, this competitive advantage does not translate into increased colony growth after 50 days. These laboratory findings are interpreted with regard to the more complex interactions in the field. [source] Refuge-mediated apparent competition in plant,consumer interactionsECOLOGY LETTERS, Issue 1 2010John L. Orrock Abstract At the intersection of consumer behaviour and plant competition is the concept of refuge-mediated apparent competition: an indirect interaction whereby plants provide a refuge for a shared consumer, subsequently increasing consumer pressure on another plant species. Here, we use a simple model and empirical examples to develop and illustrate the concept of refuge-mediated apparent competition. We find that the likelihood that an inferior competitor will succeed via refuge-mediated apparent competition is greater when competitors have similar resource requirements and when consumers exhibit a strong response to the refuge and high attack rates on the superior competitor. Refuge-mediated apparent competition may create an emergent Allee effect, such that a species invades only if it is sufficiently abundant to alter consumer impact on resident species. This indirect interaction may help explain unresolved patterns observed in biological invasion, such as the different physical structure of invasive exotic plants, the lag phase, and the failure of restoration efforts. Given the ubiquity of refuge-seeking behaviour by consumers and the ability of consumers to alter the outcome of direct competition among plants, refuge-mediated apparent competition may be an underappreciated mechanism affecting the composition and diversity of plant communities. Ecology Letters (2010) 13: 11,20 [source] Competition for food between Eurasian perch (Perca fluviatilis L.) and ruffe (Gymnocephalus cernuus [L.]) over different substrate typesECOLOGY OF FRESHWATER FISH, Issue 4 2004A. Dieterich Abstract,,, Food consumption by Eurasian perch (Perca fluviatilis L.) and ruffe (Gymnocephalus cernuus [L.]) was studied in single and mixed-species treatments in the laboratory, where alternative food resources, chironomids and zooplankton, were offered simultaneously. The effects of structural complexity, which was represented by substrate grain size, and of feeding level on food consumption were analysed. Across all experiments, the outcome of competition between perch and ruffe depended on food abundance and on the structural complexity of the environment. Perch and ruffe both changed their food consumption in the presence of a heterospecific competitor. With high food supply, perch consumed more benthic food than ruffe. With low food supply, the consumption of perch decreased strongly, while that of ruffe remained high on fine sediment. Under all conditions tested, the mechanism of competition appeared to be of interference rather than of exploitative nature. It is suggested that with decreasing lake productivity caused by re-oligotrophication, habitat shifts of both species will occur, which will alleviate interspecific competition. Ruffe will forage over fine sediment and perch over coarse sediment, whereby both species will achieve the highest foraging efficiency under conditions of low food supply. Resumen 1. Hemos estudiado el consumo alimenticio de Perca fluviatilis L. y Gymnocephalus cernuus (L.) en condiciones de laboratorio. Bajo tratamientos de especies individuales y mezcladas, les ofrecimos, simultáneamente, varios recursos alimenticios alternativos (quironómidos y zooplancton). 2. Analizamos los efectos de la complejidad estructural - representada por el tamaño del sustrato (arena, grava, y guijo) - y del nivel alimenticio, sobre el consumo alimenticio. Pusimos especial atención a la potencial influencia de competidores hetero-específicos sobre los patrones alimenticios de ambas especies, tanto en términos cualitativos como cuantitativos. Además, dado que en un futuro cercano una menor productividad general en lagos debida a re-oligotrofia, probablemente aumente la competición por el alimento en muchos lagos donde ambas especies co-existen, los experimentos se llevaron a cabo bajo niveles de abastecimiento alimenticio alto y bajo. 3. En los experimentos, la aparición de competencia entre P. fluviatilis y G. cernuus dependió de la abundancia del alimento y de la complejidad estructural del ambiente. El consumo de quironómidos por P. fluviatilis dependió del tipo de sustrato a niveles altos de abastecimiento alimenticio pero no a niveles bajos, mientras que en G. cernuus observamos lo contrario. 4. Ambas especies cambiaron el consumo alimenticio en presencia de un competidor hetero-específico. A altos niveles de abastecimiento alimenticio, P. fluviatilis consumió más bentos que G. cernuus. A niveles bajos, el consumo de P. fluviatilis decreció substancialmente mientras que el de G. cernuus permaneció alto en sedimento fino. Bajo todas las condiciones experimentales analizadas, los mecanismos de competición parecieron ser de interferencia más que de naturaleza explotativa. 5. Finalmente, presentamos un escenario sobre como P. fluviatilis y G. cernuus pueden competir por alimento bentónico en lagos con variado sustrato de fondo. Sugerimos que a altos niveles de abastecimiento alimenticio, G. cernuus forrajee más sobre arena y grava mientras que P. fluviatilis puede utilizar todos los sustratos disponibles. Al decrecer el abastecimiento alimenticio por re-oligotrofia, pueden producirse cambios en el hábitat de ambas especies que minimizarán la competencia inter-específica. G. cernuus forrajeará básicamente sobre sedimento fino, allá donde sea claramente superior a P. fluviatilis. Esta última especie forrajeará predominantemente sobre sedimento más grueso donde se enfrentará a competencia intra- e inter-específica. A través de estos cambios de hábitat, ambas especies podrían alcanzar la mayor eficiencia de forrajeo bajo condiciones de bajo abastecimiento alimenticio. [source] Helical- and ahelical-dependent chiral recognition mechanisms in capillary electrophoresis using amylose as the selectorELECTROPHORESIS, Issue 8 2009Weili Wei Abstract The present study discovered that helical structures of amylose were not always responsible for its chiral recognition abilities in CE. Several enantiomers with different structures were selected as models. Based on ultraviolet,visible spectroscopy and 13C NMR measurements, it was found that helical structures were gradually destroyed by temperature elevation and almost entirely transformed to extended ahelical structures above 60°C. Then, CE and 1H NMR chiral recognitions were investigated at different temperatures; chiral selectivity of the enantiomers varied in two different ways. Summarily, helical structures were necessary only for chiral separations of the enantiomers with small (<0.78,nm) and flexible molecular structures. However, for the gauche enantiomers (>0.78,nm) with high steric hindrances over their chiral centers, ahelical structures alone can realize chiral recognitions. By using iodine as a helix including competitor, it was further proved that helical structures functioned through the inclusive complexations only in the chiral separations of small enantiomers and had no effect for the others. The underlying mechanisms of the functions of helical and ahelical structures in molecular level were discussed as well. [source] Transcription factor binding study by capillary zone electrophoretic mobility shift assayELECTROPHORESIS, Issue 1-2 2003Zsolt Ronai Abstract Regulation of gene expression through interaction of proteins with specific DNA sequences is a central issue in functional genomics. Capillary electrophoretic mobility shift assay is an efficient novel method for the investigation of sequence specific protein-DNA interactions, allowing rapid and sensitive quantification of the complex formation. In this paper, we present a pilot study on capillary zone electrophoretic mobility shift assay (CZEMSA) to investigate the interaction between the transcription factors of HeLa nuclear extract and Sp1-specific fluorescein-labeled oligonucleotide, using the unlabeled probe as competitor. The mobility shift assay was accomplished by CZE in coated capillaries without polymeric buffer additives. Specificity of the DNA protein complex formation was verified by competition experiments, as well as by supershift assay with an anti-Sp1 antibody. The applied electric field strength did not affect the stability of DNA-protein complex during the electrophoretic analysis, allowing rapid identification and quantification of the protein DNA interaction. A practical application to study the interaction between Oryza sativa MADS-box transcription factor 4 (OsMADS4) and its consensus sequence is also reported. [source] Development of specific oligonucleotide probes for the identification and in situ detection of hydrocarbon-degrading Alcanivorax strainsENVIRONMENTAL MICROBIOLOGY, Issue 6 2001Kazuaki Syutsubo The genus Alcanivorax comprises diverse hydrocarbon-degrading marine bacteria. Novel 16S rRNA-targeted oligonucleotide DNA probes (ALV735 and ALV735-b) were developed to quantify two subgroups of the Alcanivorax/Fundibacter group by fluorescence in situ hybridization (FISH), and the conditions for the single-mismatch discrimination of the probes were optimized. The specificity of the probes was improved further using a singly mismatched oligonucleotide as a competitor. The growth of Alcanivorax cells in crude oil-contaminated sea water under the biostimulation condition was investigated by FISH with the probe ALV735, which targeted the main cluster of the Alcanivorax/Fundibacter group. The size of the Alcanivorax population increased with increasing incubation time and accounted for 91% of the 4,,6-diamidino-2-phenylindole (DAPI) count after incubation for 2 weeks. The probes developed in this study are useful for detecting Alcanivorax populations in petroleum hydrocarbon-degrading microbial consortia. [source] The Smell of New Competitors: The Response of American Mink, Mustela vison, to the Odours of Otter, Lutra lutra and Polecat, M. putoriusETHOLOGY, Issue 5 2009Lauren A. Harrington We tested the response of wild American mink (an established alien species in the UK), to the odours of unfamiliar mink, European polecat and Eurasian otter. Polecats are similar in size and habits to mink, otters are larger than mink and a dominant competitor; both are native to the UK and both were absent during the original colonization by mink but are now undergoing natural population recoveries. The response of mink to experimental odours was assessed by counting the numbers of tracks (footprints) on rafts treated with anal gland secretions, and compared with response to a control raft, on two rivers in the Upper Thames valley, UK. Remote video showed that the number of tracks was positively correlated with the time that mink spent investigating an odour. We found that mink were attracted to the odours of both unfamiliar mink and polecats. There was little evidence that mink avoided the odour of otters. We suggest that, during an encounter with a polecat, mink may behave much as they would to a conspecific. We infer from the response of mink to the odour of otters, that, if mink do avoid otters, the mechanism of avoidance is likely to be complex, situation-dependent and perhaps affected by prior experience. [source] Innate and Learned Components of Defence by Flickers Against a Novel Nest Competitor, the European StarlingETHOLOGY, Issue 10 2004Karen L. Wiebe Defence against predators is an important component of fitness in wild birds but the first step of defence, predator recognition, is not well understood. Anti-predator behaviour may innate, in which case the individual responds without prior contact with that predator, and/or there may be a learned component that develops only after direct experience. In the wild, the development of anti-predator behaviour is studied by exposing naive individuals to novel predators. I studied responses of 71 naive and experienced northern flickers Colaptes auratus, to a novel nest predator and competitor, the European starling Sturnus vulgaris that was introduced to North America. Naive individuals responded more intensely to the model starling than to the control model suggesting an innate component to recognition. However, there was also a learned component to defence because flickers nesting near to starlings reacted more aggressively than naive individuals far from starlings. Consistent with theory on life histories and optimal defence levels, no significant differences in aggression were found between the sexes or between age classes. Selection should favour more intense, and possibly innate, defence against the introduced starling. Variation in responses of naive individuals suggests that there may already be some alleles in the population associated with higher defence, but that these may not be uniform within the population. [source] Foraging Behaviour of Subordinate Great Tits (Parus major).ETHOLOGY, Issue 10 2001Can Morphology Reduce the Cost of Subordination? This paper studies the magnitude of the behavioural shift, from forage standing to forage hanging, of subordinate great tits (Parus major) in two different social contexts: feeding solitarily vs. feeding with a dominant conspecific. The aim is to test the hypothesis that differences in morphological design provide subordinates with varying abilities to reduce the presumed costs of subordination. We find that different subordinate individuals change the foraging behaviour, occupying a different niche when an intra-specific competitor is present. Morphology linked to sexual dimorphism, specifically body mass, is the factor responsible for the different magnitudes of change. Lighter subordinates can remain longer than heavier ones at the feeding patch without interrupting their foraging. Thereby, the former reduce the costs of being subordinate more than the latter. Among subordinates, females are lighter than males; they also spend more time feeding in the presence of a dominant conspecific than males do. No differences are found between age categories. We find no relationship between tarsus length and individual ecological plasticity. Our results support the idea that the ecological plasticity due to morphological differences is a mechanism that allows subordinate individuals to overcome costs associated with subordination. [source] VIRULENCE AND COMPETITIVENESS: TESTING THE RELATIONSHIP DURING INTER- AND INTRASPECIFIC MIXED INFECTIONSEVOLUTION, Issue 9 2010Peter A. Staves Understanding the reasons why different parasites cause different degrees of harm to their hosts is an important objective in evolutionary biology. One group of models predicts that if hosts are infected with more than one strain or species of parasite, then competition between the parasites will select for higher virulence. While this idea makes intuitive sense, empirical data to support it are rare and equivocal. We investigated the relationship between fitness and virulence during both inter- and intraspecific competition for a fungal parasite of insects, Metarhizium anisopliae. Contrary to theoretical expectations, competition favored parasite strains with either a lower or a higher virulence depending on the competitor: when in interspecific competition with an entomopathogenic nematode, Steinernema feltiae, less virulent strains of the fungus were more successful, but when competing against conspecific fungi, more virulent strains were better competitors. We suggest that the nature of competition (direct via toxin production when competing against the nematode, indirect via exploitation of the host when competing against conspecific fungal strains) determines the relationship between virulence and competitive ability. [source] CHARACTER DISPLACEMENT AS THE "BEST OF A BAD SITUATION": FITNESS TRADE-OFFS RESULTING FROM SELECTION TO MINIMIZE RESOURCE AND MATE COMPETITIONEVOLUTION, Issue 10 2005Karin S. Pfennig Abstract Character displacement has long been considered a major cause of adaptive diversification. When species compete for resources or mates, character displacement minimizes competition by promoting divergence in phenotypes associated with resource use (ecological character displacement) or mate attraction (reproductive character displacement). In this study, we investigated whether character displacement can also have pleiotropic effects that lead to fitness trade-offs between the benefits of avoiding competition and costs accrued in other fitness components. We show that both reproductive and ecological character displacement have caused spadefoot toads to evolve smaller body size in the presence of a heterospecific competitor. Although this shift in size likely arose as a by-product of character displacement acting to promote divergence between species in mating behavior and larval development, it concomitantly reduces offspring survival, female fecundity, and sexual selection on males. Thus, character displacement may represent the "best of a bad situation" in that it lessens competition, but at a cost. Individuals in sympatry with the displaced phenotype will have higher fitness than those without the displaced trait because they experience reduced competition, but they may have reduced fitness relative to individuals in allopatry. Such a fitness trade-off can limit the conditions under which character displacement evolves and may even increase the risk of "Darwinian extinction" in sympatric populations. Consequently, character displacement may not always promote diversification in the manner that is often expected. [source] DIVERSIFYING COEVOLUTION BETWEEN CROSSBILLS AND BLACK SPRUCE ON NEWFOUNDLANDEVOLUTION, Issue 8 2002Thomas L. Parchman Abstract Coevolution is increasingly recognized as an important process structuring geographic variation in the form of selection for many populations. Here we consider the importance of a geographic mosaic of coevolution to patterns of crossbill (Loxia) diversity in the northern boreal forests of North America. We examine the relationships between geographic variation in cone morphology, bill morphology, and feeding performance to test the hypothesis that, in the absence of red squirrels (Tamiasciurus hudsonicus), black spruce (Picea mariana) has lost seed defenses directed at Tamiasciurus and that red crossbills (L. curvirostra) and black spruce have coevolved in an evolutionary arms race. Comparisons of cone morphology and several indirect lines of evidence suggest that black spruce has evolved defenses in response to Tamiasciurus on mainland North America but has lost these defenses on Newfoundland. Cone traits that deter crossbills, including thicker scales that require larger forces to separate, are elevated in black spruce on Newfoundland, and larger billed crossbills have higher feeding performances than smaller billed crossbills on black spruce cones from Newfoundland. These results imply that the large bill of the Newfoundland crossbill (L. c. percna) evolved as an adaptation to the elevated cone defenses on Newfoundland and that crossbills and black spruce coevolved in an evolutionary arms race on Newfoundland during the last 9000 years since glaciers retreated. On the mainland where black spruce is not as well defended against crossbills, the small-billed white-winged crossbill (L. leucoptera leucoptera) is more efficient and specializes on seeds in the partially closed cones. Finally, reciprocal adaptations between crossbills and conifers are replicated in black spruce and Rocky Mountain lodgepole pine (Pinus contorta ssp. latifolia), with coevolution most pronounced in isolated populations where Tamiasciurus are absent as a competitor. This study further supports the role of Tamiasciurus in determining the selection mosaic for crossbills and suggests that a geographic mosaic of coevolution has been a prominent factor underlying the diversification of North American crossbills. [source] CHARACTER DISPLACEMENT IN POLYPHENIC TADPOLESEVOLUTION, Issue 5 2000David W. Pfennig Abstract Biologists have long known that closely related species are often phenotypically different where they occur together, but are indistinguishable where they occur alone. The causes of such character displacement are controversial, however. We used polyphenic spadefoot toad tadpoles (Spea bombifrons and S. multiplicata) to test the hypothesis that character displacement evolves to minimize competition for food. We also sought to evaluate the role of phenotypic plasticity in the mediation of competitive interactions between these species. Depending on their diet, individuals of both species develop into either a small-headed omnivore morph, which feeds mostly on detritus, or a large-headed carnivore morph, which specializes on shrimp. Laboratory experiments and surveys of natural ponds revealed that the two species were more dissimilar in their tendency to produce carnivores when they occurred together than when they occurred alone. This divergence in carnivore production was expressed as both character displacement (where S. multiplicata's propensity to produce carnivores was lower in sympatry than in allopatry) and as phenotypic plasticity (where S. multiplicata facultatively enhanced carnivore production in S. bombifrons, and S. bombifrons facultatively suppressed carnivore production in S. multiplicata). In separate experiments, we established that S. bombifrons (the species for which carnivore production was enhanced) was the superior competitor for shrimp. Conversely, S. multiplicata (the species for which carnivore production was suppressed and omnivore production enhanced) was the superior competitor for detritus. These results therefore demonstrate that selection to minimize competition for food can cause character displacement. They also suggest that both character displacement and phenotypic plasticity may mediate competitive interactions between species. [source] Predator-released compounds, ambient temperature and competitive exclusion among differently sized Daphnia speciesFRESHWATER BIOLOGY, Issue 4 2006JACOBUS VIJVERBERG Summary 1. We studied the effects of fish water and temperature on mechanisms of competitive exclusion among two Daphnia species in flow-through microcosms. The large-bodied D. pulicaria outcompeted the medium sized D. galeata × hyalina in fish water, but not in the control treatment. Daphnia galeata × hyalina was competitively displaced 36 days earlier at 18 °C than at 12 °C. 2. It is likely that the high phosphorus content of fish water increased the nutritional value of detrital seston particles by stimulating bacterial growth. Daphnia pulicaria was presumably better able to use these as food and hence showed a more rapid somatic growth than its competitor. This led to very high density of D. pulicaria in fish water, but not in the controls. The elevated D. pulicaria density coincided with high mortality and reduced fecundity in D. galeata × hyalina, resulting in competitive displacement of the hybrid. 3. It is clear that the daphnids competed for a limiting resource, as grazing caused a strong decrease in their seston food concentration. However, interference may also have played a role, as earlier studies have shown larger Daphnia species to be dominant in this respect. The high density of large-bodied D. pulicaria in fish water may have had an allelopathic effect on the hybrid. Our data are inconclusive with respect to whether the reached seston concentration was below the threshold resource level (R*) of the hybrid, where population growth rate and mortality exactly balance, as it would be set in the absence of interference, or whether interference actually raised the hybrid's R* to a value above this equilibrium particle concentration. 4. Our results do clearly show that fish-released compounds mediated competitive exclusion among zooplankton species and that such displacement occurred at a greatly enhanced rate at an elevated temperature. Fish may thus not only structure zooplankton communities directly through size-selective predation, but also indirectly through the compounds they release. [source] Is competition important to arctic zooplankton community structure?FRESHWATER BIOLOGY, Issue 9 2004Andrew R. Dzialowski Summary 1. Daphnia pulex and Daphnia middendorffiana are commonly found in the Toolik Lake region of arctic Alaska. These two species are very similar morphologically, although their natural distributions differ markedly: D. pulex is restricted to shallow ponds, while D. middendorffiana is widely distributed and found in a variety of ponds and lakes. We compared the reproductive capabilities of D. pulex and D. middendorffiana grown under similar resource conditions and in the absence of the invertebrate predator Heterocope septentrionalis. In situ life table and mesocosm experiments were conducted in Toolik Lake and Dam Pond, habitats that have historically contained natural populations of D. middendorffiana, but never D. pulex. 2. Daphnia pulex exhibited a significantly higher net growth rate than D. middendorffiana in both life table and mesocosm experiments although D. pulex has never been found in either Toolik Lake or Dam Pond. Daphnia middendorffiana exhibited a negative net growth rate in Dam Pond, which had lower resource levels then Toolik Lake. Therefore, the smaller D. pulex appears to have a lower food threshold concentration than the larger D. middendorffiana. 3. Our results indicate that D. pulex is a superior resource competitor in the Toolik Lake region. These results combined with distributional patterns suggest that the restricted distribution of D. pulex in these arctic lakes and ponds cannot be explained by resource competition alone. We suggest that in the presence of H. septentrionalis, predation is an important factor structuring arctic zooplankton communities in the Toolik Lake region. [source] On the different nature of top-down and bottom-up effects in pelagic food websFRESHWATER BIOLOGY, Issue 12 2002Z. Maciej Gliwicz SUMMARY 1.,Each individual planktonic plant or animal is exposed to the hazards of starvation and risk of predation, and each planktonic population is under the control of resource limitation from the bottom up (growth and reproduction) and by predation from the top down (mortality). While the bottom-up and top-down impacts are traditionally conceived as compatible with each other, field population-density data on two coexisting Daphnia species suggest that the nature of the two impacts is different. Rates of change, such as the rate of individual body growth, rate of reproduction, and each species' population growth rate, are controlled from the bottom up. State variables, such as biomass, individual body size and population density, are controlled from the top down and are fixed at a specific level regardless of the rate at which they are produced. 2.,According to the theory of functional responses, carnivorous and herbivorous predators react to prey density rather than to the rate at which prey are produced or reproduced. The predator's feeding rate (and thus the magnitude of its effect on prey density) should hence be regarded as a functional response to increasing resource concentration. 3.,The disparity between the bottom-up and top-down effects is also apparent in individual decision making, where a choice must be made between accepting the hazards of hunger and the risks of predation (lost calories versus loss of life). 4.,As long as top-down forces are effective, the disparity with bottom-up effects seems evident. In the absence of predation, however, all efforts of an individual become subordinate to the competition for resources. Biomass becomes limited from the bottom up as soon as the density of a superior competitor has increased to the carrying capacity of a given habitat. Such a shift in the importance of bottom-up control can be seen in zooplankton in habitats from which fish have been excluded. [source] Factors influencing Willow Tit Poecile montanus site occupancy: a comparison of abandoned and occupied woodsIBIS, Issue 2007ALEX J. G. LEWIS The British Willow Tit Poecile montanus kleinschmidti underwent a decline of 85% between 1970 and 2003. The cause of this decline is unknown. However, several hypotheses have been put forward to account for it: competition from other tit species, predation by Great Spotted Woodpeckers Dendrocopos major and habitat change. In order to test these, woods that are currently occupied by Willow Tits were paired with woods (within 50 km) that had been abandoned by Willow Tits five or more years previously. Point counts for other tit species (potential competitors) and woodpecker species (potential predators) were carried out at ten evenly spaced points throughout each wood. Habitat variables were collected within a 50-m radius of where a Willow Tit was located (in the occupied woods) or where maps showed a Willow Tit had been located (for abandoned woods). No evidence was found for differences in numbers of potential competitor or potential predator species in abandoned and occupied woods. Soil water content was found to be higher at occupied sites. No other habitat features differed between the two categories of site. The drying up of British woods could therefore be implicated in the Willow Tit decline and this warrants further investigation. [source] The concept of the gas turbine-based hybrid vehicle: system, design and configuration issuesINTERNATIONAL JOURNAL OF ENERGY RESEARCH, Issue 9 2006Roberto Capata Abstract The object of this study is a theoretical and experimental analysis of a new hybrid propulsion system for a passenger sedan in which the thermal engine is a small gas turbine set. Some preliminary results on the possibility of replacing the conventional ICE of a hybrid ,series' configuration by a turbogas were presented and discussed in previous papers by the same authors: several possible designs were examined under both a thermodynamic and an operative point of view. This paper presents a summary of the project and constitutes an attempt to put things in a proper engineering perspective: the technical feasibility of the project is assessed via a calculation of the required mission loads, a preliminary design of the most important elements of the propulsive system, the choice of the mission control strategy and the implementation of a numerical system simulator. The experiments that provided a verification for the assumed component efficiencies were carried out, in cooperation with the Research Centre of ENEA-Casaccia, on an ELLIOTT TA-45 group. Our results, though only preliminary, allow for a direct comparison between a GT-hybrid vehicle and a modern diesel car, and indicate that the GT-hybrid may be actually a competitor for the FC-powered vehicle concept. Our ,optimal' configuration is a combination of a 100 kg battery pack and two turbogas set of 5 and 16 kW, respectively. Copyright © 2005 John Wiley & Sons, Ltd. [source] | ||||||||||||||||||||||||||||||||||||||||||||||||||||